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Figure 38 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 38 - Coryphellina lotos sp. n. Japan, Pacific Honshu, Osezaki. ZMMU Op-515, living specimen 15.5 mm in length: A dorsal view B lateral view C ventral view D details of cerata E rhinophores, close up F jaw SEM G details of masticatory process of jaw, SEM H details of masticatory process of jaw, light microscopy I radular teeth, posterior part, SEM J reproductive system, light microscopy. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nd discontinuous notal edge pvd prostatic vas deferens r rhinophores rsp proximal receptaculum seminis. Scale bars: F = 100 μm; G = 20 μm; I =50 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 41 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 41 - Pacifia amica sp. n., NE Pacific, Port Orchard, holotype ZMMU Op-614: A live, dorsal view, 6.5 mm in length B lateral view C details of cerata D jaws, lateral view E details of masticatory processes of jaw F posterior rows of radula, SEM G anterior rows of radula H details of posterior radular teeth I reproductive system in situ, light microscopy. Abbreviations: am ampulla fgm female gland mass go genital opening pr prostate ps penial sheath rsd distal receptaculum seminis rsp proximal receptaculum seminis vd vas deferens. Scale bars: D = 100 μm E =2 μm; F, G, H = 10 μm. Photos of living specimens by K. Fletcher, other photos and SEM images by A.V. Martynov.

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Figure 39 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 39 - Coryphellina rubrolineata O'Donoghue, 1929. Vietnam, Nhatrang Bay. ZMMU Op-132, living specimen 15 mm in length: A dorsal view B ventral view (fixed) C details of cerata D dissected anterior part (pharynx removed) and rhinophores, SEM, scale bar 1 mm E rhinophores, close up, SEM F jaw, SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I, J reproductive system, light microscopy K bilobed proximal receptaculum seminis. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nd discontinuous notal edge ogp oral gland penetrating into basis of cerata ps penial sheath rsp proximal receptaculum seminis pvd prostatic vas deferens r rhinophores. Scale bars: D, F = 300 μm; G, H = 30 μm. Photos of living specimens by O.V. Savinkin, other photos and SEM images by A.V. Martynov.

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Figure 36 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 36 - Carronella pellucida (Alder & Hancock, 1843), comb. n. Norwegian Sea, Gulen Dive Center. ZMMU Op-513, living specimen 18.5 mm in length: A dorsal view B lateral view C ventral view D details of cerata E dissected anterior part (pharynx removed) and rhinophores, SEM F jaw SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I reproductive system, SEM J reproductive system, light microscopy K dissected penial sheath and penis, SEM. Abbreviations: a anus am ampulla ce ceratal elevations fgm female gland mass go genital opening nd discontinuous notal edge ogp oral gland penetrating into basis of cerata r rhinophores p penis ps penial sheath pvd prostatic vas deferens rsd distal receptaculum seminis rsp proximal receptaculum seminis. Scale bars: E = 1 mm; F, K = 300 μm; G = 30 μm; H = 30 μm; I = 1 mm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 34 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 34 - Schematic outline of the reproductive systems of the taxa of the family Flabellinidae integrated with molecular phylogenetic data. Colour indication of reproductive system characters: ampulla – green; body wall – gray; distal receptaculum seminis – red; female gland mass – yellow; female genital opening – orange; penis and male genital opening – dark blue; penial sheath – pale blue; prostatic vas deferens – turquoise; proximal receptaculum seminis – pink.

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Figure 30 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 30 - Microchlamylla gracilis gracilis (Alder & Hancock, 1844), comb. n. Norwegian Sea, Gulen Dive Center. ZMMU Op-502, living specimen 23 mm length: A dorsal view B lateral view C ventral view D details of cerata E dissected anterior part (pharynx removed) and rhinophores, SEM F jaw, SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I reproductive system, SEM. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nd discontinuous notal edge og oral glands ps penial sheath pvd prostatic vas deferens r rhinophores rsd distal receptaculum seminis. Scale bars: E, I = 1 mm; F = 300 μm; G, H = 30 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 35 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 35 - Carronella enne sp. n. NE Atlantic, off Ireland. ZMMU Op-526, specimen 5 mm (fixed) in length: A dorsal view (live); B details of cerata C rhinophore D dorsal view (fixed) E ventro-lateral (fixed) F jaw, SEM G details of masticatory process of jaw, SEM H details of masticatory process of jaw, light microscopy I radular teeth, posterior part, SEM J reproductive system, light microscopy. Abbreviations: a anus am ampulla ce ceratal elevations fgm female gland mass g gonad go genital opening nd discontinuous notal edge ps penial sheath pvd prostatic vas deferens r rhinophores rsd distal receptaculum seminis rsp proximal receptaculum seminis. Scale bars: F = 300 μm; G, I = 30 μm. Photos of living specimens by E. Schwabe, other photos and SEM images by A.V. Martynov.

