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Fig. 7 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 7 (next page). A. Plagigeyeria mostarensis, Kuščer 1933, holotype, from Kuščer 1933, assigned to Izvor Buna in Blagaj but rather represents the form from near spring Bunica in Hodbina. B. P. mostarensis, hypotype of Schütt, 1970, from spring Bunica (SMF 221236). C. P. mostarensis, likely topotype from spring Bunica (JG F1146). D–E. P. mostarensis var., from spring of Buna assigned as type locality (JG F0536). Fa–Fb. Paladilhiopsis (Costellina) turrita (Kuščer, 1933) paratype and holotype from Kuščer 1933, spring Izvor in valley of river Jadro near Split. G. P. pseudocostellina sp. nov., left bank side spring of river Buna 400 m from main spring (holotype HNHM-MOLL-104164). H. P. pseudocostellina sp. nov., left bank side spring of river Buna 200 m from main spring (paratype JG F1140). I. Travunijana ovalis (Kuščer, 1933), holotype from Kuščer, assigned to Izvor Buna in Blagaj but not found in assigned locality, only in spring Bunica in Hodbina. J–K. T. ovalis (Kuščer, 1933), spring Bunica near Hodbina (JG F1145). L. T. robusta (Schütt, 1959), Trebinje district, Tučevac spring at right bank of Trebišnjica River (JG F1163). M. T. robusta, Trebinje district, Gorica, cave Vruljak 1 (JG F0831). N. T. robusta asculpta (Schütt, 1972), spring Ombla (Dubrovačka Rijeka) at Komolac near Dubrovnik (JG F1080).

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Fig. 5 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 5 (next page). A. Plagigeyeria plagiostoma (A. J. Wagner, 1914), from Vrelo Bosne in Ilidža, (topotype JG F1116). B. Travunijana vruljakensis Grego & Glöer, 2019, Gorica near Trebinje, spring Studenac (holotype HNHM-MOLL-104416). C. P. inflata (A. J. Wagner, 1928), from Vrelo Bosne in Ilidža (topotype JG F1117). D. P. ljutaensis sp. nov., Konjic district, spring of River Ljuta (holotype HNHM-MOLL-104180). E. P. konjicensis sp. nov., small spring at left bank of Ljuta River, 600 m downstream of the main spring (holotype HNHM-MOLL-104174); F–G. P. olsavskyi sp. nov., Studenci, spring Kajtazovina F. Holotype (HNHM-MOLL-104178). G. Paratype (JG F1192). H–I. P. olsavskyi sp. nov., Ljubuški district, Donji Proložac, spring Mali Prokop (JG F1209).

