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Figure 5. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 5. A, holotype of Hemiphyllodactyus tehtarik sp. nov. (LSUHC 10904) from Gunung Tebu, Terengganu, Malaysia; photo by L. Grismer. B, adult female Hemiphyllodactylus larutensis (LSUHC 11294) from Bukit Larut, Perak, Malaysia; photo by L. Grismer. C, adult male holotype of Hemiphyllodactylus larutensis (holotype BM 1901.3.20.2) from Bukit Larut, Perak, Malaysia; photo by G. Zug. D, adult male Hemiphylodactylus larutensis (LSUHC 11298) from Bukit Larut, Perak, Malaysia; photo by L. Grismer. E, adult male Hemiphyllodactylus harterti (LSUHC 10383) from Bukit Larut, Perak, Malaysia; photo by L. Grismer. F, adult female Hemiphyllodactylus harterti (holotype ZMB 1360) from Gunung Hijau, Perak, Peninsular Malaysia; photo by M.-O. Rödel.

Figure 7. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 7. A, adult male Hemiphyllodactylus sp. nov. 2 (LSUHC 5759) from Pulau Sibu, Johor, Malaysia; photo by L. Grismer. B, adult female Hemiphyllodactylus sp. nov. 3 (MVZ 239345) from Pulau Enggano, Sumatra, Indonesia; photo by J. McGuire.

Figure 8. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 8. A, adult male Hemiphyllodactylus sp. nov. 5 (PNM 7666) from Palaui Island; photo by R.M. Brown. B, adult male Hemiphyllodactylus insularis (KU uncatalogued) from Pasonanca, Zamboanga, Western Mindanao, Philippines. C, adult female Hemiphyllodactylus sp. nov. 4 (KU 331843) from Mount Lanoy, Municipality of Argao, Cebu Province, Philippines; photo by R.M. Brown.

Figure 2 in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 2. Distribution of the species of Hemiphyllodactylus examined (see the Appendix for a list of the Hemiphyllodactylus typus examined). Hemiphyllodactylus aurantiacus from Yercaud, Salem District (Shevaroyan Hill Range), Tamil Nadu, India; Hemiphyllodactylus ganoklonis from Palau Ngercheu; Hemiphyllodactylus harterti and Hemiphyllodactylus larutensis from Bukit Larut, Perak, Malaysia; Hemiphyllodactylus insularis from the Municipality of Pasonaca, Zamboanga City, Zamboanga City Province, Philippines; Hemiphyllodactylus tehtarik from Gunung Tebu, Terengganu, Malaysia; Hemiphyllodactylus titiwangsaensis from the Cameron Highlands, Pahang, Malaysia; Hemiphyllodactylus sp. nov. 1 from Fraser's Hill and the Genting Highlands, Pahang; Hemiphyllodactylus sp. nov. 2 from Pulau Sibu, Johor, Malaysia; Hemiphyllodactylus sp. nov. 3 from Pulau Enggano, Bengkulu Province, Sumatra, Indonesia; Hemiphyllodactylus sp. nov. 4 from Mount Lantoy, Municipality of Argao, Cebu Province, Cebu, Philippines; Hemiphyllodactylus sp. nov. 5 from Pulaui Island, Barangay Palaui, Municipality of Santa Ana, Cagayan Province, Philippines; Hemiphyllodactylus sp. nov. 6 from Barangay Kaim, Municipality of San Francisco, Agusan del Sur Province, Mindanao, Philippines; Hemiphyllodactylus sp. nov. 7 from Chiang Mai, Chiang Mai Province, Thailand; Hemiphyllodactylus sp. nov. 8 from Pyin-Oo-Lwin, Mandalay Division, Myanmar; Hemiphyllodactylus sp. nov. 9 from Pakxong District, Champasak Province, Laos; Hemiphyllodacylus zugi from Vinh Phu, Vietnam; and Hemiphyllodactylus sp. nov. 10 from Ba Na-Nui Chua Nature Reserve, Da Nang Province, Vietnam. Localities of Hemiphyllodactylus dushnanesis (closed yellow circles), Hemiphyllodactylus jinpingensis (closed blue circles), Hemiphyllodactylus longlingensis (closed orange circles), and Hemiphyllodactylus yunnanensis (closed green circles) follow Zhou et al. (1981), except for the southern specimen of H. dushanensis from Vinh Phuc, Vinh Phuc Province, Vietnam reported here.

Figure 1 in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 1. Maximum-likelihood phylogram (–lnL 22097.690183) of the genus Hemiphyllodactylus Bleeker, 1860 with Bayesian posterior probabilities and maximum-likelihood bootstrap values, respectively.

Figure 6. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 6. A, adult male Hemiphyllodactylus titiwangsaensis (LSUDPC 4506) from Cameron Highlands, Pahang, Peninsular Malaysia. B, adult female Hemiphyllodactylus sp. nov. 1 from Genting Highlands, Pahang, Peninsular Malaysia. C, adult male Hemiphyllodactylus sp. nov. 1 from Fraser's Hill, Pahang, Peninsular Malaysia.

