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Figure 13. A–C, Unio bruguierianus. A in Species boundaries, geographic distribution and evolutionary history of the Western Palaearctic freshwater mussels Unio (Bivalvia: Unionidae)

Figure 13. A–C, Unio bruguierianus. A, Pinios River, Greece. B, Axios River, Greece. C, Pinios River, Greece. D–E, Unio ionicus. D, River at Lake Lysimacheia, Greece. E, Perroi i Bistrices, Albania. F–H, Unio crassus. F, Sperchios River, Greece. G, H, Aliakmonas River, Greece. Scale bar 2 cm.

Distribution. Balabac, Ramos, and Bugsuk Is, Palawan region, Phillipines. In the late 1990s, a small stock of eight Balabac Chevrotains escaped from their enclosures on Calauit Island, a small island off the coast of Busuanga Island, north of Palawan, where the species had been maintained and bred since 1982. These animals were reported to have increased to at least 21 free-living individuals by 2006. Apparently the species was also introduced to the larger island of Palawan. No recent records confirm that it is still extant there, although unconfirmed reports suggest that it may survive in southern Palawan. in Tragulidae

Distribution. Balabac, Ramos, and Bugsuk Is, Palawan region, Phillipines. In the late 1990s, a small stock of eight Balabac Chevrotains escaped from their enclosures on Calauit Island, a small island off the coast of Busuanga Island, north of Palawan, where the species had been maintained and bred since 1982. These animals were reported to have increased to at least 21 free-living individuals by 2006. Apparently the species was also introduced to the larger island of Palawan. No recent records confirm that it is still extant there, although unconfirmed reports suggest that it may survive in southern Palawan.

Distribution. Java. Several biogeographic reviews failed to list the genus for Bali, an island directly east of Java with strong biogeographic affinities to the latter. A sighting was, however, reported from Bali Barat National Park, in a birdwatching trip report. Given the live-animal trade of this species on Java, further records are needed to confirm whether or not there is a native population of the genus on Bali. If there is, biogeographic considerations suggest it would be most closely related to the Javan population and quite probably conspecific. in Tragulidae

Distribution. Java. Several biogeographic reviews failed to list the genus for Bali, an island directly east of Java with strong biogeographic affinities to the latter. A sighting was, however, reported from Bali Barat National Park, in a birdwatching trip report. Given the live-animal trade of this species on Java, further records are needed to confirm whether or not there is a native population of the genus on Bali. If there is, biogeographic considerations suggest it would be most closely related to the Javan population and quite probably conspecific.

Subspecies and Distribution. T.n.napuF.Cuvier,1822—SMyanmar,Thai/MalayPeninsula,islandsoffWMalayPeninsula(Langkawi&Pangkor),Borneo,SSumatra,BangkaI,islandsoffBorneo(Laut&Serasan). T.n.bangue:Chasen&Kloss,1931—BanggiIandBalembanganI,offNBorneo. T.n.bunguranensisMiller,1901—NatunaIs(=Bunguran),oftWBorneo. T.n.neubronneriSody,1931—NSumatra. T.n.nmiasisLyon,1916—NiasI,offWSumatra. T.n.rufulusMiller,1900—TiomanI,offEMalayPeninsula,RiauandLinggaArchipelagos. T. n. terutus Thomas & Wroughton, 1909 — Terutau I, off W Malay Peninsula. The species was recently reconfirmed for Singapore. Maps that include Vietnam, Cambodia, and Laos in the distribution range are based on the earlier assumption that 7. versicolor was a subspecies of 1. napu. Subsequent studies have indicated that 7. versicolor is a distinct species, and that the range of 1. napu therefore does not extend into Cambodia, Laos, and Vietnam. The northern limit on the Thai-Malay peninsula is not well defined. Specimens of 1. napu have been collected from as far north as Bankachon in southern Myanmar (10° 08" N), but despite fairly intensive camera-trapping in Kui Buri National Park, Thailand (12° N), 7. napu has not been photographed there. At the northern margin ofits range, it is generally rare. It has been reported, for example, that during the flooding of the Chiew Larn Reservoir (Surat Thani Province; about 9° N, 98° 45' E), only six 7. napu were rescued compared with 172 71. kanchil. This area is the transition zone from wetter evergreen forest to drier deciduous types, and it might be that 7° napu is not well adapted to the drier forest types towards the northern limit ofits range. There are unconfirmed reports of the species on Java, where it may have been confused with one of the two color morphs of 7. javanicus. As explained in the Taxonomy section, the subspecific status of the populations of several islands remains unclear. in Tragulidae