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Fig. 1 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 1 (next page). Studied localities of Plagigeyeria and Travunijana in Hercegovina and phreatic hydrology of the related karstic aquifers with flow directions of the groundwater. Hydrological situation after Stevanović et al. 2014. 1–14. Newly studied localities (red dot ●). 1. Konjic, spring Ljuta (LT of Plagigeyeria ljutaensis sp. nov.). 2. Konjic spring at left side of Ljuta stream (LT of P. konjicensis sp. nov.). 3. Donja Jablanica, Komadinovo Vrelo (LT of P. erossi sp. nov.). 4. Široki Brijeg, spring Lištica, Bilo Vrilo (LT of P. listicaensis sp. nov.). 5. Studencí, spring Kajtazovina (LT of P. olsavskyi sp. nov.). 6. Studencí, Vrilo Guljevina (LT of P. jakabi sp. nov.). 7. Donji Proboj near Ljubuški, spring Mali Prokop (P. olsavskyi sp. nov.). 8. Vitina, spring Vrioštica (LT of P. vriosticaensis sp. nov. and P. ozimeci sp. nov.). 9. Lukende, Jakšenica spring (LT of P. angyaldorkae sp. nov.). 10. Studenci, spring Vakuf (P. cf. jakabi). 11. Dabarsko Polje, spring Vrijeka (LT of P. reischuetzorum sp. nov. and Travunijana gloeri sp. nov.). 12. Trebinje, spring Tučevac (T. robusta, Schütt, 1959 and P. cf. lewarnei sp. nov.). 13. Trebinje, Gorica, spring Vruljak (T. robusta and LT of P. lewarnei sp. nov.). 14. Vučija, Sušićka Jama Estavelle (P. lewarnei sp. nov.). 15–32. Historical localities (blue dot ●): 15. Blagaj, spring of Buna (P. mostarensis var. Kuščer, 1933 and LT of P. pseudocostellina sp. nov.). 16. Hodbina, spring of Bunica (P. mostarensis Kuščer, 1933 and Travunijana ovalis (Kuščer, 1933)). 17. Bileća, spring Trebišnjica (T. robusta (Schütt, 1959) and T. tribunicae (Schütt, 1953)). 18. Čepelica, spring Čepo (T. robusta (Schütt, 1959)). 19. Bileća, spring Miruše (T. robusta (Schütt, 1959) and T. tribunicae (Schütt, 1953), P. zetatridyma Schütt, 1959). 20. Bileća, spring Dobriševo (T. robusta, (Schütt, 1959)). 21. Fatničko Polje, Obod Cave (T. robusta (Schütt, 1959), P. reischuetzorum sp. nov., T. gloeri sp. nov.). 22. Gacko, Cave Stepen (T. robusta). 23. Mlini, spring south of Plat (T. robusta asculpta Schütt, 1972). 24. Komolac, spring Ombla (T. robusta asculpta Schütt, 1972 and T. angelovi Schütt, 1972,). 25. Mali Zaton, spring Mlin (T. robusta asculpta (Schütt, 1972), T. angelovi (Schütt, 197)). 26. Svitavsko Blato, spring Sjekoše (T. edlaueri (Schütt, 1961), T. nitida (Schütt, 1963)). 27. Svitavsko Blato, spring Sopot Mlin (T. edlaueri (Schütt, 1961), T. nitida (Schütt 1963)). 28. Svitavsko Blato, spring Bajovci (T. edlaueri (Schütt, 1961)). 29. Metković, spring Glušči (T. edlaueri and T. nitida (Schütt, 1963)). 30. Ploče, spring Ocuša at bank of Bačinsko Jezero Lake (T. klemmi (Schütt, 1961)). 31. Vrgorac, spring Stenjevac (T. klemmi (Schütt, 1961)). 32. Ilidža, Vrelo Bosne (P. plagiostoma (A. J. Wagner, 1914) and P. inflata (A. J. Wagner, 1928)). 1–43. Karst basins – Poljes of the studied region (green square ■): 1. Konavosko Polje, 50 m a.s.l. 2. Zubačko Polje, 672 m a.s.l. 3. Konjsko and Carevo P., 829–875 m a.s.l. 4. Mokro Polje, (Popovo Polje) 269 m a.s.l. 5. Popovo Polje, 227 m a.s.l. 6. Jasen Polje, 556 m a.s.l. 7. Ljubomirsko Polje, 506 m a.s.l. 8. Ljubinjsko Polje, 396 m a.s.l. 9. Plansko Polje, 609 m a.s.l. 10. Fatničko Polje, 452 m a.s.l. 11. Dabarsko Polje, 472 m a.s.l. 12. Lukavačko Polje, 895 m a.s.l. 13. Slato Polje, 1012 m a.s.l. 14. Cerničko Polje, 816 m a.s.l. 15. Gatačko Polje, 936 m a.s.l. 16. Nevesinjsko Polje, 817 m a.s.l. 17. Trusinsko Polje, 866 m a.s.l. 18. Stolačko Polje, 45 m a.s.l. 19. Rotimlja Polje, 422 m a.s.l. 20. Crničko Polje, 212 m a.s.l. 21. Hutovo Blato, 2 m a.s.l. 22. Svitavsko Blato, 2 m a.s.l. 23. Gradac Polje, 88 m a.s.l. 24. Vrgorsko (Jezerac) Polje, 20 m a.s.l. 25. Rastok Polje, 72 m a.s.l. 26. Ljubuško Polje, 58 m a.s.l. 27. Studeničko Polje, 25 m a.s.l. 28. Čitluško Polje, 217 m a.s.l. 29. Mostarsko Blato, 223 m a.s.l. 30. Mokro Polje, 230 m a.s.l. 31. Rašanjsko Polje, 344 m a.s.l. 32. Kočerinsko Polje, 302 m a.s.l. 33. Imotsko & Grudsko P., 251–280 m a.s.l. 34. Posuško Polje, 578 m a.s.l. 35. Dugo (Dugorudo) Polje, 1206 m a.s.l. 36. Ravanjsko Polje, 1131 m a.s.l.; 37. Rakitno Polje, 890 m a.s.l. 38. Vučipolje, 977 m a.s.l.; 39. Roško Polje, 894 m a.s.l. 40. Duvanjsko Polje, 865 m a.s.l. 41. Džepi Polje, 830 m a.s.l. 42. Prečko Polje, 1002 m a.s.l. 43. Hansko Polje, 835 m a.s.l. 44. Donje Ziemlje, 507 m a.s.l.

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Fig. 19 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 19. Localities of species of Travunijana in Hercegovina and adjacent regions and the most likely distribution range of the known species within the related aquifers (colour blotches) and sampling localities (dots with numbers). Freshwater springs are in circles and brackish or submarine springs are as marked by diamonds, arrows show main karst conduits and locality numbers are according to the legend from Fig. 1.

