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FIGURE 3 in A new species of Leptolalax (Anura: Megophryidae) from the highest mountain in Indochina

FIGURE 3. Adult female Leptolalax botsfordi sp. nov. AMS R 176540 in life. Note metallic copper flecks, particularly along snout, supratympanic fold and flanks.

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FIGURE 2 in A new species of Leptolalax (Anura: Megophryidae) from the highest mountain in Indochina

FIGURE 2. Leptolalax botsfordi sp. nov. (A) dorsal and (B) ventral view of adult male holotype VNMN 03682 in life, (C) ventral view of gravid female paratype AMS R 176540 in life, (D) posterior view of adult male holotype VNMN 03682 in life, showing large femoral glands in (not to scale), (E) dorsal and (F) ventral view of adult male holotype VNMN 03682 in preservative, and (G) ventral view of adult female paratype AMS R 176540 in preservative. Scale bars = 5 mm.

FIGURE 4 in A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)

FIGURE 4. Ventral view of right hand (A) and foot (B) of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 5 mm.

FIGURE 5 in A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)

FIGURE 5. Sonagram (A, C, E) and wave form (B, D, F) showing advertisement calls of a paratype of Leptolalax sabahmontanus sp. nov. (KUHE 48016) from the type locality, Silau Silau trail along Sg. Silau-Silau, Kinabalu National Park, recorded at an air temperature of 17.2°C.

FIGURE 3 in A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)

FIGURE 3. Dorsal (A) and ventral (B) views of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 10 mm.

FIGURE 2 in A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)

FIGURE 2. Dorsolateral view of a male paratype of Leptolalax sabahmontanus sp. nov. (KUHE 48014) from the type locality, Silau Silau trail along Sg. Silau-Silau, Kinabalu National Park.

FIGURE 1 in A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)

FIGURE 1. Maximum likelihood (ML) tree of a 1907 bp sequence of mitochondrial 12S and 16S rRNA for samples of Leptolalax (for sample details, see text). Numbers above or below branches represent bootstrap supports for ML inference and Bayesian posterior probability (ML-BS/BPP).

FIGURE 9 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 9. Members of Leptolalax applebyi species group. L. applebyi (Ngoc Linh N.P., Kon Tum Province, Vietnam): (A) adult male AMS R173635; (B) AMS R 173777; L. melicus (Virachey N.P., Ratanikiri Province, Cambodia): (C) adult male holotype, MVZ258198; (D) adult male paratype MVZ258199; L. bidoupensis: (E) ZMMU A-5215, Chu Yang Sin N.P., Dak Lak Province, Vietnam; (F) adult female paratype NCSM 77320, Bidoup—Nui Ba N.P., Lam Dong Province, Vietnam (photos A–D and F by J.J.L. Rowley; E—photo by N.A. Poyarkov).

FIGURE 7 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 7. Natural habitat of Leptolalax pyrrhops sp. nov. in Loc Bac woodland, Lam Dong Province, typical breeding site at small forest stream. Photo by E.A. Galoyan.

FIGURE 5 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 5. Paratypes of Leptolalax pyrrhops sp. nov. in preservative. Scale bar 10 mm. Photo by N.A. Poyarkov.

FIGURE 4 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 4. Coloration of the Leptolalax pyrrhops sp. nov. in life: female holotype (ZMMU A-5208; filed number ABV- 00148): (A) dorsolateral view, (B) ventral view; female paratype (ZMMU A-4873, filed number ABV-00158) in life: (C) dorsolateral view, (D) ventral view. Photos by E.A. Galoyan.

FIGURE 3 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 3. Holotype Leptolalax pyrrhops sp. nov. (ZMMU A-5208, female; filed number ABV-00148) in preservative: (A) dorsal view, (B) ventral view (C) head, lateral view, (D) volar view of the left hand, (E) plantar view of the left foot. Scale bar 3 mm. Photos by N.A. Poyarkov.

FIGURE 2 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 2. Bayesian inference dendrogram of Leptolalax derived from the analysis of 1083 bp of 16S rRNA mtDNA gene fragment. Voucher specimen IDs and GenBank accession numbers are given in Table 1. Brachytarsophrys carinensisis used as an outgroup. Numbers near branches represent posterior probability (PP) or bootstrap supportvalues (BS, 1000 replicates) for BI/ML/MP inferences respectively. Color of clade labels for south Indochinese species corresponds to icon colors on Figure 1. Numbers of specimens (No. 1–92) correspond to those in Table 1.