Subspecies and Distribution. T.n.napuF.Cuvier,1822—SMyanmar,Thai/MalayPeninsula,islandsoffWMalayPeninsula(Langkawi&Pangkor),Borneo,SSumatra,BangkaI,islandsoffBorneo(Laut&Serasan). T.n.bangue:Chasen&Kloss,1931—BanggiIandBalembanganI,offNBorneo. T.n.bunguranensisMiller,1901—NatunaIs(=Bunguran),oftWBorneo. T.n.neubronneriSody,1931—NSumatra. T.n.nmiasisLyon,1916—NiasI,offWSumatra. T.n.rufulusMiller,1900—TiomanI,offEMalayPeninsula,RiauandLinggaArchipelagos. T. n. terutus Thomas & Wroughton, 1909 — Terutau I, off W Malay Peninsula. The species was recently reconfirmed for Singapore. Maps that include Vietnam, Cambodia, and Laos in the distribution range are based on the earlier assumption that 7. versicolor was a subspecies of 1. napu. Subsequent studies have indicated that 7. versicolor is a distinct species, and that the range of 1. napu therefore does not extend into Cambodia, Laos, and Vietnam. The northern limit on the Thai-Malay peninsula is not well defined. Specimens of 1. napu have been collected from as far north as Bankachon in southern Myanmar (10° 08" N), but despite fairly intensive camera-trapping in Kui Buri National Park, Thailand (12° N), 7. napu has not been photographed there. At the northern margin ofits range, it is generally rare. It has been reported, for example, that during the flooding of the Chiew Larn Reservoir (Surat Thani Province; about 9° N, 98° 45' E), only six 7. napu were rescued compared with 172 71. kanchil. This area is the transition zone from wetter evergreen forest to drier deciduous types, and it might be that 7° napu is not well adapted to the drier forest types towards the northern limit ofits range. There are unconfirmed reports of the species on Java, where it may have been confused with one of the two color morphs of 7. javanicus. As explained in the Taxonomy section, the subspecific status of the populations of several islands remains unclear.

Distribution. If still extant, the species would likely occur in Vietnam, and possibly in neighboring Cambodia and Laos. The true distribution of the species is unknown, because there has been very little awareness of the species. Surveys have not sought it, because many have assumed that only one species, Tragulus kanchil, occurs in most of Indochina. in Tragulidae

Distribution. If still extant, the species would likely occur in Vietnam, and possibly in neighboring Cambodia and Laos. The true distribution of the species is unknown, because there has been very little awareness of the species. Surveys have not sought it, because many have assumed that only one species, Tragulus kanchil, occurs in most of Indochina.

