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Figure 28 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 28 Penis, distal part free in the vestibule, Wallaconchis gracile, Indonesia, Bali, scale bar 40 μm [3106] (UMIZ 00056).
Figure 39 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 39 Reproductive system, hermaphroditic (female), posterior parts, Wallaconchis ater, Indonesia A Lombok, scale bar 2 mm [2970] (UMIZ 00043) B Sulawesi, scale bar 3 mm [2220] (UMIZ 00050). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct rs receptaculum seminis sp spermatheca.
Figure 30 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 30 Live specimens, Wallaconchis nangkauriense. A Dorsal view, 28 mm long [3162], Philippines, Luzon (PNM 041204) B Dorsal view, 23 mm long [1786], Indonesia, Sumatra (UMIZ 00012) C Dorsal view, 18 mm long [2156], Indonesia, Sulawesi (UMIZ 00013) D Dorsal view, 23 mm long [1788], Indonesia, Sumatra (UMIZ 00012) E Dorsal view, 34 mm long [3269], Philippines, Bohol (PNM 041206) F Dorsal view, 24 mm long [2161], Indonesia, Sulawesi (UMIZ 00013) G Dorsal view, 32 mm long [3268], Philippines, Bohol (PNM 041206) H Dorsal view, 28 mm long [1787], Indonesia, Sumatra (UMIZ 00012) I Dorsal view, 24 mm long [3217], Philippines, Luzon (PNM 041205) J Ventral view, same as G K Ventral view, 28 mm long [3427], Philippines, Bohol (PNM 041208) L Ventral view, 22 mm long [2731], Indonesia, Ambon (UMIZ 00017).
Figure 42 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 42 Penis, Wallaconchis ater, Indonesia. A Detail of distal portion when fully relaxed (with distal penial projections) and partial view of the coil, Indonesia, Halmahera, scale bar 500 μm [3393] (PNM 041218) B Detail of distal portion when fully relaxed (with distal penial projections) and partial view of the coil, Halmahera, scale bar 500 μm [5125] (UMIZ 00054) C Partial view of the coil (with six loops), Philippines, Bohol, scale bar 500 μm [3266] (PNM 041214) D Cross-section showing the longitudinal fold, Bali, scale bar 300 μm [3137] (UMIZ 00053).
Figure 26 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 26 Radula, Wallaconchis gracile, Indonesia, Bali (A–C) [3106] (UMIZ 00056) (D) [spm #1] (UMIZ 00056). A Rachidian and innermost lateral teeth, scale bar 6 μm B Rachidian and innermost lateral teeth, scale bar 20 μm C Transition between inner lateral teeth and outer lateral teeth, scale bar 30 μm D Left lateral teeth, scale bar 30 μm E Left lateral teeth, gradually increasing in length towards the left, scale bar 40 μm F Outermost lateral teeth, scale bar 20 μm.
Figure 25 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 25 Live specimens, Wallaconchis gracile. A Dorsal view, 19 mm long [3107], Indonesia, Bali (UMIZ 00056) B Dorsal view, 15 mm long [5158], Indonesia, Halmahera (UMIZ 00057) C Dorsal view, 21 mm long [3648], Philippines, Bohol (PNM 041231) D Dorsal view, 19 mm long [2252], Indonesia, Sulawesi (UMIZ 00055) E Dorsal view, 15 mm long [5159], Indonesia, Halmahera (UMIZ 00057) F Dorsal view, 27 mm long [5932], Indonesia, Timor (UMIZ 00074) G Dorsal view, 25 mm long [3652], Philippines, Bohol (PNM 041231) H Frontal view, same as F I Ventral view, 25 mm long [3653], Philippines, Bohol (PNM 041231) J Ventral view, same as F.
Figure 2 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 2 Phylogenetic tree based on concatenated mitochondrial COI, 16S, and 12S sequences. Numbers by branches show bootstrap values (Maximum Likelihood analysis) and posterior probabilities (Bayesian analysis). Only significant numbers (>80% and >0.9) are indicated. All sequences of Wallaconchis specimens are new. Information on individually-identified specimens can be found in the lists of material examined and in Table 1. The color used for each species is the same as the color used in the other trees (Figs 1, 3–5) and on the map of species distribution (Fig. 6).
Figure 20 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 20 Live specimens, Wallaconchis buetschlii. A Dorsal view, 46 mm long [2722], Indonesia, Ambon (UMIZ 00026) B Dorsal view, 31 mm long [2853], Indonesia, Ambon (UMIZ 00027) C Dorsal view, 23 mm long [2249], Indonesia, Sulawesi (UMIZ 00024) D Dorsal view, 30 mm long [2928], Indonesia, Kei Islands (UMIZ 00029) E Dorsal view, 52 mm long [3159], Philippines, Luzon (PNM 041209) F Dorsal view, 36 mm long, [3409], Philippines, Bohol (PNM 041212) G Ventral view, 44 mm long [5898], Indonesia, Timor (UMIZ 00072) H Ventral view, same as F I Ventral view, 43 mm long [2982], Indonesia, Lombok (UMIZ 00030).
Figure 34 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 34 Mangrove habitats, Wallaconchis ater. A–B Indonesia, Bali, Pemuteran, Labuhan Lalang Harbor, coral rubble, rocks and a few Avicennia (st 157) C–D Indonesia, Lombok, Don Don, old Avicennia forest, with coral rubble (st 149) E–F Philippines, Bohol, Mabini, open Avicennia and Sonneratia mangrove with sand and coral rubble (st 196).
