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Appendix 1 in Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths
<p><b>Appendix 1</b> (continued) Number of non-otolith taxa identified to lowest taxonomic ranking within each Claiborne Group unit. Shaded areas represent the stratigraphic range of taxa within Claiborne Group units in Alabama, USA.</p><table><thead><tr><th><b>Taxon</b></th><th colspan="2"><b>Meridian lower upper Tallahatta “lower” “upper” Lisbon/ lower Tallahatta Sand Tallahatta Tallahatta /Lisbon Lisbon Lisbon Gosport Gosport Fm. Member Fm. Fm. contact Fm. Fm. contact Sand</b></th><th><b>Gosport Sand</b></th><th><b>Total</b></th></tr></thead><tbody><tr><th><i>Myliobatis</i> sp. 1</th><td>5</td><td>307</td><td></td><td><b>312</b></td></tr><tr><th><i>Myliobatis</i> sp. 2</th><td></td><td>18 207 1 48</td><td colspan="2">13 <b>287</b></td></tr><tr><th><i>Pseudaetobatus belli</i></th><td>15</td><td>210</td><td></td><td><b>225</b></td></tr><tr><th><i>Rhinoptera</i> sp. <i>Meridiania</i> cf. <i>M. convexa</i></th><td>16</td><td>14 18 181 1 5 1525 1 1</td><td colspan="2">131 <b>1891</b> 2 <b>4</b></td></tr><tr><th><i>Leidybatus jugosus</i></th><td></td><td>2 28 41</td><td></td><td><b>71</b></td></tr><tr><th><i>Burnhamia daviesi</i></th><td></td><td>6 1 1</td><td>4</td><td><b>12</b></td></tr><tr><th><i>Eoplinthicus yazooensis</i></th><td></td><td></td><td colspan="2">1 <b>1</b></td></tr><tr><th>Batomorphii indet.</th><td></td><td>4</td><td>3</td><td><b>7</b></td></tr><tr><th>Myliobatidae indet.</th><td></td><td>750 4 74 776 1 7 762</td><td>212</td><td><b>2586</b></td></tr><tr><th><i>Cylindracanthus ornatus</i></th><td></td><td>2 1</td><td></td><td><b>3</b></td></tr><tr><th><i>Cylindracanthus rectus</i></th><td></td><td>24 16 4 2</td><td></td><td><b>46</b></td></tr><tr><th><i>Cylindracanthus</i> sp.</th><td></td><td>4 4 13 45 6</td><td></td><td><b>72</b></td></tr><tr><th><i>Pycnodus</i> sp.</th><td></td><td>8 2 10</td><td></td><td><b>20</b></td></tr><tr><th>Lepisostidae indet.</th><td></td><td>1 2 10</td><td>23</td><td><b>36</b></td></tr><tr><th><i>Egertonia isodonta</i></th><td></td><td>35 12 7 37</td><td colspan="2">37 <b>128</b></td></tr><tr><th><i>Paralbula</i> aff. <i>P. marylandica</i></th><td></td><td>17 52</td><td></td><td><b>69</b></td></tr><tr><th><i>Phyllodus toliapicus Albula eppsi Albula oweni</i></th><td></td><td>8 15 3 23 2 1 6 1 15 3 1 6</td><td colspan="2"><b>26 33 25</b></td></tr><tr><th><i>Albula</i> sp.</th><td></td><td>3 1</td><td></td><td>4</td></tr></tbody></table>
Appendix 1 in Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths
<p><b>Appendix 1</b> (continued) Number of non-otolith taxa identified to lowest taxonomic ranking within each Claiborne Group unit. Shaded areas represent the stratigraphic range of taxa within Claiborne Group units in Alabama, USA.</p><table><thead><tr><th><b>Taxon</b></th><th colspan="5"><b>Meridian lower upper Tallahatta “lower” “upper” Tallahatta Sand Tallahatta Tallahatta /Lisbon Lisbon Lisbon Fm. Member Fm. Fm. contact Fm. Fm.</b></th><th><b>Lisbon/ Gosport contact</b></th><th><b>lower Gosport Sand</b></th><th><b>Gosport Sand</b></th><th><b>Total</b></th></tr></thead><tbody><tr><th><i>Physogaleus alabamensis</i> comb. nov. <i>Physogaleus secundus</i></th><td></td><td>31 131</td><td>1 1</td><td></td><td>185 232 1 53 555 1</td><td></td><td>15 38</td><td colspan="2">17 <b>507</b> 27 <b>786</b></td></tr><tr><th><i>Physogaleus</i> sp.