Subspecies and Distribution. S. s. scrofa Linnaeus, 1758 — W Europe, from Denmark, Germany, Poland, and Czech Republic to N Italy and N Iberian Peninsula; possibly also Albania. The taxonomic status of animals in Austria, Switzerland, Slovenia, and Slovakia is unclear but presumably these populations are included in scrofa, as are the populations of Sweden, Finland, and the Baltic states. However, restocking of once depleted populations, for example in Italy, has likely involved the introduction and mixing of this subspecies with other subspecies, such as attila. S. s. affinis Gray, 1847 — S India and Sri Lanka. S. s. algirus Loche, 1867 — Tunisia, Algeria, and Morocco, on the coastal side of the mountains or in the low montane areas. S. s. attila Thomas, 1912 — Hungary, Ukraine, C & S Belarus, Romania, Moldova, and S Russia towards the N flank of the Caucasus, but not including the Transcaucasian countries of Georgia, Armenia, and Azerbaijan. The range possibly extends as far S as the Mesopotamian Delta in Iraq, in which case it would likely include W & SW Iran, and possibly E Turkey and Syria, where it borders with lybicus. Such a range could not be easily reconciled with a statement by Groves that "the difference between pigs from N and S of the Caucasus is quite striking; Transcaucasian boars are certainly not attila." This subspecies may also extend into C Asia and include Kazakhstan, Uzbekistan, and Turkmenistan, but no data exist to support this. S. s. baeticus Thomas, 1912 — originally described from Coto Donana, S Spain, and later merged with meridionalis; also S Portugal. Unless evidence is found that these Italian and Iberian populations are the relics of a much larger formerly contiguous range, this subspecies should be kept as distinct. S. s. coreanus Heude, 1897 — Korean Peninsula. S. s. eristatus Wagner, 1839 — Himalayas S to C India and E to Indochina (N of the Kra Isthmus). S. s. davidi Groves, 1981 — the arid zone from E Iran to Gujarat, including Pakistan and NW India, and perhaps N to Tajikistan. S. s. leucomystax Temminck, 1842 — main Is ofJapan (Honshu, Shikoku, Kyushu, Nakadori, Hiburijima, Tojima, Kushima, and other smaller Is). S. s. lybicus Gray, 1868 — Bulgaria, Greece, Turkey, Syria, Jordan, Israel, Palestine, in the past also in Lybia, and Egypt. The former Yugoslavia was included in its range, which would suggest that now Slovenia, Serbia, Croatia, Bosnia and Herzegovina, Montenegro, and Kosovo are within the range of this subspecies, although the exact boundaries are unclear. Pigs from Albania have been assigned to S. s. scrofa. S. s. majori De Beaux & Festa, 1927 — C & S Italian Peninsula. S. s. menidionalis Forsyth Major, 1882 — Corsica and Sardinia, with the proviso that the two populations are very likely to be introduced or feral. S. s. moupinensis Milne-Edwards, 1871 — China, S to Vietnam and W to Sichuan. S. s. nigripes Blanford, 1875 — the flanks of the Tianshan mountains in Kyrgyzstan and NW China (Xinjiang). An animal photographed in NE Iran (Golestan) looked like this subspecies. S. s. nukiuanus Kuroda, 1924 — Iriomote, Ishigaki, Okinawa, Tokunoshima, Amamioshima, and Kakerome Is in the Ryukyu chain in extreme S Japan, though some of these populations have hybridized with introduced domesticates. S. s. sibiricus Staffe, 1922 — Mongolia and Transbaikal (S & E of Lake Baikal). S. s. tawvanus Swinhoe, 1863 — Taiwan. S. s. ussuricus Heude, 1888 — far E Russia and the Manchurian region (China). Korean populations were previously included in this subspecies, but based on new evidence, the Korean taxon seems more similar to moupinensis. S. s. vittatus Boie, 1828 — Malay Peninsula, S of the Isthmus of Kra, the offshore islands of Terutai and Langkawi, Sumatra, Riau Archipelago, Java, Bali, and a range of smaller islands around these, including Babi, Bakong, Batam, Bawean, Bengkalis, Bintan, Bulan, Bunguran, Cuyo, Deli, Durian, Enggano, Galang, Jambongan, Karimon (Riau Is), Kundur, Lagong, Laut, Lingga, Lingung, Mapor, Moro Kecil, North Pagai, Nias, Panaitan, Payong, Penang, Pinie, Rupat, Siantan, Siberut, Simeulue, Singkep, Sugi, Sugi Bawa, Telibon, Tinggi, Tuangku, and the Tambelan Is. This species was originally present from the British Is in the extreme W, through Eurasia from S Scandinavia to S Siberia, extending as far E as Korea and Japan, and SE into some of the Sunda Is and Taiwan. In the S the species ranged along the Nile Valley to Khartoum, and N of the Sahara in Africa, more orless following the continental coasts of S, E, and SE Asia. Within this range it was absent only from extremely dry deserts, e.g. the driest regions of Mongolia and in China W of Sichuan; and alpine zones, such as the high altitudes of Pamir and Tien Shan. In recent centuries, the range of S. scrofa has changed dramatically because of hunting and changes in available habitat. The species disappeared from the British Is in the 17" century, from Denmark in the 19" century, and was greatly reduced in range and numbers in the 20" century from areas as distant as Tunisia, Sudan, Germany, and Russia. Following these severe declines, there were some slight population recoveries in Russia, Italy, Spain, and Germany in the mid-20™ century, and natural and assisted range expansions in Denmark and Sweden. The species has also been inadvertently reintroduced in various locations in the Great Britain via escapees of mixed origin from commercial farming enterprises. Ex-S. scrofa stocks also occur as introduced feral populations in various other parts of the world, including Australia, New Zealand, the eastern Malay Archipelago, and in North, Central, and South America. In all of these areas they are now generally recognized as a major pest. in Suidae

