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Figure 2 from: Karam-Gemael M, Izzo T, Chagas-Jr A (2018) Why be red listed? Threatened Myriapoda species in Brazil with implications for their conservation. In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 255-269. https://doi.org/10.3897/zookeys.741.21971

Figure 2 Myriapoda species assessement by the current Brazilian Red List. Myriapoda threatened species according to the 2014 Brazilian Red List, which follows IUCN classification categories (CR = Critically endangered, EN = Endangered, VU = Vulnerable, NT = Near threatened, LC = Least concern, DD = Data deficient). Dashed red line indicates threatened categories.

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Figure 1 from: Karam-Gemael M, Izzo T, Chagas-Jr A (2018) Why be red listed? Threatened Myriapoda species in Brazil with implications for their conservation. In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 255-269. https://doi.org/10.3897/zookeys.741.21971

Figure 1 Myriapoda in the IUCN and Brazilian red lists. Timeline of Myriapoda species included in the IUCN Red Lists of Threatened Species (above the red line), and in the Brazilian Red Lists of Threatened Species (below the red line), highlighting the first myriapods listed and the current number of species listed.

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Figure 17 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 17 Bristles and scales on the distal tips of the lips on hypopharynx of Lithobiidae. A Lithobius (Lithobius) cyrtopus B Lithobius (Lithobius) pelidnus C Lithobius (Lithobius) validus D Eupolybothrus (Eupolybothrus) grossipes E Disphaerobius loricatus; scales F Lithobius (Lithobius) forficatus. Scale bars: 20 µm.

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Figure 4 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 4 Epipharyngeal structures of Lithobius. A Lithobius (Lithobius) macilentus; transverse bulge with a smooth surface; two rows of epidermal glandular shafts at the lateral border of the transverse bulge B Lithobius (Lithobius) lucifugus; distal transverse bulge with longitudinal striae C Lithobius (Lithobius) lucifugus; proximal transverse bulge with longitudinal striae and pores (arrow) D Lithobius (Lithobius) fagei; longitudinal striae on the lateral part of the transverse bulge (arrow); continuous branching bristle band from the distal bar to the lateral flap at the margin of the transverse bulge E Lithobius (Lithobius) cyrtopus; distinct break of branching bristle band from the distal bar to the lateral flap F Lithobius (Monotarsobius) crassipes; distinct break of branching bristle band from the distal bar to the lateral flap. bu – transverse bulge, gl – 'bottle-shaped' epidermal glandular shafts.

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Figure 7 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 7 Labral bristle bands on the distal bar of the epipharynx of Lithobius. A Lithobius (Lithobius) lucifugus; transition of simple to branching bristles from medial to lateral with a few rows of branching bristles B Lithobius (Lithobius) peregrinus; transition of simple to branching bristles from medial to lateral with several rows of branching bristles C Lithobius (Lithobius) erythrocephalus; detail of the transition of simple to branching bristles from medial to lateral D Lithobius (Lithobius) lucifugus; simple bristles on the medial part of the distal bar (top is medial).

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Figure 9 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 9 Details of branching bristles on the lateral flap on the distal bar of the epipharynx of Lithobius. A–B Lithobius (Lithobius) fagei A bristles changing from plumose laterally to 'fan-shaped' medially B detail of 9A C Lithobius (Monotarsobius) aeruginosus; bristles changing from plumose laterally to 'fan-shaped' medially D Lithobius (Lithobius) peregrinus; plumose bristles only E Lithobius (Lithobius) cyrtopus; 'fan-shaped' bristles only F Disphaerobius loricatus; simple bristles only (top is medial).

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Figure 13 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 13 Examples of number and shape of the hypopharyngeal spines and surrounding pores of Lithobiidae. A Eupolybothrus (Eupolybothrus) grossipes; several tapering spines with trifurcate spines (asterisks) in between and a continuous transition to the tuft area (arrow) B Lithobius (Lithobius) agilis; few short tapering spines; several single pores C Lithobius (Lithobius) muticus; long and tapering spines; distinct break (arrow) between hypopharyngeal spine field and branching bristles of tuft D Lithobius (Lithobius) validus; hypopharyngeal spines surrounded by cluster of up to six pores (arrow) E Lithobius (Lithobius) cyrtopus; flattened and ridged spines F Lithobius (Lithobius) castaneus; detail of a long tapering spine close to a single pore (arrow). hsp – hypopharyngeal spine field, tu – tuft of bristles.

