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FIGURE 35 in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 35. Geographic distribution of Parascombrops analis, P. glossodon, P. madagascariensis and P. yamanouei. Large symbols indicate location of holotypes. One symbol may represent several samples.
FIGURE 34 in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 34. Geographic distribution of Caraibops trispinosus, Kaperangus microlepis, Parascombrops spinosus (including fossil records of Caraibops trispinosus and Kaperangus malzi). Large symbols indicate location of holotypes. One symbol may represent several samples.
FIGURE 33A–P in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 33A–P. Fossil otoliths of the genera Kaperangus, Parascombrops and Ƒerilus: A–C, Kaperangus malzi (Schwarzhans, 2010), A, paratype, RGM 442955, Middle Miocene, Miste, Netherlands, B–C, holotype, SMF PO 64937, Middle Miocene, Hoerstgen, Germany; Parascombrops argyreus (Gilbert & Cramer, 1897), coll. Ohe, Late Pliocene, Okinawa; H–K, Parascombrops ohei n. sp., coll. Ohe, H, Late Pliocene, Dainichi, Japan; I–K, Late Pliocene, Hijikata, Japan; L–M, Ƒerilus mutinensis (Bassoli, 1906), SMF P 2797, Middle Miocene, Kostej, Romania; N–O, Parascombrops? brzobohatyi Nolf, 1988, Late Oligocene, Aquitaine, France (refigured from Nolf 2013); P, Parascombrops? epigonoides Nolf & Brzobohaty, 2000, Early Miocene, Aquitaine, France (refigured from Nolf 2013).
FIGURE 32A–D in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 32A–D. Ƒerilus pseudomicrolepis (Schultz, 1940); A–B, USNM 436691, 88 mm SL. Ƒerilus sordidus Poey, 1860; C,CAS 30647, 225 mm SL., D, USNM 12565, 186 mm SL.
FIGURE 27A–D in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 27A–D. Number of pseudobranchial filaments (A) and gill rakers (B) against specimen size in Parascombrops argyreus, P. nakayamai and P. philippinensis; and snout length versus orbital diameter (C) and snout length as % of orbital diameter as a function of SL (D) in P. nakayamai and P. philippinensis.
FIGURE 26A–D in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 26A–D. Parascombrops philippinensis (Günther, 1880); A–B, BMNH 1879.5.14.167, lectotype, 70 mm SL; C–D, Synagrops malayanus Weber, 1913, lectotype, ZMS 112.453, 52 mm SL.
FIGURE 25A–B in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 25A–B. Number of pseudobranchial filaments and gill rakers against specimen size in Parascombrops pellucidus, specifically for Eastern (type region of P. pellucidus) and Western Indian Ocean (type region of Synagrops adeni).
FIGURE 24A–D. Parascombrops pellucidus Alcock, 1889 in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 24A–D. Parascombrops pellucidus Alcock, 1889; A–B, USNM 392727, topotypic specimen of P. pellucidus, 75 mm SL; C–D, BMNH 1939.5.24.880-891, topotypic specimen of Synagrops adeni, 65 mm SL.
FIGURE 17A–B in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 17A–B. Parascombrops argyreus (Gilbert & Cramer, 1897); A, BPBM 23852, 120 mm SL; B, AMS I.42758-006, 120 mm SL.
FIGURE 15A in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 15A–AC. Otoliths (inner faces and representative ventral views): A–F, Parascombrops pellucidus (A–C = BMNH 1939.5.24.892-894, D–F = BMNH 1939.5.24.880-891); G–L, Parascombrops philippinensis (G–H = NSMT-P 59313, I = BSKU 116496, J = NSMT-P 57379, K–L = ZMA 112. 454, Synagrops malayanus paralectotype); M–O, Parascombrops spinosus (M–N = MCZ 169467, O = USNM 395230); P–R, Parascombrops serratospinosus (USNM 99324); S–U, Parascombrops yamanouei n.sp. (NMMB-P 23097); V–X, Caraibops trispinosus (15V–W = USNM 407771, 15X = USNM 407745); Y–AA, Kaperangus microlepis (CAS 222834); AB–AC, Ƒerilus pseudomicrolepis (USNM 407758).