Subspecies and Distribution. S. s. scrofa Linnaeus, 1758 — W Europe, from Denmark, Germany, Poland, and Czech Republic to N Italy and N Iberian Peninsula; possibly also Albania. The taxonomic status of animals in Austria, Switzerland, Slovenia, and Slovakia is unclear but presumably these populations are included in scrofa, as are the populations of Sweden, Finland, and the Baltic states. However, restocking of once depleted populations, for example in Italy, has likely involved the introduction and mixing of this subspecies with other subspecies, such as attila. S. s. affinis Gray, 1847 — S India and Sri Lanka. S. s. algirus Loche, 1867 — Tunisia, Algeria, and Morocco, on the coastal side of the mountains or in the low montane areas. S. s. attila Thomas, 1912 — Hungary, Ukraine, C & S Belarus, Romania, Moldova, and S Russia towards the N flank of the Caucasus, but not including the Transcaucasian countries of Georgia, Armenia, and Azerbaijan. The range possibly extends as far S as the Mesopotamian Delta in Iraq, in which case it would likely include W & SW Iran, and possibly E Turkey and Syria, where it borders with lybicus. Such a range could not be easily reconciled with a statement by Groves that "the difference between pigs from N and S of the Caucasus is quite striking; Transcaucasian boars are certainly not attila." This subspecies may also extend into C Asia and include Kazakhstan, Uzbekistan, and Turkmenistan, but no data exist to support this. S. s. baeticus Thomas, 1912 — originally described from Coto Donana, S Spain, and later merged with meridionalis; also S Portugal. Unless evidence is found that these Italian and Iberian populations are the relics of a much larger formerly contiguous range, this subspecies should be kept as distinct. S. s. coreanus Heude, 1897 — Korean Peninsula. S. s. eristatus Wagner, 1839 — Himalayas S to C India and E to Indochina (N of the Kra Isthmus). S. s. davidi Groves, 1981 — the arid zone from E Iran to Gujarat, including Pakistan and NW India, and perhaps N to Tajikistan. S. s. leucomystax Temminck, 1842 — main Is ofJapan (Honshu, Shikoku, Kyushu, Nakadori, Hiburijima, Tojima, Kushima, and other smaller Is). S. s. lybicus Gray, 1868 — Bulgaria, Greece, Turkey, Syria, Jordan, Israel, Palestine, in the past also in Lybia, and Egypt. The former Yugoslavia was included in its range, which would suggest that now Slovenia, Serbia, Croatia, Bosnia and Herzegovina, Montenegro, and Kosovo are within the range of this subspecies, although the exact boundaries are unclear. Pigs from Albania have been assigned to S. s. scrofa. S. s. majori De Beaux & Festa, 1927 — C & S Italian Peninsula. S. s. menidionalis Forsyth Major, 1882 — Corsica and Sardinia, with the proviso that the two populations are very likely to be introduced or feral. S. s. moupinensis Milne-Edwards, 1871 — China, S to Vietnam and W to Sichuan. S. s. nigripes Blanford, 1875 — the flanks of the Tianshan mountains in Kyrgyzstan and NW China (Xinjiang). An animal photographed in NE Iran (Golestan) looked like this subspecies. S. s. nukiuanus Kuroda, 1924 — Iriomote, Ishigaki, Okinawa, Tokunoshima, Amamioshima, and Kakerome Is in the Ryukyu chain in extreme S Japan, though some of these populations have hybridized with introduced domesticates. S. s. sibiricus Staffe, 1922 — Mongolia and Transbaikal (S & E of Lake Baikal). S. s. tawvanus Swinhoe, 1863 — Taiwan. S. s. ussuricus Heude, 1888 — far E Russia and the Manchurian region (China). Korean populations were previously included in this subspecies, but based on new evidence, the Korean taxon seems more similar to moupinensis. S. s. vittatus Boie, 1828 — Malay Peninsula, S of the Isthmus of Kra, the offshore islands of Terutai and Langkawi, Sumatra, Riau Archipelago, Java, Bali, and a range of smaller islands around these, including Babi, Bakong, Batam, Bawean, Bengkalis, Bintan, Bulan, Bunguran, Cuyo, Deli, Durian, Enggano, Galang, Jambongan, Karimon (Riau Is), Kundur, Lagong, Laut, Lingga, Lingung, Mapor, Moro Kecil, North Pagai, Nias, Panaitan, Payong, Penang, Pinie, Rupat, Siantan, Siberut, Simeulue, Singkep, Sugi, Sugi Bawa, Telibon, Tinggi, Tuangku, and the Tambelan Is. This species was originally present from the British Is in the extreme W, through Eurasia from S Scandinavia to S Siberia, extending as far E as Korea and Japan, and SE into some of the Sunda Is and Taiwan. In the S the species ranged along the Nile Valley to Khartoum, and N of the Sahara in Africa, more orless following the continental coasts of S, E, and SE Asia. Within this range it was absent only from extremely dry deserts, e.g. the driest regions of Mongolia and in China W of Sichuan; and alpine zones, such as the high altitudes of Pamir and Tien Shan. In recent centuries, the range of S. scrofa has changed dramatically because of hunting and changes in available habitat. The species disappeared from the British Is in the 17" century, from Denmark in the 19" century, and was greatly reduced in range and numbers in the 20" century from areas as distant as Tunisia, Sudan, Germany, and Russia. Following these severe declines, there were some slight population recoveries in Russia, Italy, Spain, and Germany in the mid-20™ century, and natural and assisted range expansions in Denmark and Sweden. The species has also been inadvertently reintroduced in various locations in the Great Britain via escapees of mixed origin from commercial farming enterprises. Ex-S. scrofa stocks also occur as introduced feral populations in various other parts of the world, including Australia, New Zealand, the eastern Malay Archipelago, and in North, Central, and South America. In all of these areas they are now generally recognized as a major pest.