</th><td></td><td></td><td></td><td></td><td>1 1</td><td></td><td></td><td></td><td><b>2</b></td></tr><tr><th><i>Galeocerdo eaglesomei</i></th><td></td><td></td><td></td><td></td><td>4 13</td><td>1</td><td>2</td><td>18</td><td><b>38</b></td></tr><tr><th><i>Galeocerdo clarkensis</i></th><td></td><td></td><td></td><td></td><td></td><td></td><td>83</td><td colspan="2">3 <b>86</b></td></tr><tr><th><i>Galeocerdo</i> sp.</th><td></td><td></td><td></td><td></td><td></td><td></td><td>16</td><td></td><td><b>16</b></td></tr><tr><th>Galeomorphii indet.</th><td></td><td></td><td></td><td></td><td>1</td><td></td><td></td><td></td><td><b>1</b></td></tr><tr><th>Pristidae indet.</th><td></td><td></td><td></td><td></td><td>4</td><td></td><td>2</td><td colspan="2">7 <b>13</b></td></tr><tr><th><i>Anoxypristis</i> sp. <i>Pristis</i> sp.</th><td></td><td>1 27</td><td></td><td></td><td>11 4 91 27</td><td></td><td>2 36</td><td>66</td><td><b>18 247</b></td></tr><tr><th><i>Propristis schweinfurthi</i></th><td></td><td></td><td></td><td></td><td></td><td></td><td>2</td><td colspan="2">8 <b>10</b></td></tr><tr><th>Myliobatiformes indet.</th><td></td><td></td><td></td><td></td><td>1 288 3</td><td></td><td></td><td>60</td><td><b>352</b></td></tr><tr><th>“ <i>Dasyatis</i> ” aff. <i>D. charlisae</i></th><td></td><td></td><td></td><td></td><td>4 4</td><td></td><td></td><td></td><td><b>8</b></td></tr><tr><th>“ <i>Dasyatis</i> ” <i>jaekeli</i></th><td></td><td>3</td><td></td><td></td><td>13</td><td></td><td></td><td></td><td><b>16</b></td></tr><tr><th>“ <i>Dasyatis</i> ” sp.</th><td></td><td></td><td></td><td></td><td>6</td><td></td><td>2</td><td></td><td><b>8</b></td></tr><tr><th><i>Aturobatis</i> aff. <i>A. aquensis</i></th><td></td><td></td><td></td><td></td><td>2</td><td></td><td></td><td></td><td><b>2</b></td></tr><tr><th><i>Coupatezia</i> sp.</th><td></td><td>1</td><td></td><td></td><td>17</td><td></td><td></td><td></td><td><b>18</b></td></tr><tr><th><i>Hypolophodon sylvestris</i></th><td></td><td>7</td><td></td><td></td><td></td><td></td><td></td><td></td><td><b>7</b></td></tr><tr><th><i>Jacquhermania duponti</i></th><td></td><td></td><td></td><td></td><td>11</td><td></td><td>16</td><td colspan="2">1 <b>28</b></td></tr><tr><th><i>Aetobatis</i> sp.</th><td></td><td></td><td></td><td></td><td>12 103</td><td>1</td><td></td><td>11</td><td><b>127</b></td></tr><tr><th><i>Aetomylaeus</i> sp.</th><td>11</td><td>80</td><td></td><td></td><td>12 123 4</td><td></td><td>153</td><td>9</td><td>392</td></tr></tbody></table>
Evaluating the suitability of close-kin mark-recapture as a demographic modelling tool for a critically endangered elasmobranch population
<p>Estimating the demographic parameters of contemporary populations is essential to the success of elasmobranch conservation programmes, and to understanding their recent evolutionary history. For benthic elasmobranchs such as skates, traditional fisheries-independent approaches are often unsuitable as the data may be subject to various sources of bias, whilst low recapture rates can render mark-recapture programmes ineffectual. Close-kin mark-recapture (CKMR), a novel demographic modelling approach based on the genetic identification of close relatives within a sample, represents a promising alternative approach as it does not require physical