FIGURE 1 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 1. Distribution of Leptolalax species in southern Indochina: Annamite Mountains and adjacent regions. Black dots in the center of an icon indicate the type locality of new species. Locality information. Laos: 1—Khammouan Province, Huey Kun Luang (Ohler et al. 2011); Savannakhet Province: 2—Vilabouli, Sepon Mining Tenement (Rowley et al. 2010); 3—Vilabouli, Sangi (Rowley et al. 2010); Vietnam: 4—Ha Tinh Province (Ohler et al. 2011); Quang Binh Province:5—Phong Nha - Ke Bang NP (Ohler et al. 2011); 6– U Bo (Ohler et al. 2011); 7– Cha Noi (Ohler et al. 2011); 8—Da Nang, Ba Na N.P. (ZMMU A-5221); 9—Quang Nam Province, Song Thanh Proposed Nature Reserve (Rowley et al. 2010a); Kon Tum Province: 10—Dac Lei (Rowley et al. 2010b); 11, 12,—Dac Lei environs, Ngoc Linh Nature Reserve (Rowley et al. 2010b); 13—Dac Lei environs, Ngoc Linh Nature Reserve, Ngoc Linh Mt. (ZISP 12051–12058, ZISP 12091–12093; ZISP 12086–12088; ZISP 12089–12090; ZMMU A-5210; ZMMU A-5217–A-5218); 14—Ngoc Linh Nature Reserve (Rowley et al. 2012); 15—Kon Plong Nature Reserve (ZMMU A-4110; ZMMU A-5213; ZISP 12094–12095); 16—Chu Mom Ray Nature Reserve (Jestrzemski et al. 2013; the authors report two species of Leptolalax, preliminary identified as L. cf. applebyi and L. cf. firthi); Gia Lai Province: 17—Kon Cha Rang Nature Reserve (Inger et al. 1999);18—Binh Dinh Province, An Toan Nature Reserve (Hong Minh Duc, personal communication); 19—Dak Lak Province, Chu Yang Sin National Park (ZMMU A-5215); Lam Dong Province: 20—Bidoup—Nui Ba National Park, Hon Giao Mt. and environs of Giang Ly station (Rowley et al. 2011a; ZMMU A-4717; ZMMU A-4797); 21—Bidoup—Nui Ba National Park, Bidoup Mt. (Paiarkov [Poyarkov] & Vasilieva 2012; ZMMU A-4797; ZMMU A-5211); 22—Loc Bac forestry (this paper); Cambodia: 23, 24—Ratanakiri Province, Virachey National Park (Rowley et al. 2010c); 25—Kampong Speu Province, Phnom Aural Wildlife Sanctuary, Phnom Aural Mt. (Ohler et al. 2002; authors record unidentified Leptolalax larvae and metamorphs).

FIGURE 8 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 8. Advertisement call of the male paratype advertisement calls of Leptolalax pyrrhops sp. nov. (ZMMU A-4873, field number ABV-00213) recordered at air temperature of 25ºC. (A) waveform of high-pass filtered call, (B) wave form and (C) corresponding spectrogram of the same call, unfiltered.

FIGURE 6 in A new species of Leptolalax (Anura: Megophryidae) from the western Langbian Plateau, southern Vietnam

FIGURE 6. Leptolalax pyrrhops sp. nov. in situ: (A) a pair in amplexus (male ZMMU A-4873, field number ABV-00215; female ZISP 12041, field number ABV-00212); (B) calling male (ZMMU A-4873, field number ABV-00213) showing night coloration. Photos by E.A. Galoyan.

FIGURE 6 in A new species of the genus Leptolalax (Anura: Megophryidae) from Mt. Gaoligongshan of western Yunnan Province, China

FIGURE 6. Paratypes of Leptolalax tengchongensis sp. nov. in preservative, note the variation in dorsal and ventral patterns: SYS a004596, SYS a004597, SYS a004599, SYS a004601, SYS a004598, SYS a004602 (from left to right). Photos by J.H. Ya ng.

FIGURE 5 in A new species of the genus Leptolalax (Anura: Megophryidae) from Mt. Gaoligongshan of western Yunnan Province, China

FIGURE 5. Leptolalax tengchongensis sp. nov.: (A) A pair in amplexus (male SYS a004601; female SYS a004602); (B &C) dorsal and ventral views of male paratype SYS a004596 in life; (D & E) dorsal and ventral views of female paratype SYS a004598 in life. Photos by J.H. Yang.

FIGURE 2 in A new species of the genus Leptolalax (Anura: Megophryidae) from Mt. Gaoligongshan of western Yunnan Province, China

FIGURE 2. Bayesian inference (BI) tree derived from partial sequences of the mitochondrial 16S rRNA gene. Numbers above branches are bootstrap support values for maximum likelihood analyses (>70 retained) and numbers below branches indicate Bayesian posterior probabilities (> 70% retained).

FIGURE 3 in A new species of the genus Leptolalax (Anura: Megophryidae) from Mt. Gaoligongshan of western Yunnan Province, China

FIGURE 3. Holotype of Leptolalax tengchongensis sp. nov. (SYS a004600) in preservative: (A) dorsal view; (B) ventral view; (C) plantar view of the left foot; (D) volar view of the left hand; (E) lateral view of head. Photos by J.H. Yang.