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Figure 1 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 1 Multifocus light-micrographs and SEM-photographs of peristomatic structures in Lithobiidae. A Epipharynx of Lithobius (Lithobius) validus; posterior view (top is ventral) B Hypopharynx of Lithobius (Lithobius) carinatus; anterior view (top is dorsal) C Epipharynx of Eupolybothrus (Eupolybothrus) grossipes; posterior view (top is ventral) D Hypopharynx of Lithobius (Lithobius) forficatus; anterodorsal view (left ventrolateral bar broken). bdb – labral bristles on distal bar, blf – labral bristles on labral flap, bsc – 'button-shaped' sensilla, bu – single transverse bulge, bud – distal transverse bulge, bup – proximal transverse bulge, db – distal bar, gl – 'bottle-shaped' epidermal glandular shafts, hb – hypopharyngeal bar, hsp – hypopharyngeal spine field, lf – lateral flap, lsp – lateral spine field, lmc – paired lips forming median crest, mo – mouth opening, msc – median sensilla cluster, msp – median spine field, nsc – cluster of 'nipple-shaped' sensilla, pb – proximal bar, pp – pharyngeal plate, smc – spines flanking median crest, tp – tooth plate, tu – tuft of bristles, tub – tubercles on distal bar, vlb – ventrolateral bar. Scale bars: 200 µm.

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Figure 16 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 16 Examples of bristles transitioning in shape and length along the median crest margin on hypopharynx of Lithobius. A–B Lithobius (Lithobius) forficatus B 'button-shaped' sensilla between branching bristles on the distal part of the lips C Lithobius (Lithobius) pyrenaicus D Lithobius (Lithobius) erythrocephalus; Inset: detail of brush-tufts surrounding 'button-shaped' sensilla E Lithobius (Lithobius) pelidnus F Lithobius (Lithobius) carinatus.

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Figure 11 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 11 'Nipple-shaped' sensilla on pharyngeal plate and hypopharyngeal spines of hypopharynx of Lithobius. A Lithobius (Lithobius) dentatus; pairwise hypopharyngeal spine fields laterally to pharyngeal plate; transverse line of several 'nipple-shaped' sensilla B Lithobius (Monotarsobius) aeruginosus; transverse line of a few 'nipple-shaped' sensilla on the pharyngeal plate; arrow indicates a pore C Lithobius (Lithobius) pyrenaicus; 'nipple-shaped' sensilla arranged in a zig-zag-pattern D Lithobius (Lithobius) validus; several 'nipple-shaped' sensilla arranged in a transverse line with some offset sensilla E–F Lithobius (Lithobius) forficatus E clustered 'nipple-shaped' sensilla F high magnification of 'nipple-shaped' sensilla from Fig. 11E. hsp – hypopharyngeal spine field, mo – mouth opening, nsc – cluster of 'nipple-shaped' sensilla, pp – pharyngeal plate.

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Figure 12 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 12 Shapes of branching bristles forming a tuft distally to the pharyngeal plate of the hypopharynx of Lithobiidae. A Lithobius (Lithobius) forficatus; ramified branching bristles with a roundish shaft and hypopharyngeal spines laterally to pharyngeal plate (top is dorsal) B Lithobius (Lithobius) calcaratus; close-up of 'fan-shaped' and flattened branching bristles (top is medial) C Eupolybothrus (Eupolybothrus) grossipes; ramified and more flattened branching bristles (top is dorsal) D Lithobius (Lithobius) latro; ramified and flat branching bristles (top is medial). hsp – hypopharyngeal spine field.

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Figure 5 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 5 Epipharyngeal structures of Lithobius. A Lithobius (Lithobius) pyrenaicus; one row of 'bottle-shaped' epidermal glandular shafts; laterally narrowing transverse bulge B–C Lithobius (Lithobius) validus B two rows of 'bottle-shaped' epidermal glandular shafts C pore of an epidermal glandular shaft (arrow) D Lithobius (Ezembius) electus; broad median spine field with several rows of branching bristles and a subequal width medially and laterally. bu – transverse bulge, gl – 'bottle-shaped' epidermal glandular shafts.

