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Figure 63 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 63 Reproductive system, hermaphroditic (female), posterior parts, Wallaconchis melanesiensis. A Indonesia, Halmahera, scale bar 2 mm [5131] (UMIZ 00069) B Vanuatu, scale bar 1 mm [5484] (IM-2013-62406) C Indonesia, Lombok, scale bar 2.4 mm [2963] (UMIZ 00070). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct rs receptaculum seminis sp spermatheca.
Figure 54 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 54 Habitats, Wallaconchis comendadori. A Type locality, Philippines, Bohol, Guindulman, rocks and coral rubble near a few Avicennia trees (st 197) B View of the shore (same locality as A) C Indonesia, Bali, Pemuteran, Labuhan Lalang Harbor, coral rubble, rocks and a few Avicennia trees (st 157) D Indonesia, Lombok, Don Don, old Avicennia forest, with coral rubble (st 149).
Figure 55 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 55 Live specimens, Wallaconchis comendadori. A Dorsal view, 16 mm long [3133], Indonesia, Bali (UMIZ 00064) B Dorsal view, 18 mm long [2985], Indonesia, Lombok (UMIZ 00063) C Dorsal view, 13 mm long [3131], Indonesia, Bali (UMIZ 00064) D Dorsal view, 12 mm long [2725], Indonesia, Ambon (UMIZ 00061) E Dorsal view, 7 mm long [2983], Indonesia, Lombok (UMIZ 00063) F Dorsal view, holotype, 12 mm long [3626], Philippines, Bohol (PNM 041232) G Ventral view, same as B H Ventral view, 15 mm long [2315], Indonesia, Sulawesi (UMIZ 00060) I Ventral view, 22 mm long [3385], Philippines, Bohol (PNM 041234).
Figure 52 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 52 Reproductive system, Wallaconchis achleitneri, Australia, Queensland (MTQ st117). A Hermaphroditic (female), posterior parts, scale bar 1 mm [#4] B Anterior, male copulatory parts, scale bar 1 mm [#1]. Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca v vestibule.
Figure 53 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 53 Penis, Wallaconchis achleitneri, Australia, Queensland (MTQ st117). A Scale bar 100 μm [2645] B Holotype, scale bar 100 μm [3534].
Figure 6 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 6 Geographic distribution of Wallaconchis species. The colors used for each species are the same as those used in phylogenetic trees (Figs 1–5), but note that for W. ater, W. graniferum, and W. melanesiensis, both units #1 and #2 are included within the distribution of the species and that only the color of unit #1 is being used. Black dots show sites sampled by the authors, and grey dots (in Papua New Guinea and the Pacific) show localities where samples were borrowed from a museum collection.
Figure 51 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 51 Radula, Wallaconchis achleitneri, Australia, Queensland (MTQ st117). A Rachidian and innermost lateral teeth, scale bar 6 μm [#1] B Rachidian and innermost lateral teeth, holotype, scale bar 20 μm [3534] C Transition between inner lateral teeth and outer lateral teeth, holotype, scale bar 30 μm [3534] D Outermost lateral teeth, holotype, scale bar 10 μm [3534] E Lateral teeth with basal lateral spine decreasing in size from right to left, scale bar 10 μm [#1] F Left lateral teeth, scale bar 100 μm [#2].
Figure 5 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 5 Maximum parsimony bootstrap consensus tree based on concatenated nuclear ITS1 and ITS2 sequences. Only bootstrap values >70% are indicated. All sequences of Wallaconchis specimens are new. Information on individually-identified specimens can be found in the lists of material examined and in Table 1. The color used for each species is the same as the color used in the other trees (Figs 1–4) and on the map of species distribution (Fig. 6).
Figure 50 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 50 Habitats and live specimens, Wallaconchis achleitneri, Australia, Queensland, Bowen, Doughty Creek. A–B Type locality, sandy area with Avicennia and Rhizophora trees in narrow band by the creek (st 117) C–D Dorsal view, approximately 14 mm long (MTQ st117) E Ventral view, approximately 14 mm long (MTQ st 117).
Figure 9 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 9 Radula, Wallaconchis sinanui, Indonesia, Ambon [5845] (UMIZ 00059). A Rachidian and innermost lateral teeth B Rachidian and innermost lateral teethC Transition between inner lateral teeth and outer lateral teeth D Outermost lateral teeth E Right lateral teeth with basal lateral spine F Outermost lateral teeth. Abbreviations: rt rachidian tooth mc median cusp lc lateral cusp 1llt first left lateral tooth 1rlt first right lateral tooth 2llt second left lateral tooth 2rlt second right lateral tooth bls basal lateral spine (of lateral tooth) nls no lateral spine hlt hook of lateral tooth ollt outermost left lateral tooth. Scale bars: 10 μm (A); 20 μm (B, D, E); 40 μm (C), 10 μm (F).
Figure 8 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 8 External morphology and digestive system, Wallaconchis sinanui, Indonesia, Ambon (A–C) holotype [2737] (UMIZ 00058) (D–F) [2738] (UMIZ 00059). A Dorsal notum B Anterior region, ventral view C Posterior region, ventral view D Digestive system, dorsal view E Digestive system, dorsal view (digestive gland removed) F Digestive system, ventral view (digestive gland removed). Abbreviations: a anus ddg dorsal digestive gland f foot fo female opening hn hyponotum i intestine mo male opening oddg opening of the dorsal lobe of the digestive gland oldg opening of the lateral lobe of the digestive gland opdg opening of the posterior lobe of the digestive gland ol oral lobe ot ocular tentacle pn pneumostome pdg posterior lobe of the digestive gland ppg peripodial groove st1 first stomach chamber st2 second stomach chamber st3 third stomach chamber st4 fourth stomach chamber. Scale bars: 2 mm (A, B, C); 1 mm (D, E, F).
