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Figures 16 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 16 - Line drawings of ring sclerite of Guatemalan Geocharidius species, male genitalia, dorsal aspect. A Geocharidius gimlii (GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype B Geocharidius erwini (GUATEMALA, Quiché, Los Encuentros), paratype C Geocharidius minimus (GUATEMALA, Sacatepéquez, Antigua), paratype D Geocharidius jalapensis (GUATEMALA, Jalapa, Mataquescuintla), paratype E Geocharidius balini (GUATEMALA, Suchitepéquez, Volcano Atitlán), paratype F Geocharidius longinoi (GUATEMALA, El Progreso, Cerro Pinalón), paratype. Scale = 0.1mm.

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Figures 13 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 13 - Line drawings of aedeagus of Guatemalan Geocharidius species. A–C Geocharidius gimlii (GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype: A median lobe with internal sac and dorsal sclerites, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect D–F Geocharidius erwini (GUATEMALA, Quiché, Los Encuentros), paratype: D median lobe with internal sac and dorsal sclerites, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect G–J Geocharidius minimus (GUATEMALA, Sacatepéquez, Antigua), paratype: G median lobe with internal sac and dorsal sclerites, right lateral aspect H dorsal sclerite of median lobe, dorsal aspect I left paramere, left lateral aspect J right paramere, right lateral aspect K–N Geocharidius jalapensis (GUATEMALA, Jalapa, Mataquescuintla), paratype: K median lobe with internal sac and dorsal sclerites, right lateral aspect L dorsal sclerite of median lobe, dorsal aspect M left paramere, left lateral aspect N right paramere, right lateral aspect O–Q Geocharidius balini (GUATEMALA, Suchitepéquez, Volcano Atitlán), paratype: O median lobe with internal sac and dorsal sclerites, right lateral aspect P left paramere, left lateral aspect Q right paramere, right lateral aspect R–T Geocharidius longinoi (GUATEMALA, El Progreso, Cerro Pinalón), paratype: R median lobe with internal sac and dorsal sclerites, right lateral aspect S left paramere, left lateral aspect T right paramere, right lateral aspect. Scale = 0.05mm.

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Figures 12 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 12 - Digital images of habitus of Guatemalan Geocharidius species. A Geocharidius gimlii (GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype B Geocharidius integripennis (GUATEMALA, Totonicapán, "Totonicapam"), lectotype C Geocharidius longinoi (GUATEMALA, El Progreso, Cerro Pinalón), paratype D Geocharidius jalapensis (GUATEMALA, Jalapa, Mataquescuintla), paratype E Geocharidius erwini (GUATEMALA, Quiché, Los Encuentros), paratype F Geocharidius balini (GUATEMALA, Suchitepéquez, Volcano Atitlán), paratype G Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), paratype H Geocharidius minimus (GUATEMALA, Sacatepéquez, Antigua), paratype. Scale = 0.5mm.

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Figures 1 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 1 - SEM illustrations of structural features of body parts of Geocharidius species. A–C head, dorsal aspect: A Geocharidius balini B Geocharidius zullinii C G. andersoni. D–F right half of pronotum, dorsal aspect: D Geocharidius balini E Geocharidius zullinii F Geocharidius andersoni. G–I basal part of right elytron: G Geocharidius balini H Geocharidius zullinii I Geocharidius andersoni Legend: bm – basal margin; cl – clypeus; ft – frontal tubercle; lb – labrum; mp3 – maxillary palpomere 3; mp4 – maxillary palpomere 4. Scale = 0.05mm.

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Figures 2 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 2 - SEM illustrations of structural features and shape of elytra of Geocharidius species, left lateral aspect. A Geocharidius erwini B Geocharidius jalapensis C Geocharidius comayaguanus D Geocharidius phineus. Legend: ed2 – scutellar seta; ed8 – apical seta; eo1-9 – setae 1-9 from the umbilicate series. Scale = 0.2mm.

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Figures 4 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 4 - SEM illustrations of structural features of mandibles of Geocharidius species. A–B Geocharidius zullinii: dorsal aspect of left and right mandibles, respectively C–D Geocharidius jalapensis: dorsal aspect of left and right mandibles, respectively E–F Geocharidius zullinii:–ventral aspect of left and right mandibles, respectively G–H Geocharidius jalapensis: ventral aspect of left and right mandibles, respectively (right mandible with Λ-shaped crack apically from retinacular ridge). Legend: art – anterior retinacular tooth; bemr – molar ridge, basal extension; it – incisor tooth; mr – molar ridge; mt – molar tooth; pog – posterior occlusal grove; prt – posterior retinacular tooth; rr – retinacular ridge; tr – terebral ridge; tt – terebral tooth; vg – ventral groove; vm – microtrichia. Scale = 0.05mm.

