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zenodo40/100

FIG. 5 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 5. — Cyathea decrescens Mett. ex Kuhn var. decrescens: A, pinnae abaxially with a fragment of the rachis, sori only partly indicated; B, habit; C, pinnules abaxially with a fragment of the costa; D, pinnule fragment abaxially with sori, note the dense indument of patent, hyaline, multicellular hairs on the costula, veins and indusia; E, scale from the base of the petiole (left: lateral view, not to scale; right: dorsal view); F, basal part of the leaf (from the petiole base up to the first several pinna pairs), lateral view, note gradual transition of the decrescent pinnae into aphlebia; G, a multicellular, hyaline hair, part of this taxon's typical indument; H, leaf scars and trunk surface. A, C, F, Janssen et al. 2541 (P); B, uncollected, photograph at P; D, E, G, Cours 4505 (P); H, Janssen et al. 2578 (P). Scale bars: A, C, H, 1 cm; B, 1 m; D, E, 0.1 cm; F, 5 cm; G, 0.1 mm.

opencc-by-4.0Dec 2008View details →
zenodo40/100

FIG. 16 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 16. — Cyathea costularis Bonap.: A, pinnules abaxially with a fragment of the costa, sori omitted from one pinnule; B, strongly indurated scale from the abaxial face of the base of the petiole; C, habit; D, scale from the lateral face of the upper half of the petiole; E, basal part of the leaf (from the petiole base up to the first pinna pair), dorsal view, lateral view in upper half, one pinna pruned; F, leaf scar and trunk surface; G, basal part of the leaf of a juvenile plant, ventral view, note the strongly reflexed decrescent pinnules and soft scales. A, B, D, E, Rakotondrainibe 3301 (P); C, uncollected, photograph at P; F, Janssen et al. 2489 (P); G, Janssen et al. 2486 (P). Scale bars: A, F, G, 1 cm; B, D, 0.5 cm; C, 1 m; E, 5 cm.

opencc-by-4.0Dec 2008View details →
zenodo40/100

FIG. 13 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 13. — Cyathea appendiculata Baker: A, pinnules abaxially with a fragment of the costa; B, pinnules abaxially with a fragment of the costa, small and weakly auriculate form, veins not indicated, sori only partly indicated; C, pinnules abaxially with a fragment of the costa, crenate form, veins not indicated, sori only partly indicated; D, trunk surface with close standing leaf scars and rudiments of dead petioles; E, habit; F, trunk surface, common aspect with close standing appressed petiole bases in pseudo-whorls; G, pinnae abaxially with a fragment of the rachis, sori only partly indicated; H, basal part of the leaf (from the petiole base up to the second pinna pair), lateral view, scales caducous; I, scale from the base of the petiole. A, G-I, Janssen et al. 2766 (P); B, Janssen et al. 2765 (P); C, F, Janssen et al. 2764 (P); D, Janssen et al. 2755 (P); E, uncollected, photograph at P. Scale bars: A-D, F, H, 1 cm; E, 1 m; G, 5 cm; I, 0.5 cm.

opencc-by-4.0Dec 2008View details →
zenodo40/100

FIG. 8 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 8. — Cyathea ivohibensis (C.Chr.) Janssen & Rakotondr.: A, pinnules abaxially with a fragment of the costa, large form with crenate margins; B, pinnules abaxially with a fragment of the costa, small form with subentire margins; C, habit; D, pinnae abaxially with a fragment of the rachis, sori only partly indicated, large form with crenate margins; E, leaf scars and trunk surface; F, pinnae abaxially with a fragment of the rachis, sori only partly indicated, small form with subentire margins; G, basal part of the leaf (from the petiole base up to the first eight pinna pairs), lateral view; H, scale from the base of the petiole. A, D, Janssen et al. 2795 (P); B, C, E-H, Janssen et al. 2790 (P). Scale bars: A, B, H, 0.5 cm; C, 1 m; D-F, 1 cm; G, 5 cm.

opencc-by-4.0Dec 2008View details →
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FIG. 7 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 7. — Cyathea dilatata Rakotondr. & Janssen: A, pinna abaxially with a fragment of the rachis; B, leaf scar and trunk surface; C, pinnules abaxially with a fragment of the costa; D, scale from the dorso-lateral face of the petiole, about 5 cm above its base, lateral view; E, scale from the dorso-lateral face of the petiole, about 3 cm above its base, dorsal view; F, deltoid scale fragment from the dorsal face of the base of the petiole, dorsal view; G, basal part of the leaf (from the petiole base up to the first several pinna pairs), lateral view, folded upper part in dorsal view, note the damaged (fragmentary) and potentially caducous scales. A-G, Rakotondrainibe 3165 (P). Scale bars A, G, 5 cm; B, C, 1 cm; D-F, 0.1 cm.