Figure 10. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 10. A, adult Hemiphyllodactylus auriantiacus (LSUDPC 7006) from Yercaud, Salem District (Shevaroyan Hill Range), Tamil Nadu, India. B, adult Hemphyllodactylus sp. nov. 10 (ITBCZ 2450). C, male Hemphyllodactylus sp. nov. 10 (LSUDPC 6675) from Ba Na-Nui Chua Nature Reserve, Hoa Vang District, Da Nang City Province, Vietnam; photos B and C by Ngo Van Tri. D, adult male Hemiphyllodactylus zugi (LSUDPC 8089) from Cao Bang Province, Vietnam; Photo by T. Q. Nguyen and T. Lehmann.

Figure 18 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 18. Sabellomma cupoculata sp. nov., colour micrographs. A–D, live specimens. A, anterior end, ventral view. B, specimen missing posterior abdominal chaetigers, lateral view. C, radiolar crown and anterior thoracic chaetigers, lateral view. D, same, dorsal view. E–M, preserved specimens. E–G, detail of radiolar eyes along radiolar lateral margins. H, base of crown, ventral view, with crown opened showing ventral lips, dorsal lips with long radiolar appendages, and anterior thoracic chaetigers with ventral shields in contact with neuropodial tori and a dark pigment spot in between. I, J, half of radiolar crown showing dorsal lips with long radiolar appendages and pinnular appendages (arrow); dyed with methylene blue. K, thoracic chaetigers, holotype, lateral view, lacking inter-ramal eyes. L, thoracic chaetigers and collar. M, posterior thoracic and anterior abdominal chaetigers, lacking conspicuous inter-ramal eyes. A–D, AM W.37060; E–H, AM W.37060; I, J: AM W.37029; K, AM W.47193 (holotype); L, M, AM W.47189.

Figure 17 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 17. Parasabella sp. cf. Parasabella rugosa, scanning electron microscope photographs. A, anterior thoracic chaetigers and base of radiolar crown, dorsal view, showing large collar dorsal margins. B, midthoracic parapodium with elongate, narrowly hooded (arrow) superior chaetae and broadly hooded (type B) in inferior group. C, detail of inferior thoracic broadly hooded chaetae of type B. D, thoracic uncini. E, companion chaetae, lateral view. F, companion chaetae, frontal view. G, midabdominal neurochaetae narrowly hooded. H, abdominal uncini. I, posterior abdominal chaetigers and (damaged) pygidium. A–I, AM W.36431.

Figure 15 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 15. Parasabella sp. cf. Parasabella japonica, scanning electron microscope photographs. A, anterior thoracic chaetigers and base of radiolar crown, ventral view, showing ventral shields separated from neuropodial tori, and ventral lappets. B, anterior chaetigers, lateral view. C, same, dorsal view. D, midthoracic parapodium, elongate, narrowly hooded superi- or chaetae and broadly hooded type B chaetae in inferior group. E, detail of inferior, broadly hooded type B chaeta. F, thoracic uncini and companion chaetae. G, thoracic uncini, detail. H, companion chaetae showing the laterally compressed hood. I, midabdominal narrowly hooded neurochaetae. J, abdominal uncini. A–J, AM W.36450.

Figure 16 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 16. Parasabella sp. cf. Parasabella rugosa, colour micrographs of preserved specimens. A, anterior thoracic chaetigers, ventral view, showing posterior peristomial ring collar, and ventral shields in contact with neuropodial tori. B, complete thorax, lateral view. C, same, dorsal view. D, posterior abdominal chaetigers and pygidium, ventral view. A, NMV F.108844; B–D, AM W.36431.

Figure 12 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 12. Parasabella crassichaetae sp. nov. complex, colour micrographs of preserved specimens. A, whole specimen, lateral view. B, anterior chaetigers, showing the base of crown and the collar margins. C, specimens with detached crown showing the peristomial eyes (arrows) near the insertion site of the radiolar lobes. A–B, AM W.31103; C, AM W.37028.

Figure 10 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 10. Parasabella bioculata sp. nov. colour micrographs. A–C, live specimen. A, radiolar crown and anterior thoracic chaetigers, lateral view. B, same, dorsal view. C, whole specimens with opened radiolar crown. D–I, preserved specimens. D, anterior thoracic chaetigers and base of radiolar crown, lateral view. E, same, ventrolateral view. F, specimen with detached crown, dorsal view. G, tip of lateral radioles showing the subdistal radiolar eyes. H, arrangement of radiolar eyes on lateral radiole. I, magnified detail showing cup-shaped radiolar eye. A–C, AM W.37053; D, E, G, H, AM W.46997; F, I, AM W.37056.

