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FIGURE 17. Alpheus richardsoni Yaldwyn, 1971 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 17. Alpheus richardsoni Yaldwyn, 1971: male (cl 8.3 mm) from Loders Creek, Moreton Bay, Queensland, Australia (FLMNH UF 18694) [A]; male (cl 9.3 mm) from Cleveland, Queensland, Australia (OUMNH.ZC 2009-09-0015); A, B— habitus, dorsal view; C—same, lateral view. Photographs by the author.
FIGURE 20 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 20. Alpheus microrhynchus De Man, 1897: male (cl 17.1 mm) from near Ho Chi Minh City, Vietnam (MNHN-IU- 2013-14789) [A–D]; neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193) [E]; A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped in situ, mesial view; D—minor (left) cheliped in situ, lateral view; E—minor (left) cheliped, lateral view. Refer to A for scale in B–D, and to Fig. 19A for scale in E.
FIGURE 11 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 11. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—cephalothorax and chelipeds, dorsal view; B—frontal region, dorsal view; C, D— major (left) cheliped, mesial view, with slightly different angles and illuminations (slightly more ventral and enhancing mesial subdistal ridge on pollex in D). Refer to A for scale.
FIGURE 8 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 8. Alpheus euphrosyne De Man, 1897: male (cl indet.) from Satun, Thailand, in lateral (A) and dorsal (B) views (deposition unknown). Photographs courtesy of Rueangrit Promdam.
FIGURE 2 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 2. Alpheus microrhynchus De Man, 1897: syntype, incomplete male (cl indet., tl 40.0 mm) from Pontianak, Kalimantan, Indonesia (after De Man 1897, as Alpheus sp. and A. microrhynchus, specimen lost) [A–C]; female (cl indet., tl 43.0 mm) from Bangkok, Thailand (after De Man 1898) [D, E]; female (tl 85.0 mm) from Makassar, Sulawesi, Indonesia (after De Man 1911), specimen tentatively assigned to A. takla sp. nov. (see text) [F, G]: A—frontal region, dorsal view; B—male minor (left) chela, dorsal view; C—same, lateral view; D—female minor (right) cheliped, mesial view; E—female major (left) cheliped, lateral view; F—frontal area of carapace, dorsal view; G—telson, dorsal view. Not to scale.
FIGURE 4 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 4. Alpheus euphrosyne De Man, 1897: chelipeds of male (cl indet.) from Sumatra, Indonesia (ZMA Crus.D.202726) showing main features and terminology used in the present study; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (left) cheliped, mesial view; D—same, lateral view. Abbreviations (in alphabetical order): ad—adhesive disks (plaques) on dactylus and palm (small on major chela, highly reduced in minor chela); be—balaeniceps expansion of minor chela dactylus; ddr—dorsal ridge of major chela dactylus (dactylar dorsal ridge); dla—dorsolateral angle of major chela pollex; dlt—dorsolateral tooth of major / minor chela palm; dma—dorsomesial angle of major chela pollex; dmt—distomesial tooth of major / minor chela palm; dp—plunger of major chela dactylus (dactylar plunger); ds—dorsal shoulder of major / minor chela palm; dtg—dorsal transverse groove of major / minor chela palm (also known as dorsal notch); lbd—lateral balaeniceps ridge of minor chela dactylus; lbp—lateral balaeniceps ridge of minor chela pollex; li—linea impressa of major / minor chela palm; lld—lateral longitudinal depression (groove) of major / minor chela palm; ltr—smooth lateral transverse ridge of major chela palm (often poorly developed); mbd—mesial balaeniceps ridge of minor chela dactylus; mbp—mesial balaeniceps ridge of minor chela pollex; mld—mesial longitudinal depression (groove) of major / minor chela palm; msr—mesial subdistal ridge on major chela pollex; mtr—smooth mesial transverse ridge of major chela palm (often poorly developed); pf—fossa (socket) of major chela pollex; vs—ventral shoulder of major / minor chela palm; vtg—ventral transverse groove of major / minor chela palm (also known as ventral notch).
FIGURE 7 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view.
FIGURE 5 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale.
FIGURE 19 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 19. Alpheus microrhynchus De Man, 1897: neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped, lateral view; D—same, mesial view; E—minor (left) cheliped, lateral view; F—same, mesial view. Refer to A for scale.
FIGURE 13 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 13. Alpheus eurydactylus De Man, 1920: female (cl 21.3 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49664): A—habitus, lateral view; B—minor (left) chela in situ, mesial view; C—major (right) cheliped in situ, mesial view; D—same, lateral view. Refer to A for scale.
FIGURE 25 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 25. Alpheus nomurai sp. nov.: holotype, male (cl 8.8 mm) from Tanabe, Wakayama, Japan (MNHN-IU-2018-5590); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) cheliped, lateral view; D—same, mesial view; E— minor (right) cheliped, mesial view; F—same, dorsolateral view; G—same, lateral view. Refer to A for scale.