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Figure 6 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 6 Details of spines from the median spine field on the epipharynx of Lithobius. A Lithobius (Monotarsobius) aeruginosus; narrow median spine field with a few rows of branching spines B Lithobius (Lithobius) macilentus C Lithobius (Lithobius) peregrinus D Lithobius (Lithobius) tricuspis. msp – median spine field. Scale bars: 5 µm.

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Figure 8 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 8 Details of branching bristles on the outer row of the labral bristle bands on the distal bar of the epipharynx of Lithobius. A–B 'hassock-like' branching bristles with a broad base A Lithobius (Lithobius) mutabilis B Lithobius (Ezembius) electus C Lithobius (Lithobius) pyrenaicus; 'hassock-like' branching bristles with a narrow base D Lithobius (Lithobius) validus; 'palmleaf-like' bristles E Lithobius (Sigibius) trebinjanus; 'comb-like' bristles (top is medial).

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Figure 15 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 15 Examples of spines and bristles flanking the median crest margins of hypopharynx of Lithobius. A–C flattened spines with a transition to branching bristles on the inner rows A Lithobius (Lithobius) pelidnus B Lithobius (Monotarsobius) franciscorum C Lithobius (Lithobius) muticus; Inset: detail of structured surface of flattened spines D–E continuously branching bristles flanking the median crest D Lithobius (Sigibius) microps; stout and short branching bristles E Lithobius (Lithobius) piceus; slender and long branching bristles F Lithobius (Lithobius) forficatus; flattened spines flanking median crest margin. A–B, D–E top is medial; C top is ventral; F top is dorsal.

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Figure 10 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 10 Median sensilla cluster and lateral spine fields on the epipharynx of Lithobiidae. A Lithobius (Lithobius) validus; sensilla cluster arranged in an offset pattern; spine field arranged as single oblique row; Inset: Lithobius (Monotarsobius) aeruginosus; sensilla cluster arranged in line B Lithobius (Lithobius) tenebrosus; sensilla cluster arranged symmetrically; spine field arranged as single oblique row with trifurcate spines (arrow) C Neolithobius aztecus; sensilla cluster arranged in an offset pattern; spine field arranged as single oblique row D Eupolybothrus (Eupolybothrus) grossipes; sensilla cluster arranged in an offset-pattern and strongly overlapping with lateral spine field proximolaterally; spine field arranged as single oblique row E Lithobius (Lithobius) pelidnus; spine field arranged as two rows (tendency of clustering) with bi- or trifurcate spines (arrows) and pores (asterisks) F Lithobius (Monotarsobius) curtipes; spine field arranged as single oblique row G Lithobius (Monotarsobius) aeruginosus; spine field arranged as single oblique row H Lithobius (Monotarsobius) aeruginosus; tubercles on distal bar. lsp – lateral spine field, msc – median sensilla cluster.

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Figure 14 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 14 Examples of 'button-shaped' sensilla on the lips of hypopharynx of Lithobiidae. A Lithobius (Lithobius) forficatus; proximal part of lips forming median crest with cluster of 'button-shaped' sensilla; flattened spines flanking median crest margin B Eupolybothrus (Eupolybothrus) grossipes; left lip with cluster of 'button-shaped' sensilla C Lithobius (Lithobius) validus D Lithobius (Sigibius) burzenlandicus E Lithobius (Lithobius) muticus F Lithobius (Lithobius) carinatus. bsc – 'button-shaped' sensilla, smc – spines flanking median crest, tu – tuft of bristles. A top is dorsal; B–F top is medial.