FIGURE 12A–E in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 12A–E. Gills and pseudobranchial filaments: A, gill arch showing many developed rakers (Parascombrops glossodon n.sp., USNM 436685, paratype, 64 mm SL); B, gill arch with a low number of developed rakers and atrophied remnants of rakers at each end of the arch (Parascombrops nakayamai n.sp., MNHN 1998-0993, paratype, 100 mm SL); C–D, pseudobranch with low number of filaments (C = Kaperangus microlepis, CAS 222834, 78.5 mm SL; D = Parascombrops glossodon n.sp., USNM 436685, paratype, 64 mm SL); E, pseudobranch with many filaments (Parascombrops argyreus, BPBM 23839, 119 mm SL).
FIGURE 13A–Q in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 13A–Q. Anal-fin pterygiophores and shape of large first anal-fin pterygiophore of genera Parascombrops, Caraibops, Kaperangus, Synagrops and of Ƒerilus pseudomicrolepis. A, Parascombrops analis, BSKU 94343, 74.5 mm SL; B, Parascombrops argyreus, AMS I.42758-006, 120 mm SL; C, Parascombrops glossodon n.sp., USNM 436684, paratype, 73 mm SL; D, Parascombrops madagascariensis n.sp., MNHN 2008-1769, holotype, 68.5 mm SL; E, Parascombrops mochizukii n.sp., NMMBA-P 3887, paratype, 98 mm SL; F, Parascombrops nakayamai n.sp., BSKU 102255, holotype, 97 mm SL; G, Parascombrops ohei n.sp., BSKU 72614, holotype, 75 mm SL; H, Parascombrops parvidens n.sp., BSKU 15219, paratype, 70 mm SL; I, Parascombrops pellucidus, USNM 44428, 65.5 mm SL; J, Parascombrops philippinensis, BMNH 2016.9.14.1, paralectotype, 64 mm SL; K, Parascombrops serratospinosus, CAS 238016, 67.5 mm SL; L, Parascombrops spinosus, MNHN 1989-0411, 82 mm SL; M, Parascombrops yamanouei n.sp., NMMB-P 15571, paratype, 102 mm SL; N, Caraibops trispinosus, CAS 61040, 97.5 mm SL; O, Kaperangus microlepis, CAS 222834, 78.5 mm SL; P, Synagrops japonicus, MNHN 1998-0993, 84 mm SL; Q, Ƒerilus pseudomicrolpis, CAS 61164, 91 mm SL.
FIGURE 14A–Y in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 14A–Y. Otoliths (inner faces and representative ventral views): A–E, Parascombrops argyreus (14A–B = MNHN 1998-1041, 14C = ZMUC P44272, 14D–E = CAS 90647); F–G, Parascombrops analis (MNHN 1998-1056); H–I, Parascombrops mochizukii n.sp. (NMMB-P 3887); J–N, Parascombrops glossodon n.sp. (14J–K = NMMB-P 23095, 14L = USNM 436684, 14M–N = BMNH 1986.10.9.79); O–R, Parascombrops nakayamai n.sp. (14O = MNHN 1998-0993, 14P = BSKU 102255, 14Q–R = NMMB-P 23092); S–U, Parascombrops ohei n.sp. (14S = BSKU 72614, 14T–U = BSKU 116498); V–Y, Parascombrops parvidens n.sp. (14V–W = BSKU 98883, 14X–Y = BSKU 98886).