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Figures 17 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 17 - Line drawings of spermatheca of Guatemalan Geocharidius species. A Geocharidius integripennis (GUATEMALA, Totonicapán, "Totonicapam"), holotype B Geocharidius erwini (GUATEMALA, Quiché, Los Encuentros), paratype C Geocharidius minimus (GUATEMALA, Sacatepéquez, Antigua), paratype D Geocharidius jalapensis (GUATEMALA, Jalapa, Mataquescuintla), paratype E Geocharidius balini (GUATEMALA, Suchitepéquez, Volcano Atitlán), paratype F Geocharidius longinoi (GUATEMALA, El Progreso, Cerro Pinalón), paratype. Scale = 0.05mm.

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Figure 22 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figure 22 - Map of southern Mexico, Guatemala and adjacent part of Honduras, showing positions of locality records for the species of Geocharidius: white diamond, Geocharidius andersoni; white circles (black point in a circle shows "terra typica" for the species), Geocharidius zullinii; white squares, Geocharidius vignatagliantii; white triangle, Geocharidius gimlii; black and white triangle, Geocharidius integripennis; green squares, Geocharidius longinoi; green circles, Geocharidius erwini; green flowers, Geocharidius minimus; green diamonds, Geocharidius balini; green triangle, Geocharidius jalapensis; yellow quadrangle, Geocharidius antigua; yellow diamond, Geocharidius lencanus; yellow flower, Geocharidius celaquensis; yellow triangles, Geocharidius disjunctus; yellow circles, Geocharidius comayaguanus. Physiographic features: CH, Chiapas Highlands; GC, Guatemalan Cordillera; HIH, Honduran Interior Highlands; MTZ, Motagua Fault Zone; SC, Sierra de los Cuchumatanes; SMC, Sierra Madre de Chiapas. Elevation scale bar is given in meters.

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Figures 15 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 15 - Schematic line drawings of median lobe and dorsal sclerites of internal sac of two previously described Guatemalan Geocharidius species. A Geocharidius integripennis (GUATEMALA, Totonicapán, "Totonicapam"), holotype B Geocharidius gimlii (GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype, both left lateral aspect.

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Figures 11 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 11 - Line drawings of spermatheca of Mexican Geocharidius species. A Geocharidius zullinii (MEXICO, Chiapas, Reserva Huitepec) B Geocharidius vignatagliantii (MEXICO, Chiapas, Motozintla), paratype C Geocharidius andersoni (MEXICO, Chiapas, Cerro Huitepec), paratype. Legend: c – cornu; n – nodulus; sd – spermathecal duct; sg – spermathecal gland; sp – spermatheca. Scale = 0.05mm.

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Figures 10 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 10 - Line drawings of ring sclerite of Mexican Geocharidius species, male genitalia, dorsal aspect. A Geocharidius zullinii (MEXICO, Chiapas, Reserva Huitepec) B Geocharidius vignatagliantii (MEXICO, Chiapas, Motozintla), paratype C Geocharidius andersoni (MEXICO, Chiapas, Cerro Huitepec), holotype. Legend: hd – handle of ring sclerite. Scale = 0.1mm.

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Figures 21 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 21 - Line drawings of spermatheca of Guatemalan and Honduran Geocharidius species. A Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), paratype B Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), paratype C Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype D Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype E Geocharidius disjunctus (HONDURAS, Yoro, Pico Pijol National Park). Scale = 0.05mm.

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Figures 14 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 14 - Photographs of labels for type specimens of Geocharidius species. A Geocharidius gimlii, holotype B Geocharidius integripennis, lectotype C Geocharidius integripennis, paralectotype.

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Figures 20 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 20 - Line drawings of ring sclerite of Guatemalan and Honduran Geocharidius species, male genitalia, dorsal aspect. A Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), holotype B Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), holotype C Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype D Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype E Geocharidius disjunctus (HONDURAS, Francisco Morazán, La Tigra National Park), holotype. Scale = 0.1mm.