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Figure 2 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 2 Details of transverse bulge, 'bottle-shaped' epidermal glandular shafts and median spine field of the epipharynx of Lithobiidae. A Disphaerobius loricatus; no transverse bulge; consistently two rows of 'bottle-shaped' epidermal glandular shafts; narrow and slightly medially widening median spine field B Lithobius (Lithobius) pyrenaicus; parallel aligned margins of a single transverse bulge; one row of 'bottle-shaped' epidermal glandular shafts; rhomboid and medially widening median spine field C Lithobius (Lithobius) fagei; single transverse bulge with parallel margins; more than one row of 'bottle-shaped' epidermal glandular shafts laterally; laterally widening median spine field D Lithobius (Sigibius) microps; single transverse bulge with non-parallel margins; subequal width of median spine field E Lithobius (Lithobius) mutabilis; single transverse bulge with non-parallel margins; one row of 'bottle-shaped' epidermal glandular shafts; subequal width of median spine field F Lithobius (Monotarsobius) aeruginosus; single transverse bulge with non-parallel margins; one row of 'bottle-shaped' epidermal glandular shafts; subequal width of median spine field. bu – transverse bulge, gl – 'bottle-shaped' epidermal glandular shafts, msp – median spine field.

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Figure 3 from: Ganske A-S, Edgecombe GD, Akkari N (2018) The peristomatic structures as a source of systematic characters in the genus Lithobius Leach, 1814 (Myriapoda, Chilopoda). In: Stoev P, Edgecombe GD (Eds) Proceedings of the 17th International Congress of Myriapodology, Krabi, Thailand. ZooKeys 741: 49-75. https://doi.org/10.3897/zookeys.741.21706

Figure 3 Details of transverse bulge, 'bottle-shaped' epidermal glandular shafts and median spine field of the epipharynx of Lithobiidae. A Lithobius (Lithobius) macilentus; single transverse bulge with smooth surface (see Fig. 4A) and non-parallel aligned margins; one row of 'bottle-shaped' epidermal glandular shafts medially with a transition to two rows laterally (see Fig. 4A); weak transverse furrow distally to the transverse bulge (arrow); laterally widening median spine field B Lithobius (Lithobius) piceus; weak transverse furrow (arrow) distally to the single transverse bulge (non-parallel margins); irregular two rows of 'bottle-shaped' epidermal glandular shafts; subequal width medially and laterally of median spine field C Lithobius (Lithobius) lucifugus; distal and proximal transverse bulges with surface striation (see Fig. 4B–C) D Eupolybothrus (Eupolybothrus) grossipes; distal and proximal transverse bulges; medially widening median spine field. bu – transverse bulge, bud – distal transverse bulge, bup – proximal transverse bulge, gl – 'bottle-shaped' epidermal glandular shafts, msp – median spine field.

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Figure 4 from: Chagas-Jr A, Bichuette ME (2018) Synopsis of centipedes in Brazilian caves (Arthropoda, Myriapoda, Chilopoda), a hidden diversity to be protected. ZooKeys 737: 13-56. https://doi.org/10.3897/zookeys.737.20307

Figure 4 Outcrops and caves from Brazil with centipede records. A Limestone outcrops with pastures from Peruaçu region, Minas Gerais state. Cerrado (savannah-like vegetation) interspersed with Caatinga (xerophitic vegetation) (M.E. Bichuette) B Limestone outcrops with pastures from Serra do Ramalho, Bahia state. Caatinga (xerophitic vegetation) (M.E. Bichuette) C Atlantic Rainforest and limestone outcrops from Alto Ribeira, São Paulo state (M.E. Bichuette) D Alto Ribeira cave entrance, São Paulo state (M.E. Bichuette) E Quartizite outcrops with "Campo rupestre" vegetation from Diamantina, Minas Gerais state (M.E. Bichuette) F Sandstone cave from Igatu, Chapada Diamantina, Bahia state (E.C. Igual) G Sandstone outcrop from Igatu, Chapada Diamantina, Bahia state (J.E. Gallão) H Granite outcrop and Atlantic Rainforest from Florianópolis, Santa Catarina state (J.E. Gallão) I Iron ore outcrop with Cerrado from Caeté, Minas Gerais state (M.E. Bichuette) J Iron ore cave from Caeté, Minas Gerais state (M.E. Bichuette) K, L Iron ore caves and Canga vegetation from Canaã dos Carajás, Pará state (A. Coelho).

opencc-by-4.0Apr 2018View details →

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Last verified 2026-04-30Open record

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Last verified 2026-04-29Open record

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Last verified 2026-04-29Open record