FIGURE 11A–E in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 11A–E. Preopercular serrations: A, inner and outer edges serrated, serrations of outer edge extending onto preopercle as radial crests (Kaperangus microlepis, CAS 222834, 78.5 mm SL); B, inner and outer edges serrated, serration of outer edge extending onto preopercle as radial crests, and small spike present at upper central portion (Parascombrops analis, MNHN 1998-1056, 82 mm SL); C, inner and outer edges serrated, no serrations on preopercle, two ridges on central portion of preopercle (Parascombrops argyreus, BPBM 23839, 118 mm SL); D, inner and outer edges serrated, serrations of outer edge extending onto preopercle as radial crests, and one ridge on central portion of preopercle (Parascombrops parvidens n.sp., BSKU 15221, paratype, 74 mm SL); E, inner and outer edges serrated, no serrations or ridges on preopercle (Parascombrops philippinensis, BMNH 1879.5.14.167, lectotype, 70 mm SL). Right images show characters enhanced by drawing and annotation.
FIGURE 10A–H in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 10A–H. Dentition of vomer, palatines and ectopterygoids: A, Parascombrops pellucidus, USNM 392727, 84 mm SL; B, Parascombrops philippinensis, BMNH 2016.9.14.1, paralectotype, 74 mm SL; C, Parascombrops serratospinosus, BSKU 16827, 67 mm SL; 10D, Parascombrops spinosus, MNHN 1989-0411, 82 mm SL; E, Kaperangus microlepis, CAS 222834, 96.5 mm SL; F, Parascombrops yamanouei n.sp., NMMB-P 11343, paratype, 82 mm SL; G, Caraibops trispinosus, CAS 61040, 97.5 mm SL; H, Ƒerilus pseudomicrolepis, CAS 61164, 91 mm SL.
FIGURE 7A–Q in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 7A–Q. Head profiles of Parascombrops, Caraibops, Kaperangus, Synagrops and Ƒerilus pseudomicrolepis. A, Parascombrops analis, MNHN 1998-1056, 82 mm SL; B, Parascombrops argyreus, ZMUC P44272, 122 mm SL; C, Parascombrops glossodon n.sp., BMNH 1986.10.9.79.85, paratype, 4 mm SL; D, Parascombrops madagascariensis n.sp., MNHN 2008-1769, holotype, 68.5 mm SL; E, Parascombrops mochizukii n.sp., NMMBA-P 782, holotype, 166 mm SL; F, Parascombrops nakayamai n.sp., BSKU 102255, holotype, SL 97 mm; G, Parascombrops ohei n.sp., BSKU 72614, holotype, 75 mm SL; H, Parascombrops parvidens n.sp., BSKU 98886, holotype, 57.5 mm SL; I, Parascombrops pellucidus, USNM 44428, 64.5 mm SL; J, Parascombrops philippinensis, ZMA 112.453 (paralectotype of P. malyanus), 52 mm SL; K, Parascombrops serratospinosus, CAS 238016, 67.5 mm SL; L, Parascombrops spinosus, USNM 395230, 95.5 mm SL; M, Parascombrops yamanouei n.sp., NMMB-P 23097, paratype, 70 mm SL; N, Caraibops trispinosus, CAS 61040, 97.5 mm SL; O, Kaperangus microlepis, CAS 222834, 78.5 mm SL; P, Synagrops japonicus, ZMUC P44324, 159 mm SL; 7Q, Ƒerilus pseudomicrolpis, CAS 61164, 91 mm SL.
FIGURE 8A–F in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 8A–F. Dentition and shape of maxillary: A, long, fang-like teeth on symphyses and laterally on dentary, maxillary with straight posterior edge (Parascombrops philippinensis, BMNH 1879.5.14.167, lectotype, 70 mm SL); B, no long teeth on premaxillary or dentary, maxillary with straight posterior edge (Parascombrops parvidens n.sp., BSKU 98888, paratype, 62 mm SL); C, D, mouth cavity with teeth patches on vomer, palatine and ectopterygoid (8c = Parascombrops nakayamai n.sp., CAS 34870, 81.5 mm SL; 8D = Parascombrops serratospinosus, AMS I.22822-017, 72 mm SL); E, maxillary with ventroposterior extension and concave posterior edge (Parascombrops argyreus, BPBM 23839, 118 mm SL); F, teeth patch on tongue (Parascombrops glossodon n.sp., ZMMGU 23812, paratype, 80 mm SL).