Distribution. Sulawesi and adjacent Is (Buton, Kabaena, Muna, Peleng, Lembeh, and on some of the Togian Is); thought to be extinct on Selayar I. Pigs have been widely domesticated through the Indonesian archipelago and beyond. This primarily involved the Eurasian Wild Pig (S. scrofa), but also S. celebensis, the only other species of pig successfully domesticated. Mitochondrial DNA studies of the dispersion of these domesticated forms agree on three major dispersal events, two involving S. scrofa and one S. celebensis. Evidence supports an early human-mediated translocation of S. celebensis to Flores and Timor and two later, separate human-mediated dispersals of domestic pig through islands of SE Asia into Oceania. In addition to Flores and Timor, S. celebensis is also thought to occur in its domesticated form on Halmahera, Lendu, Roti, and Savur Is, and even on Simeulue and Nias Is to the W of Sumatra and far from its island of origin, Sulawesi. In the Moluccas, and possibly elsewhere in this region, introduced S. celebensis are thought to have hybridized with other introduced pigs of S. scrofa derivation, and apparent hybrids between these species are now reported to survive on a number of islands, including Salawatti, Great Kei, Dobu, Seram, Ambon, Bacan, Ternate, Morotai, and New Guinea. It is also reported that in the 19" century the sows of domestic pigs in Sulawesi frequently mated with wild animals, after which they returned to their villages. in Suidae