recaptures. We evaluated the suitability of CKMR as a demographic modelling tool for the critically endangered blue skate (<em>Dipturus batis</em>) in the Celtic Sea using samples collected during fisheries-dependent trammel-net surveys that ran from 2011 to 2017. We identified three full-sibling and 16 half-sibling pairs among 662 skates, which were genotyped across 6,291 genome-wide single nucleotide polymorphisms (SNPs), 15 of which were cross-cohort half-sibling pairs that were included in a CKMR model. Despite limitations owing to a lack of validated life-history trait parameters for the species, we produced the first estimates of adult breeding abundance, population growth rate, and annual adult survival rate for <em>D. batis</em> in the Celtic Sea. The results were compared to estimates of genetic diversity, effective population size (N<sub>e</sub>), and catch per unit effort (CPUE) estimates from the trammel-net survey. Although each method was characterised by wide uncertainty bounds, together they suggested a stable population size across the time-series. Recommendations for the implementation of CKMR as a conservation tool for data-limited elasmobranchs are discussed. In addition, the spatio-temporal distribution of the 19 sibling pairs revealed a pattern of site-fidelity in <em>D</em>. <em>batis</em>, and supported field observations suggesting an area of critical habitat that could qualify for protection might occur near the Isles of Scilly.</p>
Figure 3 in Elasmobranch diversity across a remote coral reef atoll revealed through environmental DNA metabarcoding
Figure 3. Spatial variation in elasmobranch abundance and diversity inferred from eDNA metabarcoding of surface (A) and deep (40 m) (B) water samples collected around Diego Garcia. Negaprion acutidens is not visible in the charts as a result of low copy number, but was detected at site 8 in surface samples. Numbers correspond to the site numbers detailed in Figure 1.
Figure 4 in Elasmobranch diversity across a remote coral reef atoll revealed through environmental DNA metabarcoding
Figure 4. Venn diagram showing the overlap of shark species detected in previous UVC and BRUVS surveys in the MPA and the eDNA samples from around Diego Garcia analysed in this study.
Fig. 2 in Vulnerability of elasmobranchs caught as bycatch in the grouper longline fishery in the Gulf of Gabès, Tunisia Abstract
Fig. 2: Productivity, susceptibility and vulnerability scores of elasmobranch species caught by the grouper demersal longline fishery in the Gulf of Gabès. Numbers correspond to elasmobranch species as listed in Table 4. The colors represent the relative vulnerability: the green areas being the lowest, the yellow ones being the moderate and the red areas being the highest.
Fig. 1 in Vulnerability of elasmobranchs caught as bycatch in the grouper longline fishery in the Gulf of Gabès, Tunisia Abstract
Fig. 1: Map showing the location of the grouper demersal longline sets surveyed during 2016 () and 2017 () in the Gulf of Gabès.
FIGURE 5 in Trophic relationships in the Early Miocene Upper Marine Molasse of Baden-Württemberg, Southwest Germany, with special emphasis on the elasmobranch fauna
FIGURE 5. Similarity analyses (cluster analyses) of the shark and ray fauna of the OMM deposits in Baden-Württemberg, Bavaria and Switzerland. Redrawn after Höltke et al. (2020; 2022b).