opencc-by-4.0Dec 2008View details →
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FIG. 12. — A-H in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 12. — A-H, Cyathea acutula (R.M.Tryon) Janssen & Rakotondr. var. acutula; A, leaf scars and trunk surface; B, pinna abaxially with a fragment of the rachis, sori only partly indicated; C, habit (left: common form; right: form of the Manongarivo massif with dense pinnules near the petiole bases clustered above the trunk apex); D, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view; E, pinnules abaxially with a fragment of the costa, crenate form; F, pinnules abaxially with a fragment of the costa, common form; G, sori with a fragment of the costula carrying acaroid scales; H, scale from the petiole base; I-K, C. acutula var. deltoidea Janssen & Rakotondr.; I, leaf scars and trunk surface; J, basal part of the petiole with contiguous deltoid scales; K, scales from the petiole (left: from the base of the petiole; right: from about 20 cm above the base of the petiole); L, C. acutula var. rufescens Janssen & Rakotondr., scales from the base of the petiole of two different specimens. A, B, D, F-H, Janssen et al. 2849 (P); C, uncollected, photographs at P; E, Rakotondrainibe 4124 (P); I, Janssen et al. 2496 (P); J, K, Rabarimanarivo 74 (P); L (left), Rakotondrainibe et al. 5039bis (P); L (right), Rasolohery 108 (P). Scale bars: A, E, F, I, 1 cm; B, D, J, 5 cm; C, 1 m; G, H, K, L, 0.1 cm.

opencc-by-4.0Dec 2008View details →
zenodo40/100

FIG. 6. — A, B in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 6. — A, B, Cyathea decrescens Mett. ex Kuhn var. cristata Janssen & Rakotondr.; A, pinna abaxially with a fragment of the rachis, sori only partly indicated; B, pinnules abaxially with a fragment of the costula; C-E, C. decrescens var. manongarivensis Janssen & Rakotondr.; C, scale from the base of the petiole (left: lateral view, not to scale; right: dorsal view); D, fragment of the rachis abaxially, note the hyaline hairs being generally more than 10 cells long and having an indurated basal cell; E, pinnules abaxially with a fragment of the costula; F-J, C. decrescens var. quadrata (Baker) Janssen & Rakotondr.; F, pinna abaxially with a fragment of the rachis, sori only partly indicated; G, pinnules abaxially with a fragment of the costula, form of the Andasibe region, cf. "zakamenensis"-morphotype; H, pinnules abaxially with a fragment of the costula, "quadrata"-morphotype; I, pinnules abaxially with a fragment of the costula, "zakamenensis"-morphotype;J, pinnules abaxially with a fragment of the costula, "pilosula"-morphotype. A, B, Rakotondrainibe 3030 (P); C-E, Rakotondrainibe 1711 (P); F, G, Janssen et al. 2578 (P); H, Rasolohery 232 (P); I, van der Werff 12854 (P); J, Capuron 24 (P). Scale bars: A, F, 5 cm; B, E, G-J, 1 cm; C, D, 0.1 cm.

opencc-by-4.0Dec 2008View details →
zenodo40/100

FIG. 15. — A-E in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 15. — A-E, Cyathea conferta Janssen & Rakotondr.; A, pinnules adaxially with a fragment of the costa, note that the auricles overlap the costa; B, sori, note that these are small and very close standing; C, scale from the base of the petiole; D, petiole, note that the filiform scales are restricted to its base; E, pinna abaxially with a fragment of the rachis, sori and veins only partly indicated, note the close standing pinnules; F-J, C. hebes Janssen & Rakotondr.; F, sori, note that these are big and spaced from each other; G, pinnules abaxially with a fragment of the costa; H, basal part of the leaf (from the petiole base up to the first pinna pair), lateral view; I, scale from the base of the petiole; J, pinna abaxially with a fragment of the rachis, sori only partly indicated. A-E, van der Werff et al. 12850 (P); F-J, Janssen et al. 2480 (P). Scale bars: A, C, G, 1 cm; B, F, 0.1 cm; D, E, H, J, 5 cm; I, 0.5 cm. D, E, Drawings by B. Raufeisen.

opencc-by-4.0Dec 2008View details →
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FIG. 3 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 3. — Distribution of tree fern diversity in Madagascar. The number of collected taxa per grid cell of 0.25° × 0.25° is indicated on the varietal level and takes into account all currently recognized taxa of Cyathea sect. Alsophila (this treatment) and Cyathea sect. Gymnosphaera (Janssen & Rakotondrainibe 2007). This grid is superposed on a map indicating the remaining primary vegetation of Madagascar in dark green (DuPuy & Moat 1999) and outlining the bioclimatic regions of Madagascar according to Figure 2. All dark green areas in the humid and subhumid zones present high probabilities for the occurrence of Cyathea. Most specimens and hence most taxa have,however, been collected from the major mountain massifs and other well-known forests in the humid and subhumid zones (names indicated).