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Figure 28 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 28 - Himatina trophina (Bergh, 1894). North West Pacific, Kamchatka. ZMMU Op-29, living specimen 32 mm length: A dorsal view B ventral view ZMMU Op-532, fixed specimen 27 mm length C lateral view D ventral view E details of cerata ZMMU Op-29 F dissected anterior part (pharynx removed) and rhinophores, SEM G jaw, SEM H details of masticatory process of jaw SEM I radular teeth, posterior part, SEM J reproductive system, light microscopy K reproductive system SEM L penis, SEM. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nc continuous notal edge p penis ps penial sheath r rhinophores rsd distal receptaculum seminis rsp proximal receptaculum seminis. Scale bars: G, K, L = 1 mm; H = 30 μm; I =10 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 37 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 37 - Coryphellina exoptata (Gosliner & Willan, 1991), comb. n. Vietnam, Nhatrang Bay. ZMMU Op-116, living specimen 9 mm in length: A dorsal view B ventral view (fixed) C details of cerata D dissected anterior part (pharynx removed) and rhinophores, SEM E jaw, SEM F details of masticatory process of jaw, SEM, scale bar = 30 μm G radular teeth, posterior part, SEM H, I reproductive system, light microscopy. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nd discontinuous notal edge ogp oral gland penetrating into basis of cerata ps penial sheath r rhinophores rsp proximal receptaculum seminis pvd prostatic vas deferens. Scale bars: D = 1 mm; E = 300 μm; F = 30 μm. Photos of living specimens by O.V. Savinkin, other photos and SEM images by A.V. Martynov.

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Figure 27 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 27 - Gulenia orjani sp. n. Norwegian Sea, Gulen Dive Center. Holotype NTNU-VM-72482, living specimen 22 mm length: A dorsal view B lateral view C ventral view F details of cerata D Paratype ZMMU Op-538, living specimen 19 mm length, dorsal view E living specimens (not collected) on a hydroid colony G radular teeth, posterior part, SEM ZMMU Op-407 H jaw, SEM ZMMU Op-407 I details of masticatory process of jaw, SEM ZMMU Op-407 J reproductive system, SEM ZMMU Op-409 K penis, SEM ZMMU Op-409 L seminal reservoirs ZMMU Op-409. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nc continuous notal edge p penis ps penial sheath pvd prostatic vas deferens rsd distal receptaculum seminis rsp proximal receptaculum seminis. Scale bars: G = 30 μm; H, J, K, L = 300 μm; I =30 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 24 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 24 - Fjordia lineata (Lovén, 1846), comb. n. A neotype 11 mm length (fixed), dorsal view (live), NTNU-VM-72483, Idefjorden, Sverige B living specimen 21 mm length dorsal view (ZMMU Op-507) F same, details of cerata C ZMMU Op-508, lateral view D same, ventral view E living specimens (not collected) on a hydroid colony G jaw, SEM (ZMMU Op-507) H details of masticatory process of jaw, SEM (ZMMU Op-507) I radular teeth, posterior part, SEM (ZMMU Op-508) J reproductive system, SEM, scale bar = 1 mm (ZMMU Op-507) K penis, SEM (ZMMU Op-507) L seminal reservoirs (ZMMU Op-507). Abbreviations: a anus am ampulla rsd distal receptaculum seminis fgm female gland mass go genital opening p penis ps penial sheath pvd prostatic vas deferens rsp proximal receptaculum seminis. Scale bars: G, I, K = 300 μm H = 10 μm; J = 1 mm; L = 100 μm. Photos of living specimens by T.A. Korshunova (A, C, D), and B. Picton (B, E), SEM images by A.V. Martynov.

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Figure 4 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 4 - Chlamylla borealis orientalis (Volodchenko, 1941), comb. n. ZMMU Op-478. The Sea of Japan, Vostok Bay, 15 mm in length: A living animal, dorsal view B fixed animal, ventral view of anterior part C fixed animal, lateral view D dissected anterior part (pharynx removed) and rhinophores, SEM E jaw, SEM F details of masticatory process of jaw, SEM G details of masticatory process of jaw, light microscopy H radular teeth, posterior part, SEM I reproductive system, SEM. Abbreviations: a anus gf genital fold fgm female gland mass nc continuous notal edge pc penial collar (external) pr prostate ps penial sheath r rhinophores vd vas deferens (muscular part). Scale bars: D = 1 mm; E, I = 300 μm; F = 30 μm; H = 100 μm. Photos and SEM images by A.V. Martynov.

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Figure 2 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 2 - Phylogenetic tree of aeolids with integrated data on external morphology. The "Flabellinidae" polyphyletic family complex is highlighted by different colours; names of the family- and genus-level taxa are provided. Abbreviations: CPA cleioproctic anus APA acleioproctic anus DTA dorso-terminal anus MSR multiserial radula PPA pleuroproctic anus TSR triserial radula USR uniserial radula.