Figure 18 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 18 Penis, Wallaconchis uncinus, Indonesia. A Two most distal loops of fully evaginated penis (the loop on the right is proximal to the loop on the left), Halmahera, scale bar 0.5 mm [5079] (UMIZ 00011) B Two most distal loops of (nearly fully evaginated) penis (the loop on the right is proximal to the loop on the left), Ambon, scale bar 300 μm [2843] (UMIZ 00007) C Proximal region of penis, detail view of curved hooks, Sulawesi, scale bar 100 μm [2268] (UMIZ 00005) D Distal region of penis which is not fully evaginated, scale bar 300 μm, same as C. Abbreviations: ch curved hooks ih immature hooks rh rounded hooks.
Figure 33 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 33 Penis, Wallaconchis nangkauriense, Philippines. A Luzon, scale bar 200 μm [3208] (PNM 041205) B Bohol, scale bar 200 μm [3268] (PNM 041206).
Figure 17 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 17 Reproductive system, Wallaconchis uncinus, Indonesia. A Hermaphroditic (female), posterior parts, Sulawesi, scale bar 2 mm [2268] (UMIZ 00005) B Anterior, male copulatory parts, Halmahera, scale bar 3 mm [5079] (UMIZ 00011) C Anterior, male copulatory parts, Bali, scale bar 1 mm [3138] (UMIZ 00008) D Penis (fully evaginated), Halmahera, scale bar 0.5 mm [5079] (UMIZ 00011). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca v vestibule.
Figure 15 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 15 External morphology and digestive system, Wallaconchis uncinus, Indonesia. A Anterior region, ventral view, Sulawesi, scale bar 2 mm [2268] (UMIZ 00005) B Posterior region, ventral view, Sulawesi, scale bar 2 mm [2268] (UMIZ 00005) C Dorsal notum (papillae with dorsal eyes), Halmahera, scale bar 2.6 mm [5056] (UMIZ 00009) D Digestive system, dorsal view, Sulawesi, scale bar 1 mm [2268] (UMIZ 00005). Abbreviations: a anus ddg dorsal lobe of digestive gland f foot fo female opening hn hyponotum i intestine mo male opening ol oral lobe ot ocular tentacle pdg posterior lobe of the digestive gland pn pneumostome ppg peripodial groove st2 stomach chamber 2.
Figure 22 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 22 Reproductive system, Wallaconchis buetschlii. A Hermaphroditic (female), posterior parts, Indonesia, Sulawesi, scale bar 2 mm [2120] (UMIZ 00021) B Anterior, male copulatory parts, Australia, Queensland, scale bar 1 mm [2555] (MTQ st100) C Anterior, male copulatory parts, Indonesia, Kei, scale bar 1 mm [2933] (UMIZ 00029). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland of oviduct fibers attaching to the inner wall of the visceral cavity ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
Figure 14 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 14 Live specimens, Wallaconchis uncinus, Indonesia. A Dorsal view, 25 mm long [5070], Halmahera (UMIZ 00010) B Holotype, dorsal view, 22 mm long [2751], Ambon (UMIZ 00004) C Dorsal view, 30 mm long [2752], Ambon (UMIZ 00006) D Dorsal view, 28 mm long [2843], Ambon (UMIZ 00008) E Dorsal view, 15 mm long [3138], Bali (UMIZ 00005) F Dorsal view, 25 mm long [5080], Halmahera (UMIZ 00011) G Ventral view, same as F H Ventral view, same as B.
Figure 27 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 27 Reproductive system, Wallaconchis gracile, Indonesia. A Hermaphroditic (female), posterior parts, Sulawesi, scale bar 2 mm [2280] (UMIZ 00055) B Anterior, male copulatory parts, Bali, scale bar 2 mm [3106] (UMIZ 00056). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland of oviduct fibers attaching to the inner wall of the visceral cavity ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
Figure 12 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 12 Penis, Wallaconchis sinanui, Indonesia, Ambon. A Penis evaginated outside the male opening, scale bar 200 μm [5846] (UMIZ 00059) B Penis removed from vestibule, scale bar 20 μm [5844] (UMIZ 00059).
Figure 24 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 24 Habitats, Wallaconchis gracile, Indonesia. A–B Bali, Gilimanuk, muddy-sandy beach near a mangrove (st 155) C–D Ternate, sandy mudflat adjacent to a rocky beach (st 220) E–F Timor, sandy area of mangrove with ripples in the sand and near crab pellets (st254).
Figure 13 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 13 Habitats, Wallaconchis uncinus, Indonesia. A, B Sulawesi, Wori, tall mangrove forest of Sonneratia and some Avicennia with a rocky area and dead logs (st 90) C Ambon, Lateri, muddy mangrove with Rhizophora and Avicennia (st 131) D Type locality, Ambon, Lateri, mudflat adjacent to a shallow river through a mangrove (st 128) E Halmahera, Sofifi, mangrove with Sonneratia trees, dense roots and hard mud (st 205) F Ternate, Bastiong, muddy rocks nearby a mangrove patch (st 203).
Figure 11 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 11 Reproductive system, Wallaconchis sinanui, holotype, Indonesia, Ambon [2737] (UMIZ 00058). A Hermaphroditic (female), posterior parts, scale bar 0.8 mm B Anterior, male copulatory parts, scale bar 2 mm. Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca v vestibule.
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