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Fig. 14. A in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 14. A. Travunijana vruljakensis Grego & Glöer, 2019, Hercegovina, Vrelo Studenac, Gorica near Trebinje, SBMNH 625961 (SEM SBMNH Vanessa Delnavaz). B. T. angelovi (Schütt, 1972), Spring Ombla, Komolac near Dubrovnik, paratype (SMF 318892). C–D. T. tribunicae (Schütt, 1963), Hercegovina, Bileća, main spring of Trebijšnica. C. SMF 191518. D. SMF 318909. Scale bars = 1 mm (B–D SEM SMF Sigrid Hof).

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Fig. 16. A–B in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 16. A–B. Travunijana klemmi (Schütt, 1961), Croatia, spring in Stenjevac near Vrgorac. A. SMF 318873. B. Paratype SMF 164345. C. T. edlaueri (Schütt, 1961), Hercegovina, Sopot Mlin at south bank of Svitavsko Blato near Metkovic, SMF 171671. D. T. edlaueri, Hercegovina, Sjekoše at south bank of Svitavsko Blato near Metkovic, SMF 318858. Scale bars = 1 mm (SEM SMF Sigrid Hof).

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Fig. 11. A in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 11. A. Plagigeyeria ozimeci sp. nov., Vitina, Vrilo Vrioštica, paratype SBMNH 626440. B. Plagigeyeria lewarnei sp. nov., spring Tučevac in Trebinje, SBMNH 626406. Scale bars = 1 mm (SEM SBMNH Vanessa Delnavaz).

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Fig. 8. A in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 8. A. Plagigeyeria reischuetzorum sp. nov., Bjeljani, Dabarsko Polje, Vrelo Vruljak, paratype (SBMNH 632724). B. P. pseudocostellina sp. nov., Blagaj, side spring of Buna at left bank, paratype (SBMNH 626414). C. P. mostarensis Kuščer, 1933, Blagaj Izvor Bune (SBMNH 626411). D. P. mostarensis Kuščer, 1933, Hodbina, Izvor Bunice (SBMNH 626405). Scale bars = 1 mm (SEM SBMNH Vanessa Delnavaz).

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Fig. 2 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 2. Photos of studied localities in Hercegovina. A. Konjic. Ljuta spring (LT of Plagigeyeria ljutaensis sp. nov.). B. Blagaj, spring Buna (P. mostarensis var.). C. Konjic, spring at left bank of Ljuta Rivulet (LT of P. konjicensis sp. nov.). D. Donja Jablanica, Komadinovo Vrelo (LT of P. erossi sp. nov.). E. Vučija, Sušicka Jama (P. cf. lewarnei sp. nov.). F. Trebinje, Gorica, spring Vruljak 1 (P. robusta and P. lewarnei sp. nov.). Photos: Jozef Grego, Dorottya Angyal and Mário Olšavský.

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Fig. 3 in Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions

Fig. 3. Photos of studied localities in Hercegovina. A. Dabarsko Polje, Sušicka Peć cave near spring Vrijeka (LT of Plagigeyeria reischuetzorum sp. nov. and Travunijana gloeri sp. nov.). B. Tihaljina, spring Tihaljina in Peć Mlini Cave (Belgrandia torifera Schütt, 1961). C. Hodbina, spring Bunica (likely LT of P. mostarensis Kuščer, 1933, P. ovalis Kuščer, 1933). D. Studenci, spring Kajtazovina (LT of P. olsavskyi sp. nov.) E. Vitina, spring Vrioštica (LT of P. vriosticaensis sp. nov. and P. ozimeci sp. nov.) F. Lukende, Jakšenica, Vrelo Jakšenica (LT of P. angyaldorkae sp. nov.) Photos: Jozef Grego, Dorottya Angyal and Mário Olšavský.

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Figures 3-8 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figures 3-8 - 3–4 Paladilhiopsis prekalensis sp. n. (holotype) 5–6 Paladilhiopsis lozeki sp. n. (holotype) 7–8 Paladilhiopsis szekeresi sp. n. (holotype); (photos: Peter Glöer, Jozef Grego).

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Figures 9-14 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figures 9-14 - 9–10 Paladilhiopsis falniowskii sp. n. (holotype) 11 Paladilhiopsis gittenbergeri (holotype) 12 Paladilhiopsis szarowskae (holotype) 13–14 Paladilhiopsis wohlberedti sp. n. (holotype); (photos: Peter Glöer, Jozef Grego, Zoltán Fehér).