FIGURE 3. A in Trophic relationships in the Early Miocene Upper Marine Molasse of Baden-Württemberg, Southwest Germany, with special emphasis on the elasmobranch fauna
FIGURE 3. A. Tearing type [Odontaspis molassica (Probst, 1878)]. Tooth height: c. 9. 8 mm. Locality: Messkirch-Walbertsweiler. Redrawn after Pfeil (1991, pl. 2, fig. 2). B. Tearing type [Carcharias contortidens (Agassiz, 1843)]. Tooth height: c. 17. 9 mm. Locality: Messkirch-Walbertsweiler. Redrawn after Pfeil (1991, pl. 2, fig. 3, named here as "Synodontaspis acutissima"). C. Cutting type - sensu stricto cutting subtype [Galeocerdo aduncus (Agassiz, 1843)]. Tooth width: c. 14. 2 mm. Locality: Messkirch-Walbertsweiler. Redrawn after Pfeil (1991, pl. 3, fig. 16). D. Cutting type - sensu stricto cutting subtype [Otodus (Megaselachus) chubutensis (Ameghino, 1901)]. Tooth height: c. 65 mm. Locality: Messkirch-Walbertsweiler. Redrawn after Pfeil (1991, pl. 2, fig. 10, named here as "Procarcharodon megalodon"). E. Cutting type – cutting-clutching subtype [Carcharhinus similis (Probst, 1878)]. Tooth height: c. 12. 9 mm. Locality: Baltringen. Redrawn after Probst (1878, pl. 1, fig. 19), Size according to Reinecke et al. (2011, Text-Fig. 21). F. Cutting type – cutting-clutching subtype [Paragaleus tenuis (Probst, 1878)]. Tooth height: c. 4 mm. Locality: Messkirch-Walbertsweiler. Redrawn after Pfeil (1991, pl. 3, fig. 8). G. Crushing type [Dasyatis rugosa (Probst, 1877)]. Tooth height: c. 3 mm. Locality: Messkirch-Walbertsweiler. Redrawn after Pfeil (1991, pl. 4, fig. 9)]. H. Crushing type [Rynchobatus pristinus (Probst, 1877)]. Tooth height: c. 4 mm. Locality: Baltringen. Redrawn after Probst (1877, pl. 1, fig. 19)]. I. Clutching type [Pachyscyllium dachiardii (Lawley, 1876)]. Tooth height c. 6 mm. Locality: Ursendorf. Redrawn after Höltke et al. (2020, pl. 7, fig. 1]. J. Clutching type [Ginglymostoma delfortriei Daimeries, 1889)]. Tooth width. c. 7. 6 mm. Locality: Baltringen. Redrawn after Pfeil [1991, pl. 2, fig. 1 named here as "Ginglymostoma grandis"). K. Grinding type [Aeobatus arcuatus (Agassiz, 1843)]. Tooth width. C. 14 mm. Locality: Ursendorf. Redrawn after Höltke et al. (2020, pl. 9, fig. 8). L. Grinding type [Rhinoptera cf. studeri (Agassiz, 1843). Tooth width: c. 23 mm. Locality: Ursendorf. Redrawn after Höltke et al. (2020, pl. 9, fig. 11).
FIGURE 1 in Trophic relationships in the Early Miocene Upper Marine Molasse of Baden-Württemberg, Southwest Germany, with special emphasis on the elasmobranch fauna
FIGURE 1. New lithostratigraphic terminology for the Ottnangian deposits of the OMM in Southwest Germany. Modified after Heckeberg et al. (2010).
FIGURE 4 in Trophic relationships in the Early Miocene Upper Marine Molasse of Baden-Württemberg, Southwest Germany, with special emphasis on the elasmobranch fauna
FIGURE 4. Generalized trophic interaction scheme of the five OMM deposits discussed in this paper. Some of the fish feeding taxa also feed on invertebrates. A definitive separation in exclusive fish and invertebrate feeders is often not possible. The same problem exists for members of the shark genus Isistius, which are ectoparasites of large marine animals but also feed on cephalopods.