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Figures 19 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 19 - Line drawings of aedeagus of Guatemalan and Honduran Geocharidius species. A–D Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), holotype: A median lobe with internal sac and dorsal sclerites, right lateral aspect B dorsal sclerite of median lobe, dorsal aspect C left paramere, left lateral aspect D right paramere, right lateral aspect E–G Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), holotype: E median lobe with internal sac and dorsal sclerites, right lateral aspect F left paramere, left lateral aspect G right paramere, right lateral aspect H–J Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype: H median lobe with internal sac and dorsal sclerites, right lateral aspect I left paramere, left lateral aspect J right paramere, right lateral aspect K–N Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype: K median lobe with internal sac and dorsal sclerites, right lateral aspect L variation in a shape of dorsal sclerite of internal sac, right lateral aspect M left paramere, left lateral aspect N right paramere, right lateral aspect O–P Geocharidius comayaguanus (HONDURAS, La Paz, Guajicuiro): variations in a shape of dorsal sclerite of internal sac, right lateral aspect Q–S Geocharidius disjunctus (HONDURAS, Francisco Morazán, La Tigra National Park), holotype: Q median lobe with internal sac and dorsal sclerites, right lateral aspect R left paramere, left lateral aspect S right paramere, right lateral aspect. T Geocharidius disjunctus (HONDURAS, Yoro, Pico Pijol National Park): shape of dorsal sclerite of median lobe, right lateral aspect. Scale = 0.05mm.

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Figures 3 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 3 - SEM illustrations of structural features of body parts of Geocharidius species. A–C prothorax, ventral aspect: A Geocharidius zullinii B Geocharidius erwini C Geocharidius comayaguanus. D–F pterothorax, ventral aspect: D Geocharidius jalapensis E Geocharidius minimus F Geocharidius comayaguanus. G–I abdominal ventrites 3-5: G Geocharidius minimus, male H Geocharidius jalapensis, male I Geocharidius jalapensis female. Legend: asts – abdominal sternal terminal seta; ipa –intercoxal process of abdominal ventrite 2; mscx – mesocoxa; msp – mesosternal process; mtcx – metacoxa; mtp – metasternal process; mtv – metaventrite; pas – prosternal ambulatory seta; pep – proepipleuron; pes – proepisternum; prcx – procoxa; ps – prosternum; psp – prosternal process; v3-v5 – abdominal ventrites 3-5. Scale = 0.05mm.

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Figures 5 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 5 - SEM illustrations of structural features of labial complex of Geocharidius species. A Geocharidius erwini B Geocharidius balini C Geocharidius lencanus D Geocharidius zullinii E Geocharidius longinoi F Geocharidius minimus. Legend: gsc – glossal sclerite; lms – lateral mental seta; lss – lateral submental seta; m – mentum; mss – medial submental seta; pms – paramedial mental seta; prss – primary basal submental seta; sm – submentum. Scale = 0.05mm.

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Figures 23 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 23 - Diagrams illustrating size variation for sympatric pairs of closely and remotely related species of the integripennis group at different geographical localities of Nuclear Central America. Closely related pairs (for shared characters see subchapters on Relationships in the text for corresponding species): A Chiapas Highlands, Mexico B Mataquescuintla, Guatemala C Celaque National Park, Honduras. Remotely related species: D Volcano Agua, Guatemala E Volcano Atitlan, Guatemala F Pico Pijol National Park, Honduras. Legend: black dot – median; box – 25–75% range of values; whiskers – non-outlier range of values; b – coefficient of regression.

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Figures 18 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 18 - Digital images of habitus of Honduran Geocharidius species. A Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), paratype B Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype C Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype D Geocharidius disjunctus (HONDURAS, Francisco Morazán, La Tigra National Park), holotype. Scale = 0.5mm.

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Figure 8 from: Kavanaugh D, Sokolov I, Reddell J (2014) Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. ZooKeys 417: 71-101. https://doi.org/10.3897/zookeys.417.7733

Figure 8 - Locality records for Anillinus species: Anillinus acutipennis – white stars; Anillinus affabilis – black triangle; Anillinus comalensis – white diamond; Anillinus depressus – black diamond; Anillinus forthoodensis – white quadrangle; Anillinus lescheni – black circle Anillinus sinuatus– black quadrangle; Anillinus wisemanensis– white triangles; Anillinus spp. – white circles (1 – Bell and Coryell Counties; 2 – Williamson County; 3 – Bexar County). Violet color – range of the western Anillinus species in Arkansas and Oklahoma (Sokolov et al. 2004). Light blue color – recharge zone of the Edwards Aquifer, dark blue color – the Balcones Escarpment, topographic expression of the Balcones Fault Zone (Woodruff and Abbot 1986). Contour of the Cross Timbers ecological region (highlighted) were taken from Wikipedia: http://en.wikipedia.org/wiki/Cross_Timbers. Elevation scale bar is given in meters.

opencc-by-4.0Jun 2014View details →

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allen-brain-atlas
neuroscienceopenDocumentation, web resources, and API references are available online.
Last verified 2026-04-30Open record

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Last verified 2026-04-29Open record

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Last verified 2026-04-29Open record