opencc-by-4.0Dec 2008View details →
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FIG. 2 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 2. — The bioclimatic regions of Madagascar. This map is a simplified version of the bioclimatic map of Madagascar by Cornet (1974) and redrawn from Schatz (2000). Suitable ecological conditions for the growth of tree ferns are found in the humid, subhumid and montane zones. This map has also been used as a base map for the distribution maps presented in Figures 44 to 46.

opencc-by-4.0Dec 2008View details →
zenodo40/100

FIG. 1 in A revision of the indusiate scaly tree ferns (Cyatheaceae, Cyathea subgen. Alsophila sect. Alsophila) in Madagascar, the Comoros and the Seychelles

FIG. 1. — Spores of Cyathea sect. Alsophila in Madagascar and on the Comoros: A-C, spores of taxa belonging to the bipinnate clade (Janssen et al. 2008) have a perispore formed by dense slender rodlets; D-F, spores of taxa belonging to the tripinnate clade (Janssen et al. 2008) have a perispore with prominent crests; A, C. approximata Bonap.; B, C. kirkii Hook.; C, C. serratifolia Baker; D, C. boivinii Mett. ex Kuhn var. boivinii; E, C. melleri (Baker) Domin; F, C. similis C.Chr., a young spore with a distantly verrucose perispore (Janssen & Rakotondrainibe 2006; Gastony & Tryon 1976). A, Janssen et al. 2449 (P); B, Rakotondrainibe et al. 6751 (P); C, Rakotondrainibe et al. 6230 (P); D, Janssen et al. 2481 (P); E, Janssen et al. 2536 (P); F, Janssen et al. 2576 (P). Scale bars: 10 μm.

opencc-by-4.0Dec 2008View details →
dryad36/100

Antheridiogen controls spatial dynamics of sex‐expression in naturally occurring gametophytes of the tree fern Cyathea multiflora

<p>Background and aims: Antheridiogen systems are an important and widespread mechanism by which sex expression is controlled and genetic diversity maintained in fern gametophyte populations. However, antheridiogens have rarely been studied outside of the laboratory and little is known about their function in natural populations. Combining predictions based on field and laboratory study, we document the sexual structure of tree fern gametophyte populations and test the effects of antheridiogen <em>in situ</em>. </p> <p>Key results: In all populations, sex ratios indirectly indicate antheridiogen activity. No hermaphroditic gametophytes were identified in any population. Female gametophytes are randomly distributed in each population while male gametophytes tend to cluster. In two of the populations, male sex expression is spatially dependent on females, providing direct evidence of antheridiogen function <em>in situ</em>.</p> <p>Conclusions: This study provides the first documentation of spatial sex expression in natural gametophyte populations of an antheridiogen-producing tree fern species. The profound impact of antheridiogen on gametophyte sex expression in field settings suggests this system is intimately tied to mating systems, fitness, and genetic diversity in <em>Cyathea multiflora</em>.</p>

opencc-zeroJul 2022View details →
dryad36/100

Antheridiogen controls spatial dynamics of sex‐expression in naturally occurring gametophytes of the tree fern Cyathea multiflora

Open the record for dataset details and reuse information.

publicJul 2022View details →
dryad32/100

Data from: Historical reconstruction of climatic and elevation preferences and the evolution of cloud forest-adapted tree ferns in Mesoamerica

Background. Cloud forest, characterized by a persistent, frequent or seasonal low-level cloud cover and a fragmented distribution, is one of the most threatened habitats especially in the Neotropics. Tree ferns are among the most conspicuous elements in these forests and ferns are restricted to regions in which minimum temperatures rarely drop below freezing and rainfall is high and evenly distributed around the year. Current phylogeographic data suggest that some of the cloud forest-adapted species remained in situ or expanded to the lowlands during glacial cycles and contracted allopatrically during the interglacials. Although the observed genetic signals of population size changes of cloud forest-adapted species including tree ferns correspond to predicted changes by Pleistocene climate change dynamics, the observed patterns of intraspecific lineage divergence showed temporal incongruence. Methods. Here we combined phylogenetic analyses, ancestral area reconstruction, and divergence time estimates with climatic and altitudinal data (environmental space) for phenotypic traits of tree fern species to make inferences about evolutionary processes in deep time. We used phylogenetic Bayesian inference and geographic and altitudinal distribution of tree ferns to investigate the ancestral area and elevation and environmental preferences of Mesoamerican tree ferns. The phylogeny was then used to estimate divergence times and ask whether the ancestral area and elevation and environmental shifts were linked to climatic events and historical climatic preferences. Results. Bayesian trees retrieved Cyathea, Alsophila, Gymnosphaera and Sphaeropteris in monophyletic clades. Splits for species in these genera found in the Mesoamerican cloud forests are recent, from the Neogene to the Quaternary. Australia was identified as the ancestral area for the clades of these genera, except for Gymnosphaera that was Mesoamerica. Climate tolerance was not divergent from hypothesized ancestors for the four most significant variables or elevation. For elevational shifts we found repeated changes from low to high elevations. Conclusions. Our data suggest that representatives of Cyatheaceae main lineages migrated from Australia to Mesoamerican cloud forests in different times and have persisted in these environmentally unstable areas but extant species diverged recently from their ancestors.