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Figures 29-34 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figures 29-34 - 29–32 Vinodolia matjasici: 29–30 Albania, Bajzë, Syri i Sheganit 31–32 Montenegro, spring Vitoja 33–34 Saxurinator schlickumi: Kosovo: Peja, Rugova Gorge; (photos: Peter Glöer, Jozef Grego).

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Figures 21-28 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figures 21-28 - 21–26 Plagigeyeria zetaprotogona: 21–22 Montenegro, Vitoja Spring 23–24 Albania, Bajzë, Syri i Sheganit Spring 25–26 Albania, Rraps, Vrak Spring; Plagigeyeria cf. zetaprotogona: 27–28 Albania, Dverpte Cave. (photos: Peter Glöer, Jozef Grego).

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Figure 2 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figure 2 - Various habitats where subterranean gastropods were found: A Prekal, Zhyla Cave B Shoshan Spring C Tamarë, spring above the trout farm D Krumë, Vrela Spring E Novosellë, White Drin Spring F Bajzë, Syri i Sheganit Spring G Dverpte Cave south of Tamarë H sampling from sand and fine gravel by a sieve combined with polyamide pantyhose (photos: Zoltán Fehér, Maroš Grego, Dávid Murányi).

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Figures 15-20 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figures 15-20 - 15–17 Plagigeyeria steffeki sp. n. (holotype) 18–20 Plagigeyeria gladilini (topotype from the White Drin Spring near Novosellë).

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Figure 1 from: Grego J, Glöer P, Erőss ZP, Fehér Z (2017) Six new subterranean freshwater gastropod species from northern Albania and some new records from Albania and Kosovo (Mollusca, Gastropoda, Moitessieriidae and Hydrobiidae). Subterranean Biology 23: 85-107. https://doi.org/10.3897/subtbiol.23.14930

Figure 1 - Distribution of the new species and faunal records mentioned in this paper. Red dots indicate type localities of new the species. 1 Prekal, Zhyla Cave (Paladilhiopsis prekalensis sp. n.) 2 Shoshan (Paladilhiopsis lozeki sp. n.) 3 Tamarë (Paladilhiopsis szekeresi sp. n. and P. wohlberedti sp. n.) 4 Krumë, Vrela Spring (Paladilhiopsis falniowskii sp. n., Plagigeyeria steffeki sp. n. and Paladilhiopsis szarowskae) 5 Novosellë, White Drin Spring (type locality of Plagigeyeria gladilini and Iglica illyrica) 6 Bajzë, Syri i Sheganit Spring (Plagigeyeria zetaprotogona) 7 Rrash, Vrak Spring (Plagigeyeria zetaprotogona) 8 Tamarë, Dverpte Cave (Plagigeyeria zetaprotogona) 9 Kosovo, Pejë, Rugova Gorge, (Saxurinator schlickumi) 10 Vau i Dejës (type locality of Paladilhiopsis gittenbergeri) 11 Kosovo, Decani Valley (Iglica illyrica).

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Figure 8 from: Hofman S, Grego J, Rysiewska A, Osikowski A, Falniowski A (2021) Two new species of the Balkan genus Paladilhiopsis Pavlović, 1913 (Caenogastropoda, Moitessieriidae). ZooKeys 1046: 157-176. https://doi.org/10.3897/zookeys.1046.64489

Figure 8 Renal and pallial section of the female reproductive organs of Paladilhiopsis stellatus X. Abbreviations: bc – bursa copulatrix, cbc – duct of bursa, ga – albuminoid gland, gn – nidamental gland, gp – gonoporus, ov – oviduct, ovl – loop of (renal) oviduct, rs – seminal receptacle (in the position of Radoman's (1983) rs1 – distal seminal receptacle). Scale bar: 500 μm.

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Figure 7 from: Hofman S, Grego J, Rysiewska A, Osikowski A, Falniowski A (2021) Two new species of the Balkan genus Paladilhiopsis Pavlović, 1913 (Caenogastropoda, Moitessieriidae). ZooKeys 1046: 157-176. https://doi.org/10.3897/zookeys.1046.64489

Figure 7 A teleoconch sculpture of Paladilhiopsis arionB–D radula of P. stellatus. Scale bars 50 μm (A); 10 μm (B–D).

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Figure 5 from: Hofman S, Grego J, Rysiewska A, Osikowski A, Falniowski A (2021) Two new species of the Balkan genus Paladilhiopsis Pavlović, 1913 (Caenogastropoda, Moitessieriidae). ZooKeys 1046: 157-176. https://doi.org/10.3897/zookeys.1046.64489

Figure 5 Maximum likelihood tree computed for concatenated partial sequences of COI and H3 sequences. Bootstrap supports given if ≥ 60%.

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Last verified 2026-04-29Open record

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Last verified 2026-04-29Open record