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Figure 3 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 3 - Chlamylla borealis borealis Bergh, 1899, stat.n. ZMMU Op-479. Kara Sea, living animal 46 mm in length: A dorsal view B ventral view C right lateral view D left lateral view E details of cerata and notal edge F jaws, frontal view, light microscopy G details of masticatory process of jaw, SEM H radular teeth, middle part, SEM I details of small denticles on lateral teeth, SEM J reproductive system, light microscopy. Abbreviations: a anus gf genital fold nc continuous notal edge pc penial collar (external) pr prostate ps penial sheath vd vas deferens (muscular part). Scale bars: G = 500 μm; H =100 μm; I = 10 μm. Photos of living specimens by O.L. Zimina, other photos and SEM images by A.V. Martynov.

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Figure 17 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 17 - Borealia sanamyanae sp. n. ZMMU Op-518. Kurile Islands, Matua Island, living specimen 5 mm in length: A latero-ventral view B dorsal view (fixed) C ventral view (fixed) D jaw, SEM E details of masticatory process of jaw, SEM F details of masticatory process of jaw, light microscopy G radular teeth, posterior part H reproductive system (non-mature), light microscopy. Abbreviations: a anus fgm female gland mass nc continuous notal edge pvd prostatic vas deferens. Scale bars: D = 300 μm; E, G = 30 μm. Photos of living specimens by N.P. Sanamyan, other photos and SEM images by A.V. Martynov.

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Figure 16 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 16 - Borealia nobilis (Verrill, 1880), comb. n. ZMMU Op-510. White Sea, Cape Kartesh, 33 mm in length (live): A living animal, dorsal view B same, ventral view C same, lateral view D same, details of cerata E dissected anterior part (pharynx removed) and rhinophores, light microscopy F jaw, SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I reproductive system (non mature), light microscopy. Abbreviations: a anus fgm female gland mass go genital opening nc continuous notal edge ps penial sheath pvd prostatic vas deferens r rhinophores. Scale bars: F = 300 μm; G, H = 30 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 21 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 21 - Schematic outline of the reproductive systems of the taxa of the families Coryphellidae integrated with molecular phylogenetic data. Colour indication of reproductive system characters: ampulla – green; body wall – gray; distal receptaculum seminis – red; female gland mass – yellow; female genital opening – orange; penis and male genital opening – dark blue; penial sheath – pale blue; prostatic vas deferens – turquoise; proximal distal receptaculum seminis – pink.

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Figure 42 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 42 - Unidentia nihonrossija sp. n. Japan, Pacific Honshu, Osezaki. ZMMU Op-517, specimen 4.5 mm in length (fixed): A dorsal view (live) B lateral view (live) C ventral view (live) D details of cerata (live) E particular elongate elevations at the base of cerata (fixed) F jaw, SEM G details of masticatory process of jaw, SEM H details of masticatory process of jaw, light microscopy I radular teeth, posterior part, SEM J reproductive system, light microscopy. Abbreviations: a anus am ampulla cre ceratal elevated rows fgm female gland mass ps penial sheath pvd prostatic vas deferens rsp proximal receptaculum seminis. Scale bars: F = 100 μm; G, I = 10 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 19 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 19 - Coryphella verrucosa (M. Sars, 1829), short cerata (typical) morphotype. ZMMU Op-539. Norwegian Sea, Gulen Dive Center, living specimen 38 mm in length: A dorsal view B lateral view C right ventral view D details of cerata E dissected anterior part (pharynx removed) and rhinophores, SEM F jaw, SEM G details of masticatory process of jaw, SEM H details of masticatory process of jaw, light microscopy I radular teeth, posterior part, SEM J reproductive system, SEM. Abbreviations: a anus am ampulla rsd distal receptaculum seminis fgm female gland mass go genital opening ps penial sheath pvd prostatic vas deferens r rhinophores rsp proximal receptaculum seminis. Scale bars: E, J = 1 mm; F = 300 μm; G = 30 μm I = 100 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 15 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 15 - Ziminella salmonacea (Cothouy, 1839), comb. n.: A living animal (Spitzbergen), dorsal view B preserved animal, lateral view ZMMU Op-265, 19 mm preserved C same, ventral view D details of cerata E jaw, SEM F details of masticatory process of jaw, SEM G radular teeth, posterior part, SEM H reproductive system, light microscopy. Abbreviations: a anus am ampulla fgm female gland mass go genital opening nc continuous notal edge ps penial sheath pvd prostatic vas deferens. Scale bars: E = 300 μm; F = 30 μm; G bar = 100 μm. Photos of living specimens by B. Gulliksen, other photos and SEM images by A.V. Martynov.

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Figure 14 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 14 - Ziminella japonica (Volodchenko, 1941), comb. n.: A ZMMU Op-261, (Sea of Japan, 470–528 m) dorsal view, 8.3 mm in fixed length B same, ventral view C jaw, light microsocopy D details of masticatory process of jaws, SEM E radular teeth, posterior part F details of rachidian radular tooth G reproductive system, light microscopy. Abbreviations: am ampulla fgm female gland mass go genital opening nc continuous notal edge pvd prostatic vas deferens. Scale bars: D = 30 μm; E = 100 μm. Photos and SEM images by A.V. Martynov.