opencc-zeroDec 2015View details →
zenodo32/100

FIGURE 3 in Cyathea fabiolae (Cyatheaceae, Polypodiopsida), a new scaly tree fern from the northern Andes

FIGURE 3. Croziers of A) C. fabiolae, B) C. tortuosa, and C) C. brunnescens. Note lack of villosity, near lack of scurf, and orange-brown scales in C. fabiolae in contrast to C. tortuosa (villous axes abaxially, brown scales) and C. brunnescens (markedly scurfy rachis, brown scales).

opennotspecifiedJun 2022View details →
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FIGURE 2 in Cyathea fabiolae (Cyatheaceae, Polypodiopsida), a new scaly tree fern from the northern Andes

FIGURE 2. Known localities (black circle, type locality; open circles, localities known from photographic evidence; gray circle, paratype locality) of Cyathea fabiolae along the Amazonian slope of the Andes in Peru and Ecuador. Photographic records: a. Pacto Sumaco, Napo: -0.618, -77.598; b. Mera, Pastaza: -1.438, -78.130; c. Quimi, Morona Santiago: -3.489, -78.417; by Adrian Tejedor, June, 2016.

opennotspecifiedJun 2022View details →
zenodo32/100

FIGURE 1. Cyathea fabiolae. A in Cyathea fabiolae (Cyatheaceae, Polypodiopsida), a new scaly tree fern from the northern Andes

FIGURE 1. Cyathea fabiolae. A, pinna, silhouette; B, petiole scale; C, fertile pinnule abaxially; D, fertile segment, abaxially, from G. Calatayud 6220.

opennotspecifiedJun 2022View details →
zenodo32/100

FIGURE 4. A–E in A new hybrid and further taxonomic notes on Brazilian tree ferns (Cyatheaceae)

FIGURE 4. A–E. Cyathea mexiae (Schwartsburd 2983 [VIC]): A. Base of petiole, showing aculei, scales, and scurf. B. Petiolar aculei and scurf. C. Medial pinna. D. Rachis, pinna-rachis, and pinnules, abaxially, showing sericeous axes and sessile pinnules. E. Segment, abaxially, showing sori, hairs, and bullate scales on costule. F–J. Alsophila salvinii (Schwartsburd 3369 [VIC]): F. Base of petiole, showing scales and broken-off aphlebiae. G. Entire aphlebia with laminar expansions. H. Broken-off aphlebia without laminar expansion. I. Medial pinna. J. Rachis, pinna-rachis, and pinnules, abaxially, showing scurf, scales, and segments.

opennotspecifiedOct 2015View details →
zenodo32/100

FIGURE 3. A–C in A new hybrid and further taxonomic notes on Brazilian tree ferns (Cyatheaceae)

FIGURE 3. A–C. VIC: Cyathea corcovadensis s. str. (Silva 1593): A. Departure of pinnule (tangential section) from pinna-rachis (cross section), showing the schlerenquima ring internally on pinnule articulation (arrows). B. Mesophyll in cross section. C. Stomata from abaxial surface of lamina. D–F. Cyathea ×stella-matutina (Schwartsburd 3303 [VIC]): D. Departure of pinnule (tangential section) from pinna-rachis (cross section), showing the sclerenchyma ring internally on pinnule articulation (arrows). E. Mesophyll in cross section (asterisk showing intercellular protuberances). F. Stomata from abaxial surface of lamina. G–I. Cyathea microdonta (Schwartsburd 3300 [VIC]): G. Departure of pinnule (tangential section) from pinna-rachis (cross section), showing the absence of a sclerenchyma ring internally. H. Mesophyll in cross section (asterisk showing intercellular protuberances). I. Stomata from abaxial surface of lamina.

opennotspecifiedOct 2015View details →
zenodo32/100

FIGURE 5. A in A new hybrid and further taxonomic notes on Brazilian tree ferns (Cyatheaceae)

FIGURE 5. A. Petiolar scale of Cyathea mexiae (Schwartsburd 2983 [VIC]). B. Petiolar scales of Alsophila salvinii (Schwartsburd 3369 [VIC]). Bar of 0.5 cm.

opennotspecifiedOct 2015View details →

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