FIGURE 12 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 12. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) [A–E] and paralectotype, ovigerous female (cl 15.0 mm) [F, G], both from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—major (left) cheliped, lateral view; B—same, chela, latreral view; C—same, fingers open, mesial view (see also Fig. 10C); D—minor chela in situ, dorsal view (cf. Fig. 3H); E—same, lateral view; F—frontal region, lateral view; G—frontal region and chelipeds in situ, dorsal view. Refer to Fig. 11A for scale in A–E.
FIGURE 18. Alpheus richardsoni Yaldwyn, 1971 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 18. Alpheus richardsoni Yaldwyn, 1971: male (cl indet.) from Brunswisk River, New South Wales, Australia [A, B]; male from Parengarenga (Paua), New Zealand [C]; male from Whangarei, New Zealand [D]; A, C—habitus, dorsal view; B—same, lateral view; D—tail fan, dorsal view. Specimens not deposited / deposition unknown. Photographs courtesy of Denis Riek (A, B) and Richard B. Taylor (C, D).
FIGURE 24 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 24. Alpheus cyanoteles Yeo & Ng, 1996: male (cl 9.5 mm) from Sungei Tementang, southern Johor, Malaysia (OUMNH.ZC. 2013.05.6); A—habitus, dorsal view; B—frontal region, dorsal view; C—tailfan, dorsal view; D—chelipeds (major chela widely opened), mesial view; E—H.H. Tan and S. De Grave collecting A. cyanoteles in its natural habitat, a small stream in southern Johor. Photographs by the author.
FIGURE 29 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 29. Alpheus takla sp. nov.: holotype, male (cl 25.5 mm) from Bohol, Philippines (NMCR 99110) [A–G]; two paratypes, males (cl 24.0, 25.0 mm) from the same locality (MNHN-IU-2022-2939) [H–K]; A—habitus, dorsal view; B—same, lateral view; C—frontal region, dorsal view; D—major (left) cheliped, lateral view; E—same, mesial view; F—minor (right) chela, lateral view; G—same, mesial view; H—minor (left) cheliped, lateral view; I—same, chela and carpus, dorsal view; J—major (right) cheliped, mesial view; K—same, dorsal view. Refer to A, B, H, J for scale.
FIGURE 28 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 28. Alpheus nomurai sp. nov.: male (cl not determined) from Yabu River, Nago, Okinawa-jima, Japan (not deposited) [A, B]; male paratype (cl 9.6 mm) from Tanabe, Wakayama, Japan (OUMNH.ZC. 2011.06-006) [C]; four specimens, including male holotype (cl 8.8 mm), from Tanabe, Wakayama, Japan (holotype: MNHN-IU-2018-5590) [D]; A—habitus, lateral view; B–D, same, dorsal view. Photographs courtesy of Tadafumi Maenosono (A, B) and Keiichi Nomura (C, D).
FIGURE 23 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 23. Alpheus cyanoteles Yeo & Ng, 1996: ovigerous female (cl indet.) [A, B] and female (cl indet.) [C, D] from an unknown locality in southern Johor, Malaysia; A—habitus, lateral view; B—same, dorsal view; C—same, lateral view; D—detail of frontal margin of carapace and left antenna, lateral view. Aquarium-kept specimens, probably not deposited. Photographs courtesy of C.H. Wah (A, B) and Heok-Hui Tan (C, D).
FIGURE 1 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 1. Alpheus euphrosyne De Man, 1897: syntype, female (cl 46.0 mm) from Java Sea, Indonesia (after De Man 1897, specimen lost) [A–E]; male (cl indet., tl 34.0 mm) from Bangkok, Thailand (after De Man 1898) [F, G]; A—frontal region, dorsal view; B—female minor (left) cheliped, lateral view; C—second pereiopod, lateral view; D—third pereiopod, lateral view; E—sixth pleonite, telson and uropods, dorsal view; F—male minor (right) cheliped, dorsolateral view; G—same, minor chela, lateral view. Not to scale.
FIGURE 16 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D).
FIGURE 3 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)
FIGURE 3. Three species of Alpheus described by Kemp (1915), De Man (1920) and Schmitt (1926). Alpheus paludicola Kemp, 1915: syntype, male (cl indet., tl 22.0 mm) from Chilika Lake, India [A–G]; A—frontal region, dorsal view; B—scaphocerite of left antenna, dorsal view; C—(left) major cheliped, mesial view; D—same, lateral view; E—minor (right) cheliped, distal portion, mesial view; F—second pereiopod, lateral view; G—telson, dorsal view (after Kemp 1915). Alpheus eurydactylus De Man, 1920: syntypes, male (cl 13.8 mm) [H] and ovigerous female (cl 15.0 mm) [I] from Java, Indonesia (see text for lectotype / paralectotype designation); H—male minor (right) chela, distal portion of palm and fingers, dorsal view; I—female (right) minor chela, lateral view (after De Man 1924). Alpheus langi (Schmitt, 1926), originally described as Crangon langi, presenly junior synonym of Alpheus pontederiae de Rochebrune, 1883: holotype, male (cl 11.0 mm, tl 34.0 mm) from Congo [J–M]: J—frontal region, dorsal view; K—major (left) chela, lateral view; L—minor (right) chela, lateral view; M—second pereiopod, lateral view (after Schmitt 1926). Not to scale.
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