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Figure 3 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 3 - Chlamylla borealis borealis Bergh, 1899, stat.n. ZMMU Op-479. Kara Sea, living animal 46 mm in length: A dorsal view B ventral view C right lateral view D left lateral view E details of cerata and notal edge F jaws, frontal view, light microscopy G details of masticatory process of jaw, SEM H radular teeth, middle part, SEM I details of small denticles on lateral teeth, SEM J reproductive system, light microscopy. Abbreviations: a anus gf genital fold nc continuous notal edge pc penial collar (external) pr prostate ps penial sheath vd vas deferens (muscular part). Scale bars: G = 500 μm; H =100 μm; I = 10 μm. Photos of living specimens by O.L. Zimina, other photos and SEM images by A.V. Martynov.

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Figure 17 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 17 - Borealia sanamyanae sp. n. ZMMU Op-518. Kurile Islands, Matua Island, living specimen 5 mm in length: A latero-ventral view B dorsal view (fixed) C ventral view (fixed) D jaw, SEM E details of masticatory process of jaw, SEM F details of masticatory process of jaw, light microscopy G radular teeth, posterior part H reproductive system (non-mature), light microscopy. Abbreviations: a anus fgm female gland mass nc continuous notal edge pvd prostatic vas deferens. Scale bars: D = 300 μm; E, G = 30 μm. Photos of living specimens by N.P. Sanamyan, other photos and SEM images by A.V. Martynov.

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Figure 16 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 16 - Borealia nobilis (Verrill, 1880), comb. n. ZMMU Op-510. White Sea, Cape Kartesh, 33 mm in length (live): A living animal, dorsal view B same, ventral view C same, lateral view D same, details of cerata E dissected anterior part (pharynx removed) and rhinophores, light microscopy F jaw, SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I reproductive system (non mature), light microscopy. Abbreviations: a anus fgm female gland mass go genital opening nc continuous notal edge ps penial sheath pvd prostatic vas deferens r rhinophores. Scale bars: F = 300 μm; G, H = 30 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 21 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 21 - Schematic outline of the reproductive systems of the taxa of the families Coryphellidae integrated with molecular phylogenetic data. Colour indication of reproductive system characters: ampulla – green; body wall – gray; distal receptaculum seminis – red; female gland mass – yellow; female genital opening – orange; penis and male genital opening – dark blue; penial sheath – pale blue; prostatic vas deferens – turquoise; proximal distal receptaculum seminis – pink.

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Figure 42 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 42 - Unidentia nihonrossija sp. n. Japan, Pacific Honshu, Osezaki. ZMMU Op-517, specimen 4.5 mm in length (fixed): A dorsal view (live) B lateral view (live) C ventral view (live) D details of cerata (live) E particular elongate elevations at the base of cerata (fixed) F jaw, SEM G details of masticatory process of jaw, SEM H details of masticatory process of jaw, light microscopy I radular teeth, posterior part, SEM J reproductive system, light microscopy. Abbreviations: a anus am ampulla cre ceratal elevated rows fgm female gland mass ps penial sheath pvd prostatic vas deferens rsp proximal receptaculum seminis. Scale bars: F = 100 μm; G, I = 10 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 19 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 19 - Coryphella verrucosa (M. Sars, 1829), short cerata (typical) morphotype. ZMMU Op-539. Norwegian Sea, Gulen Dive Center, living specimen 38 mm in length: A dorsal view B lateral view C right ventral view D details of cerata E dissected anterior part (pharynx removed) and rhinophores, SEM F jaw, SEM G details of masticatory process of jaw, SEM H details of masticatory process of jaw, light microscopy I radular teeth, posterior part, SEM J reproductive system, SEM. Abbreviations: a anus am ampulla rsd distal receptaculum seminis fgm female gland mass go genital opening ps penial sheath pvd prostatic vas deferens r rhinophores rsp proximal receptaculum seminis. Scale bars: E, J = 1 mm; F = 300 μm; G = 30 μm I = 100 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 15 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 15 - Ziminella salmonacea (Cothouy, 1839), comb. n.: A living animal (Spitzbergen), dorsal view B preserved animal, lateral view ZMMU Op-265, 19 mm preserved C same, ventral view D details of cerata E jaw, SEM F details of masticatory process of jaw, SEM G radular teeth, posterior part, SEM H reproductive system, light microscopy. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nc continuous notal edge ps penial sheath pvd prostatic vas deferens. Scale bars: E = 300 μm; F = 30 μm; G bar = 100 μm. Photos of living specimens by B. Gulliksen, other photos and SEM images by A.V. Martynov.

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Last verified 2026-04-29Open record