Distribution. Sulawesi and adjacent Is (Buton, Kabaena, Muna, Peleng, Lembeh, and on some of the Togian Is); thought to be extinct on Selayar I. Pigs have been widely domesticated through the Indonesian archipelago and beyond. This primarily involved the Eurasian Wild Pig (S. scrofa), but also S. celebensis, the only other species of pig successfully domesticated. Mitochondrial DNA studies of the dispersion of these domesticated forms agree on three major dispersal events, two involving S. scrofa and one S. celebensis. Evidence supports an early human-mediated translocation of S. celebensis to Flores and Timor and two later, separate human-mediated dispersals of domestic pig through islands of SE Asia into Oceania. In addition to Flores and Timor, S. celebensis is also thought to occur in its domesticated form on Halmahera, Lendu, Roti, and Savur Is, and even on Simeulue and Nias Is to the W of Sumatra and far from its island of origin, Sulawesi. In the Moluccas, and possibly elsewhere in this region, introduced S. celebensis are thought to have hybridized with other introduced pigs of S. scrofa derivation, and apparent hybrids between these species are now reported to survive on a number of islands, including Salawatti, Great Kei, Dobu, Seram, Ambon, Bacan, Ternate, Morotai, and New Guinea. It is also reported that in the 19" century the sows of domestic pigs in Sulawesi frequently mated with wild animals, after which they returned to their villages.

Distribution. Main tropical rainforest belt from Senegal and Guinea Bissau to NW Uganda and E DR Congo, reaching as S boundary N Angola (Cabinda); no recent records from Gambia or Chad and its presence is uncertain in S Sudan and SW Ethiopia. As a species that may range widely through gallery forests, it may also occur in adjacent countries. in Suidae

Distribution. Main tropical rainforest belt from Senegal and Guinea Bissau to NW Uganda and E DR Congo, reaching as S boundary N Angola (Cabinda); no recent records from Gambia or Chad and its presence is uncertain in S Sudan and SW Ethiopia. As a species that may range widely through gallery forests, it may also occur in adjacent countries.

Subspecies and Distribution. P.l.hassamaHeuglin,1863—EAfrica,withspecimensknownfromEthiopia,SSudan,EDRCongo,Uganda,Rwanda,Burundi,Kenya,andTanzania. P.l.koiropotamusDesmoulins,1831—lowerCongoRiver(leftbank),Angola,SDRCongo,Zambia,Malawi,Mozambique,Zimbabwe,Botswana,Swaziland,andSouthAfrica. P. l. somaliensis de Beaux, 1924 — Tana, Juba, and Shebelle rivers in NE Kenya and Somalia. The species is also present in Madagascar and the Comoro Is, but their taxonomic situation thereis still unresolved. in Suidae

Subspecies and Distribution. P.l.hassamaHeuglin,1863—EAfrica,withspecimensknownfromEthiopia,SSudan,EDRCongo,Uganda,Rwanda,Burundi,Kenya,andTanzania. P.l.koiropotamusDesmoulins,1831—lowerCongoRiver(leftbank),Angola,SDRCongo,Zambia,Malawi,Mozambique,Zimbabwe,Botswana,Swaziland,andSouthAfrica. P. l. somaliensis de Beaux, 1924 — Tana, Juba, and Shebelle rivers in NE Kenya and Somalia. The species is also present in Madagascar and the Comoro Is, but their taxonomic situation thereis still unresolved.