Figure 7 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 7 Habitats and live specimens, Wallaconchis sinanui, Indonesia, Ambon, Lateri. A Type locality, mudflat adjacent to a shallow river through a mangrove (st 128) B Close up of mud with a few individuals of W. sinanui (same locality as A) C Wider view of the mud with many individuals of W. sinanui (same locality as A) D Dorsal view, 9 mm long [2746] (UMIZ 00059) E Holotype, dorsal view, 9 mm long [2737] (UMIZ 00058) F Dorsal view, 9 mm long [2740] (UMIZ 00059).
Figure 48 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 48 Evaginated penis, Wallaconchis graniferum, Philippines (A–C) Luzon [3163] (PNM 041227) (D) Bohol [3636] (PNM 041228) (E) Bohol [3638] (PNM 041225). A Proximal region of evaginated penis with hooks, scale bar 400 μm B Middle region of evaginated penis with long slender hooks on one side, and shorter hooks on the other side, scale bar 400 μm C Distal region of evaginated penis with distal hooks, scale bar 300 μm D Partially evaginated penis with proximal hooks (more distal hooks are inside the rest of the invaginated penis), scale bar 200 μm E Partially evaginated penis with most proximal hooks (more distal hooks are invaginated inside), scale bar 200 μm. Abbreviations (from proximal to distal): ph proximal hooks (first penial region) lh long slender hooks (second penial region) sh short hooks (second penial region) nh no hooks (gap) dh distal hooks (third penial region).
Figure 47 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 47 Reproductive system, anterior, male copulatory parts, Wallaconchis graniferum, Philippines. A Penis with two proximal regions of hooks separated from a third distal region of hooks by a gap with no hooks (the hooks of the first, most proximal region are on the other side of the penis and cannot be seen here), Luzon, scale bar 0.7 mm [3163] (PNM 041227) B Anterior male copulatory parts scale bar 3.6 mm same as A. C Penis partially evaginated inside the vestibule (removed) Bohol scale bar 1 mm [3638] (PNM 041225). Abbreviations: dd deferent duct pr1 first most proximal penial region (with hooks on the other side) pr2 second proximal penial region (with hooks) pr3 third distal penial region (with hooks) ps penial sheath rm retractor muscle sr smooth region of the penis (with no hooks) v vestibule.
Figure 66 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 66 Phylogenetic relationships of Wallaconchis species, with shifts in habitat and character state mapped (for the details on node support and numbers of specimens included in each species, see Figs 1–5). Abbreviations L.Ovi. long oviduct NCP narrow copulatory parts Ovi.M. oviduct membrane WCP wide copulatory parts.
Figure 49 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 49 Penis, Wallaconchis graniferum, not evaginated and opened, Philippines, Bohol (A–B) [5762] (PNM 041226) (C) [5761] (PNM 041228). A Penis invaginated inside the penial sheath opened to reveal regions of penial hooks, scale bar 200 μm B Most proximal region of penis invaginated inside the penial sheath opened to reveal penial hooks, scale bar 200 μm C Penis invaginated inside penial sheath (both opened to reveal the penial hooks), scale bar 0.5 mm. Abbreviations (from proximal to distal): ph proximal hooks (first penial region) lh long slender hooks (second penial region) sh short hooks (second penial region) nh no hooks (gap) dh distal hooks (third penial region).
Figure 46 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 46 Reproductive system, Hermaphroditic (female), posterior parts, Wallaconchis graniferum, Philippines. A Luzon, scale bar 5 mm [3163] (PNM 041227) B Bohol, scale bar 2 mm [3638] (PNM 041225). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct rs receptaculum seminis sp spermatheca.
Figure 45 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 45 Radula, Wallaconchis graniferum, Philippines. A Rachidian and innermost lateral teeth, Bohol, scale bar 20 μm [3636] (PNM 041228) B Rachidian and innermost lateral teeth, Luzon, scale bar 30 μm [3163] (PNM 041227) C Transition between inner lateral teeth and outer lateral teeth, Bohol, scale bar 30 μm [3635] (PNM 041228) D Outermost lateral teeth, Bohol, scale bar 30 μm [3636] (PNM 041228).
Figure 43 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 43 Habitats, Wallaconchis graniferum. A Philippines, Bohol, Loay, mostly sand, very few rocks, and a few small Avicennia trees (st 198) B Close up of sand with shells and Avicennia roots (same locality as A) C Indonesia, Timor, Oesapa, sandy area of mangrove with Avicennia and Sonneratia trees spread out D Close up of sediment and onchidiid tracks (same locality as C).
Figure 44 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 44 Live specimens, Wallaconchis graniferum, Philippines. A Dorsal view, 46 mm long [3163], Luzon (PNM 041227) B Dorsal view, 30 mm long [5902], Indonesia, Timor (UMIZ 00073) C Dorsal view, 28 mm long [3636], Bohol (PNM 041228) D Dorsal view, 26 mm long [3635], Bohol (PNM 041228) E Ventral view, same as A, F Ventral view, same as B.
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