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Figure 12 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 12 - Ziminella abyssa sp. n.: A preserved holotype (Sea of Japan, ZSM Mol-20100647, 2676 m), dorsal view, 19 mm in fixed length B same, latero-ventral view C jaw, SEM ZMMU Op-259, depth 3460 m D details of masticatory process of jaws E radular teeth, posterior part, SEM ZMMU Op-250, 24 mm length, 3300 m F–H details of three central teeth with additional or highly aberrant lateral denticles ZMMU Op-248, 12 mm length, 3560 m I rachidian tooth of juvenile specimen 2 mm length showing additional lateral denticles ZMMU Op-264, 1525 m J reproductive system, light microscopy ZSM Mol-20100644, 3213 m. Abbreviations: a anus fgm female gland mass go genital opening nc continuous notal edge pvd prostatic vas deferens. Scale bars: C = 1 mm; E = 50 μm; F, G, H = 300 μm. Photos and SEM images by A.V. Martynov.

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Figure 29 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 29 - Itaxia falklandica (Eliot, 1907), comb. n. South Pacific, Chile, Canal Artilleria, ZSM Mol-20070592, specimen 12 mm length (fixed): A dorsal view (live) B ventral view (live) C lateral view (live) D details of cerata (live) E jaws, light microscopy F details of masticatory process of jaw, light microscopy G radular teeth, posterior part, SEM H reproductive system (non mature), light microscopy. Abbreviations: fgm female gland mass nc continuous notal edge pvd prostatic vas deferens. Scale bars: G = 30 μm. Photos of living specimens by M. Schrödl, other photos and SEM images by A.V. Martynov.

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Figure 26 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 26 - Gulenia monicae sp. n. Norwegian Sea, Gulen Dive Center. Holotype ZMMU Op-466, living specimen 34 mm length: A dorsal view B lateral view C ventral view; Paratype ZMMU Op-411, living specimen 24.5 mm length D dorsal view E details of cerata F jaw, SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I J reproductive system, light microscopy. Abbreviations: a anus am ampulla fgm female gland mass g gonad go genital opening nc continuous notal edge p penis ps penial sheath pvd prostatic vas deferens rsd distal receptaculum seminis rsp proximal receptaculum seminis. Scale bars: F = 300 μm; G, H = 30 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 11 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 11 - Polaria polaris (Volodchenko, 1946), comb. n. ZMMU Op-519. Laptev Sea, living specimen 12 mm in length: A dorsal view B latero-ventral view C ventral view D details of cerata E jaw, SEM F details of masticatory process of jaw, SEM G radular teeth, posterior part, SEM H reproductive system, light microscopy I reproductive system, SEM. Abbreviations: a anus am ampulla rsd distal receptaculum seminis fgm female gland mass; g gonad; nc continuous notal edge p penis ps penial sheath pvd prostatic vas deferens Scale bars: E, I = 300 μm; F = 30 μm; G = 100 μm. Photos of living specimens by O.L. Zimina, other photos and SEM images by A.V. Martynov.

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Figure 13 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 13 - Ziminella circapolaris sp. n.: Barents Sea, Franz-Josef Land: A living animal, dorsal view ZMMU Op-482 B living animal on a hydroid colony ZMMU Op-483, 12 mm in fixed length C same, details of cerata D dissected anterior part (pharynx removed) and rhinophores, SEM E jaw, SEM F details of masticatory process of jaw, SEM G radular teeth, posterior part, SEM ZMMU Op-482 H lateral tooth, close up, SEM ZMMU Op-482 I radular teeth, posterior part, SEM ZMMU Op-483 J lateral tooth, close up, SEM ZMMU Op-483 K reproductive system, SEM ZMMU Op-483 L reproductive system, light microscopy. Abbreviations: am ampulla fgm female gland mass nc continuous notal edge pvd prostatic vas deferens r rhinophores. Scale bars: D = 1 mm; E, K = 300 μm; F, H = 30 μm; G, I, J = 100 μm. Photos of living specimens by O.V. Savinkin, other photos and SEM images by A.V. Martynov.

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Figure 23 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 23 - Fjordia chriskaugei sp. n. A–C, F Holotype ZMMU Op-477, living specimen 45 mm length, Norwegian Sea, Gulen Dive Center: A dorsal view B lateral view C ventral view F same, details of cerata D Paratype ZMMU Op- 412, living specimen 42 mm length, dorsal view, Gulen E living specimen on a hydroid colony, ZMMU Op-499, Ireland G–L Paratype ZMMU Op-405, details of living specimen 30 mm length, Gulen G jaw, SEM H details of masticatory process of jaw I radular teeth, posterior part, SEM J reproductive system, SEM K penis, SEM L seminal reservoirs, SEM. Abbreviations: a anus am ampulla rsd distal receptaculum seminis fgm female gland mass go genital opening nd discontinuous notal edge p penis ps penial sheath pvd prostatic vas deferens rsp proximal receptaculum seminis. Scale bars: G, K, L = 300 μm; H = 30 μm; I = 100 μm; J = 1 mm. Photos of living specimens by T.A. Korshunova (A–D), and B. Picton (E), SEM images by A.V. Martynov.