FIGURE 3A–J in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 3A–J. Predorsal formulae, pterygiophore / interneural patterns and epural patterns: A, pterygiophore pattern with last D1 spine and its pterygiophore aligned, last D1 pterygiophore unmodified with short posterior portion of proximal-medial element, short distal element, and interneural gap absent (Kaperangus microlepis n.gen., CAS 223183, 110 mm SL); B, pterygiophore pattern with anteriorly shifted last D1 spine, with strongly inclined last D1 pterygiophore with extended posterior portion of proximal-medial element and elongated distal element, interneural gap present (Parascombrops nakayamai n.sp., BSKU 102255, holotype, SL 97 mm); C, predorsal formula 0/0/0+2/ (Synagrops japonicus, MNHN 1998-0993, 84 mm SL); D, predorsal formula /0+0/0+2/ (Parascombrops glossodon n.sp., USNM 99335, paratype, 64.5 mm SL); E, predorsal formula /0/0+2/ (Kaperangus microlepis n.gen., CAS 222834, 78.5 mm SL); F, epural pattern with long, straight 1st epural (Kaperangus microlepis n.gen., CAS 222834, 78.5 mm SL); G–H, epural pattern with short, curved 1st epural (G = Parascombrops ohei n.sp., BSKU 72614, holotype, 75 mm SL; H = Synagrops bellus, USNM 436688, SL 103 mm); I, 2nd and 3rd anal pterygiopjore separated by 2nd haemal spine (Kaperangus microlepis n.gen., CAS 222834, 78.5 mm SL); J, 2nd and 3rd anal-fin pterygiophore corresponding to 1st interhaemal space (Parascombrops pellucidus, USNM 44428, 62.5 mm SL). Abbreviations: DE, distal element of last D1 pterygiophore; ppMPE, posterior portion of proximal-medial element of last D1 pterygiophore.
FIGURE 9A–H in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 9A–H. Dentition on vomer, palatines and ectopterygoids: A, Parascombrops analis, AMS I.20919-038, 107.5 mm SL; B, Parascombrops argyreus (B1 = view of mouth roof, BPBM 23829, 119 mm SL; B2 = lateral close-up view of left palatine and ectopterygoid, MNHN 1998-1041, 92 mm SL); C, Parascombrops glossodon n.sp., AMS I.21835-026, paratype, 72 mm SL; D, Parascombrops madagascariensis n.sp., MNHN 1988-1443, paratype, 62 mm SL; E, Parascombrops mochizukii n.sp., NMMB-P 3887, paratype, 98 mm SL; F, Parascombrops nakayamai n.sp., AMS I.2449-008, paratype, 93.5 mm SL; G, Parascombrops parvidens n.sp., BSKU 99887, paratype, 58 mm SL; H, Parascombrops ohei n.sp., BSKU 063370, paratype, 87 mm SL.
FIGURE 6A–H in Reappraisal of Synagrops Günther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae)
FIGURE 6A–H. Spine serrations in Parascombrops: A, C, serrated first dorsal fin spine (6A = Parascombrops serratospinosus, AMS I.22822-017, 72 mm SL; 6C = Parascombrops spinosus, USNM 395230, 95.5 mm SL); B, serrated second dorsal fin spine (Parascombrops serratospinosus, AMS I.22822-017, 72 mm SL); D, second dorsal fin spines smooth (Parascombrops spinosus, USNM 395230, 95.5 mm SL); E, serrated spine in pelvic fin (Parascombrops madagascariensis n.sp., MNHN 2008-1769, holotype, 68.5 mm SL); F, in anal fin spine serrated (Parascombrops serratospinosus, AMS I.22822- 017, 72 mm SL); G, anal fin with two spines, one unbranched soft ray and six branched soft rays (Parascombrops ohei n.sp., BSKU 72614, holotype, 75 mm SL); H, anal fin with three spines and six soft rays (Parascombrops analis, MNHN 1998-1056, 82 mm SL).
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