FIGURE 14. A-G in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 14. A-G. Amamriabatis heni nov gen. nov. sp. A. anterior tooth KEB 1-218, A1. Occlusal view, A2. Basal view, A3. Lingual view, B. antero-lateral tooth KEB 1-219 (Holotype), B1. Occlusal view, B2. Lingual view, B3. Basal view, C. juvenile tooth KEB 1-220, occlusal view, D. antero-lateral tooth KEB 1-221, D1. Occlusal view, D2. Basal view, D3. Labial view, E. lateral tooth KEB 1-222, E1. Occlusal view, E2. Lingual view, E3. Basal view, F. antero-lateral tooth KEB 1-223, F1. Occlusal view, F2. Basal view, F3. Labial view, G. lateral tooth KEB 1-224, G1. Occlusal view, G2. Basal view, G3. Lingual view; H. Archaeomanta sp. KEB 1-225, H1. Lateral view, H2. Labial view.
FIGURE 12. A-C in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 12. A-C. Mecotrygon asperodentulus nov. gen nov. sp. A. anterior tooth KEB 1-188, A1. occlusal view, A2. Profile, A3. Basal view; B. lateral tooth KEB 1-189, B1. occlusal view, B2. labial view, B3. Profile; C. lateral tooth KEB 1-190, HOLOTYPE, C1. Occlusal view, C2. Lingual view, C3. Basal view, C4. Profile; D-M. Himantura souarfortuna nov. sp. D.?posterior tooth KEB 1-191, D1. Occlusal view, D2. Profile, D3. Basal view; E. antero-lateral tooth KEB 1- 192, occlusal view; F. anterior tooth KEB 1-193, F1. Occlusal view, F2. Labial view; G. antero-lateral tooth KEB 1-194, G1. Occlusal view, G2. Profile; H. anterior tooth KEB 1-195, occlusal view; I. lateral tooth KEB 1-196, occlusal view; J. A. anterior tooth KEB 1-197, occlusal view; K. lateral tooth KEB 1-198, K1. Occlusal view, K2. Basal view; L. A. anterior tooth KEB 1-199, occlusal view; M. lateral tooth KEB 1-200, occlusal view. N-O. Dasyatoid indet. N. antero-lateral tooth KEB 1-201, N1. Occlusal view, N2. Basal view; O. A. antero-lateral tooth KEB 1-202, occlusal view; P-Q. Arechia sp. P. lateral tooth KEB 1-203, occlusal view; Q. A. anterior tooth KEB 1-204, Q1. Occlusal view, Q2. Lingual view.
FIGURE 10. A-D in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 10. A-D: Propristis cf. schweinfurti. A. Rostral denticle KEB 1-172, A1. Profile, A2. dorsal view; B. Rostral denticle KEB 1-173, Profile; C. Rostral denticle KEB 1-174, C1. Profile, C2. Dorsal view, C3. basal view; D. Rostral denticle KEB 1-175, D1. profile. D2. dorsal view; E: Pristis sp. Rostral denticle, KEB 1-165, dorsal view; F-G. Rhynchobatus cf. vincenti. F. anterior tooth KEB 1-176, F1. Occlusal view, F2. Basal view; G. anterior tooth KEB 1-177, occlusal view; H-I.?Torpedo sp. H. lateral tooth KEB 1-178, occlusal view, I. lateral tooth KEB 1-179, occlusal view.