Subspecies and Distribution. P.c.capensisPallas,1766—SouthAfrica,includingLesothoandSwaziland. P.c.bamendaeBrauer,1913—CameroonandCentralAfricanRepublic. P.c.capillosaBrauer,1917—SEthiopia. P.c.erlanger:Neumann,1901—SSomalia. P.c.habessinicaHemprich&Ehrenberg,1832—Egypt,NSudan,Israel,SaudiArabia,andYemen. P.c.jacksoniThomas,1900—EKenya. P.c.jayakariThomas,1892—Oman. P.c.johnston:Thomas,1894—SWTanzania,Malawi,Mozambique,andZimbabwe. P.c.kerstingiMatschie,1899—TogoandBenin. P.c.mackinder:Thomas,1900—WKenya. P.c.matschietNeumann,1900—DRCongoandTanzania. P.c.pallidaThomas,1891—NSomalia. P.c.ruficepsHemprich&Ehrenberg,1832—NandWAfrica. P.c.scioanaGiglioli,1888—NEthiopia. P.c.sharicaThomas&Wroughton,1907—Chad. P.c.syriacaSchreber,1784—Syria,Lebanon,Jordan,andIsrael. P. c. welwitschii Gray, 1868 — SW Angola and Namibia. The distribution information for this species is still incomplete; the Rock Hyrax is also present in Eritrea, Niger, Nigeria, C & S Sudan, Uganda, Rwanda, Burundi, Zambia and E Botswana, but the subspecific identity of these populations still requires confirmation. in Procaviidae

Subspecies and Distribution. P.c.capensisPallas,1766—SouthAfrica,includingLesothoandSwaziland. P.c.bamendaeBrauer,1913—CameroonandCentralAfricanRepublic. P.c.capillosaBrauer,1917—SEthiopia. P.c.erlanger:Neumann,1901—SSomalia. P.c.habessinicaHemprich&Ehrenberg,1832—Egypt,NSudan,Israel,SaudiArabia,andYemen. P.c.jacksoniThomas,1900—EKenya. P.c.jayakariThomas,1892—Oman. P.c.johnston:Thomas,1894—SWTanzania,Malawi,Mozambique,andZimbabwe. P.c.kerstingiMatschie,1899—TogoandBenin. P.c.mackinder:Thomas,1900—WKenya. P.c.matschietNeumann,1900—DRCongoandTanzania. P.c.pallidaThomas,1891—NSomalia. P.c.ruficepsHemprich&Ehrenberg,1832—NandWAfrica. P.c.scioanaGiglioli,1888—NEthiopia. P.c.sharicaThomas&Wroughton,1907—Chad. P.c.syriacaSchreber,1784—Syria,Lebanon,Jordan,andIsrael. P. c. welwitschii Gray, 1868 — SW Angola and Namibia. The distribution information for this species is still incomplete; the Rock Hyrax is also present in Eritrea, Niger, Nigeria, C & S Sudan, Uganda, Rwanda, Burundi, Zambia and E Botswana, but the subspecific identity of these populations still requires confirmation.

FIGURE 1. Alona werestschagini Sinev, 1999 in New data on morphology and distribution of Alona werestschagini Sinev, 1999-the only Arcto-Alpine species of Chydoridae (Cladocera: Anomopoda) known to date

FIGURE 1. Alona werestschagini Sinev, 1999 from Russia, Tuva Republic, groundwater-fed flow-through pond in Hjut River floodplain. Parthenogenetic female. A, lateral view. B, ventral margin of valves. C, anterior group of ventral setae. D, posterior group of ventral setae. E, posteroventral angle of valves. F, head pores. G, labrum. H, postabdomen. I, postanal margin of postabdomen. J, antennule. K, antenna. Adult male. L, lateral view. M, postabdomen. N, antennule.

FIGURE 3. Alona werestschagini Sinev, 1999 in New data on morphology and distribution of Alona werestschagini Sinev, 1999-the only Arcto-Alpine species of Chydoridae (Cladocera: Anomopoda) known to date

FIGURE 3. Alona werestschagini Sinev, 1999 from Russia, Tuva Republic, groundwater-fed flow-through pond in Hjut River floodplain. Thoracic limbs of adult parthenogenetic female. A, limb I. B, setae a, b, d of limb I. C, ODL and IDL of limb I. D-E, limb II. F, exopodite of limb III. G, inner portion of limb III. H, exopodite of limb IV. I, inner portion of limb IV. J, limb V. K, limb VI. Adult male. L, limb I. M, copulatory hook of limb I.