Figure 9 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 9. Parasabella sp. cf. Parasabella aulaconota scanning electron microscope photographs. A, anterior thoracic chaetigers and base of radiolar crown, ventral view, with ventral shields in contact with neuropodial tori. B, same, showing the collar, lateral view. C, collar dorsal margins, lateral view. D, anterior thoracic chaetigers and base of radiolar crown, showing collar dorsal margins, dorsal view. E, collar chaetae, elongate, narrowly hooded. F, thoracic elongate, narrowly hooded superior notochaetae and inferior, broadly hooded (type B) chaetae. G, detail of inferior thoracic chaetae. H, I, thoracic uncini from specimens collected in temperate and tropical sites, respectively. J, companion chaetae. K, complete row of uncini in abdominal notopodia, showing the range in size from dorsal- to ventral-most uncini. L, midabdominal narrowly hooded neurochaetae. M, abdominal uncini. B, C, F, H, K–M, AM W.22017; A, D, E, G, I, J, AM W.22480.

Figure 11 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 11. Parasabella bioculata sp. nov. scanning electron microscope photographs. A, anterior thoracic chaetigers and base of radiolar crown, ventral view, showing ventral shields separated from neuropodial tori. B, same, lateral view. C, same, dorsal view. D, inside of radiolar crown, frontal view, showing dorsal lips and long radiolar appendages. E, elongate, narrowly hooded collar chaetae. F, midthoracic chaetiger, elongate, narrowly hooded superior thoracic chaetae, and inferior, broadly hooded type B chaetae. G, thoracic uncini. H, companion chaetae. I, midabdominal, narrowly hooded neurochaetae. J, abdominal uncini. A–J, AM W.36449.

Figure 8 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 8. Parasabella sp. cf. Parasabella aulaconota colour micrographs. A, B, live specimen, anterior end, dorsal view. C–F, preserved specimen. C, detail of remaining pigmentation on radioles. D, anterior end showing collar margins, ventral lappets, ventral shields, and neuropodial tori. E, anterior end, lateral view. F, anterior end, dorsal view, showing dorsal collar margins. A, B, AM W.35612; C, AM W.47006; D–F, AM W.22480.

Figure 6. Parasabella aberrans spp. complex, colour micrographs. A in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 6. Parasabella aberrans spp. complex, colour micrographs. A, crown and anterior thoracic segments, lateral view. B, thoracic chaetigers showing ventral shields in contact with neuropodial tori. C, anterior thoracic chaetigers and base of crown, showing collar ventral lappets and shape of ventral shields. D, anterior thoracic segments in dorsal view, showing the stiff fleshy swelling separated by the faecal groove. E, same. F, fleshy swelling continuous across dorsum. A, AM W.36946; B, D, AM W.36935; C, E, AM W.36430; F, AM W.32018.

Figure 7. Parasabella aberrans spp. complex, scanning electron microscope photographs. A in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 7. Parasabella aberrans spp. complex, scanning electron microscope photographs. A, anterior thoracic chaetigers and base of radiolar crown, ventral view. B, same, showing collar, lateral view. C, collar dorsal margins, lateral view. D, collar dorsal margins and fleshy swelling divided in two by faecal groove, dorsal view. E, fleshy swelling continuous across dorsum. F, elongate, narrowly hooded collar chaetae. G, inferior thoracic notochaetae (broadly hooded, type B). H, thoracic uncini and companion chaetae. I, J, thoracic uncini from specimens with continuous and divided dorsal swellings, respectively. K, L, companion chaetae from specimens with continuous and divided dorsal swelling, respectively. M, midabdominal, narrowly hooded neurochaetae. N, abdominal uncini. O, posterior abdominal chaetigers and pygidium, ventral view. A, B, D, F, G, H, M, N, AM W.36935; C, E, J, L, O, AM W.32018; I, K, AM W.36430.

Figure 4 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 4. Cross-sections of radioles near the base showing the supporting cartilaginous vacuolated cells in the rachis (with grey nuclei), surrounded by an extracellular cartilaginous sheath (white) and covered by columnar epithelium (grey). Upper side of drawings are the outer margin of radioles; on the bottom incomplete pinnules are sketched with a blood vessel (black) in between. A, Parasabella aberrans spp. complex. B, Parasabella sp. cf. Parasabella aulaconota. C, Parasabella fullo. D, Parasabella bioculata sp. nov. E, Parasabella crassichaetae sp. nov. complex. F, Parasabella sp. cf. Parasabella japonica. G, Parasabella sp. cf. Parasabella rugosa. H, Sabellomma cupoculata sp. nov. A, AM W.36947; B, AMW.47009; C, ZMB 5731; D, AM W.46840; E, AM W.47181; F, AM W.36450; G, AM W.36431; H, AM W.47189.

Figure 1 in Integrative taxonomy of Parasabella and Sabellomma (Sabellidae: Annelida) from Australia: description of new species, indication of cryptic diversity, and translocation of some species out of their natural distribution range

Figure 1. Maps with collecting sites in Australia. A, Parasabella aberrans spp. complex, Parasabella sp. cf. Parasabella aulaconota, Parasabella bioculata sp. nov. B, Parasabella crassichaetae sp. nov. complex, Parasabella sp. cf. Parasabella japonica, Parasabella sp. cf. Parasabella rugosa, Sabellomma cupoculata sp. nov.