FIGURE 9. A-F in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 9. A-F: Propristis cf. schweinfurti, A. Anterior oral tooth KEB 1-166, A1. Lingual view, A2. Occlusal view, A3. profile; B. Antero-lateral oral tooth KEB 1-167, B1. Occlusal view, B2. Lingual view, B3. Labial view; C. Anterior oral tooth KEB 1-168, C1. Lingual view, C2. Basal view, C3. Profile; D. lateral tooth KEB 1-169, D1. Lingual view, D2. Occlusal view, D3. Basal view, D4. Magnificence of crown-root boundary of D3; E.?male lateral tooth KEB 1-170, E1. Occlusal tooth, E2. Basal view; F.?male anterior tooth KEB 1-171, F1. Lingual view, F2. Occlusal view; G-L: Pristis sp. G. porterior tooth KEB 1-158, G1. Occlusal view, G2. Basal view; H. anterior tooth KEB 1-159, occlusal view; I. lateral tooth KEB 1-160, I1 occlusal view, I2., lingual view. J lateral tooth KEB 1-161, occlusal view; K. anterior tooth KEB 1-162, occlusal view; L. lateral tooth of?juvenile KEB 1-163, L1. Occlusal view, L2. Basal view; M. lateral tooth of juvenile KEB 1-164, M1. Occlusal view, M2. Profile.
FIGURE 1 in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 1. Paleotemperatures (ice-free deep-ocean T°/ tropical sea surface T°) and Thermic events during the "doubthouse" conditions of Eocene period (from Cramwinckel et al., 2018 modified with events dating from Hollis et al., 2019). Stratigraphically and geographical locations of the main deposits with Elasmobranch associations along the southwestern Tethys. Abbreviations: DAK: Dakhla (Adnet et al., 2010), GEN: Genam (Zouhri et al., 2017, in press); AZ: Aznag (Tabuce et al., 2005) PM: Phosphate ores (see Noubhani and Cappetta, 1997), Morocco; GAF: Gafsa basin (see Arambourg, 1952); KEBAR: Kébar (this work and Adnet et al., 2019); MBK: Mabrouk (see Sweydan et al., 2019), Tunisia; EG: ElGedida (see Strougo et al., 2007); KM: KM11 (see Adnet et al., 2011) MT: Minqar Tabaghbagh (see Zalmout et al., 2012); BQ: Birquet Qarun QS: Quar et Sa; GE: Genahamm Fm.; MI: Midawara FM. from Wadi al Hitan, see Underwood et al., 2011), Egypt; QD: Qa Faydat al Dahikya, Jordania, see Mustafat and Zalmout, 2002).
Fig. 2 in Zoogeography of Elasmobranchs in the Colombian Pacific Ocean and Caribbean Sea
Fig. 2. Relative contribution of each family to the total richness of elasmobranchs for both areas combined (values in brackets) and by area, Colombian Pacific (black bars) and Caribbean (white bars). Values in parentheses indicate the total number of genera (first entry) and species (second entry) per family.
Fig. 1 in Zoogeography of Elasmobranchs in the Colombian Pacific Ocean and Caribbean Sea
Fig. 1. Study area indicating the territorial limits of the Colombian seas. The dotted lines indicate the partitioning into the subareas (Coastal Caribbean, Oceanic Caribbean, Coastal Pacific and Oceanic Pacific) used in this study.
Fig. 4 in Zoogeography of Elasmobranchs in the Colombian Pacific Ocean and Caribbean Sea
Fig. 4. Shark and batoid richness per habitat (a) and per habit (b) in the Colombian Pacific and Caribbean waters, and relationship between the body size of Colombian marine elasmobranchs and its richness (c; values expressed in percentage) and the number of sub-areas (Oceanic Caribbean, Coastal Caribbean, Oceanic Pacific, and Coastal Pacific) occupied by them (d).
Fig. 3 in Zoogeography of Elasmobranchs in the Colombian Pacific Ocean and Caribbean Sea
Fig. 3. Similarity analysis of elasmobranch families per number of species present in the Colombian Caribbean and Pacific. A: Pacific-associated; B: Associated with both areas, Caribbean and Pacific; C: Caribbean-Associated. The subgroups in A (A1, A2) and C (C1, C2) correspond to families exclusive to the area or shared but predominant in the area, respectively.
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