FIGURE 5 in New data on morphology and distribution of Alona werestschagini Sinev, 1999-the only Arcto-Alpine species of Chydoridae (Cladocera: Anomopoda) known to date

FIGURE 5. Distribution of Alona werestschagini Sinev, 1999. The map is made based on the Marble Virtual Globe 2.2.0 available at https://marble.kde.org/.

FIGURE 2. Alona werestschagini Sinev, 1999 in New data on morphology and distribution of Alona werestschagini Sinev, 1999-the only Arcto-Alpine species of Chydoridae (Cladocera: Anomopoda) known to date

FIGURE 2. Alona werestschagini Sinev, 1999 from Chukotka Autonomous Area, Providenskii District, small tundra lake near Kivak settlement, Parthenogenetic female. A, head shield. B, head pores. Ephippial female. C, lateral view. Adult male. D, lateral view. E, head shield. F, head pores. G, postabdomen. H, antennule.

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dryad32/100

Distribution of 8 species of large-seeded pines and their primary animal seed-dispersers in China: match or mismatch?

Aim: The geographic distribution of plants influenced by seed dispersal, but this influence on plants that use animals as seed dispersers is often overlooked. Here, we took large-seeded pines and primary seed dispersers as examples to explore the effects of existing or potential seed dispersal on the distribution of plants based on the geographical distribution of the two trophic species, it required understanding of (a) the distribution range and distribution characteristics of each species, and (b) the overlapping of distribution areas of animals and plants to explore whether they match. Location: China Methods: To find the target species, we identified eight large-seeded pine species in China in terms of seed size and wing traits as well as four primary seed disperser species in terms of body size, seed-diet, food hoarding behavior and frequencies in existing studies. Then, we obtained species distribution information from books and literature and used ArcGIS for mapping. Finally, we analyzed the distribution relationship by overlapping the distribution areas and patterns comprehensively. Results: We identified eight species of large-seeded pines (Pinus fenzeliana, P. gerardiana, P. dabeshanensis, P. koraiensis, P. pumila, P. bungeana, P. armandii, and P. sibirica) and four species of primary seed dispersers (Nucifraga caryocatactes, Sciurus vulgaris, Tamias sibiricus, and Sciurotamias davidianus). These eight species of large-seeded pines interlaced from the Northeast to the Southwest of China along the mountains with an average altitude of 1000-2000 m, while each species of primary seed disperser had a wide distribution range that overlapped completely or partially with that of four or more species of the large-seeded pines. Not only that, our findings provided potential seed dispersers for pines that lack sufficient research on seed dispersal. Main conclusions: The distribution pattern of large-seeded pines and the primary seed dispersers was matched, we believed that reciprocal relationship promotes this distribution pattern. Our study highlights the importance of incorporating the ecological consequences of geographical distribution into reciprocal interactions between species and biodiversity conservation.

FIGURES 5–9 in A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa

FIGURES 5–9. Haroldius lyleae Daniel, Strümpher & Snäll, new species, holotype (TMSA). 5, prothorax; 6, pygidium; 7, aedeagus, ventral view; 8, aedeagus, lateral view; 9, aedeagus, dorsal view.

FIGURE 15 in A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa

FIGURE 15. Distribution of Haroldius lyleae Daniel, Strümpher & Snäll, new species from South Africa (Yellow circles; circle with black point indicate type locality – "Soetvlakte Farm").

FIGURES 1–4 in A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa

FIGURES 1–4. Haroldius lyleae Daniel, Strümpher & Snäll, new species, holotype (TMSA). 1, habitus, dorsal view; 2, habitus, ventral view; 3, habitus, lateral view; 4, holotype labels.

FIGURES 10–14 in A new species and new distributional records of Haroldius Boucomont, 1914 (Coleoptera: Scarabaeidae: Scarabaeinae) from southern Africa

FIGURES 10–14. Haroldius lyleae Daniel, Strümpher & Snäll, new species, paratype (ESRC). 10, habitus, dorsal view; 11, paratype labels; 12, habitus, ventral view; 13, clypeus; 14, host ant: Monomorium albopilosum, dorsal view.