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Figure 10 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 10 - Paracoryphella parva (Hadfield, 1963), comb. n. Holotype NHMD-91476, 2 mm length: A fixed animal, dorsal view B fixed animal, ventral view of anterior part C fixed animal, lateral view D rachidian radular teeth E lateral teeth F reproductive system. Abbreviations: a anus am ampulla fgm female gland mass fgo female genital opening nc continuous notal edge p penis pvd prostatic vas deferens r rhinophores rsd distal receptaculum seminis. Photos by A.V. Martynov. Drawings D, E, F from original description of P. parva from Hadfield (1963).

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Figure 1 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 1 - Phylogenetic tree of aeolidacean nudibranchs based on concatenated molecular data (COI + 16S + 28S + H3) represented by Bayesian Inference (BI). Numbers above branches represent posterior probabilities from Bayesian Inference. Numbers below branches indicate bootstrap values for Maximum Likelihood. Some branches are collapsed at species level. The "Flabellinidae" polyphyletic family complex is highlighted by different colours (see Fig. 2 for the family names).

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Figure 25 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 25 - Gulenia borealis (Odhner, 1922), comb. n. GNM Gastropoda 9417, 11 mm length (fixed), Norway (A–C, E–I), Sweden (D): A dorsal view (live) B ventro-lateral view (fixed) C details of cerata D living animals in situ on Funiculia (Pennatulacea) E jaw, SEM F, G details of masticatory process of jaw, SEM H radular teeth, posterior part I reproductive system. Abbreviations: a anus am ampulla fgm female gland mass nc continuous notal edge p penis ps penial sheath rsd distal receptaculum seminis rsp proximal receptaculum seminis vd vas deferens (muscular part). Scale bars: E = 300 μm; F = 30 μm; G = 10 μm; H = 100 μm. Photos of living specimens by T. Cedhagen (A), and K. Malmberg (D), SEM images by A.V. Martynov.

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Figure 22 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 22 - Fjordia browni (Picton, 1980), comb. n. Norwegian Sea, Gulen Dive Center. ZMMU Op-413, living specimen 48 mm in length: A dorsal view B lateral view C ventral view D living specimen on hydroid colony E details of cerata F jaw, SEM G details of masticatory process of jaw, SEM H radular teeth, posterior part, SEM I reproductive system, SEM J penis, SEM K seminal reservoirs. Abbreviations: a anus am ampulla rsd distal receptaculum seminis fgm female gland mass nd discontinuous notal edge p penis ps penial sheath pvd prostatic vas deferens rsp proximal receptaculum seminis. Scale bars: F, J, K = 300 μm; G = 30 μm; H = 100 μm; I = 1 mm. Photos of living specimens by T.A. Korshunova (A–C), and C. Skauge (D), SEM images by A.V. Martynov.

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Figure 20 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 20 - Coryphella verrucosa (M. Sars, 1829), long cerata ("rufibranchialis") morphotype. ZMMU Op-521, Barents Sea, Oskara Bay, living specimen 42 mm in length (live): A dorsal view B lateral view C ventral view D details of cerata E jaw, SEM F details of masticatory process of jaw, SEM G details of masticatory process of jaw, light microscopy H radular teeth, posterior part, SEM I reproductive system, light microscopy J reproductive system, SEM K penis partially everted, SEM. Abbreviations: a anus am ampulla fgm female gland mass go genital opening p penis pvd prostatic vas deferens. Scale bars: E bar = 300 μm; F, K = 30 μm; H = 100 μm; J = 1 mm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Figure 18 from: Korshunova T, Martynov A, Bakken T, Evertsen J, Fletcher K, Mudianta WI, Saito H, Lundin K, Schrödl M, Picton B (2017) Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1-139. https://doi.org/10.3897/zookeys.717.21885

Figure 18 - Coryphella pseudoverrucosa Martynov et al., 2015. ZMMU Op- 528. NW Pacific, Kamchatka, Avachinskaya guba, living specimen 12.5 mm in length (fixed): A dorsal view B ventral view C lateral view D details of cerata E jaw, SEM F details of masticatory process of jaw, SEM G radular teeth, posterior part, SEM H reproductive system, light microscopy. Abbreviations: a anus am ampulla fgm female gland mass p penis ps penial sheath pvd prostatic vas deferens. Scale bars: E = 300 μm; F 30 μm; G = 100 μm. Photos and SEM images by T.A. Korshunova, A.V. Martynov.

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Last verified 2026-04-30Open record

International Brain Laboratory public data

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Last verified 2026-04-29Open record

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neuroscienceopenPublished datasets are available on demand over the internet.
Last verified 2026-04-29Open record