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Fig. 31 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 31 – Aedeagi and spicula gastrales, a–c, Synarmostes antsingyi (Tsingy de Bemaraha); d–f, Synarmostes beankae sp. nov., holotype; g–i, Synarmostes bekaraokae sp. nov., holotype; j–l, Synarmostes caterinae sp. nov., holotype; m–o, Synarmostes monoculus sp. nov., holotype.

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Fig. 11 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 11 – Synarmostes tibialis, mouthparts (specimen from « Madagascar »), a–b, mandibles in dorsal view; c–d, mandibles in ventral view; e, labium and labial palpi; f, maxilla and maxillary palpus; g, detail of galeal brush; h, distal epipharynx.

opencc-by-4.0Nov 2021View details →
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Fig. 30 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 30 – Aedeagi and spicula gastrales, a–c, Synarmostes aes sp. nov., holotype; d–f, Synarmostes separatus sp. nov., holotype; g–i, Synarmostes compactus sp. nov., holotype; j–l, Synarmostes fortis sp. nov., holotype; m–o, Synarmostes makirovanae sp. nov., holotype; p–r, Synarmostes lavasoae sp. nov., holotype.

opencc-by-4.0Nov 2021View details →
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Fig. 22 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 22 – Synarmostes compactus sp. nov., a, dorsal view of distended paratype; b, ventral view of rolled up paratype; c, dorsal view of rolled up paratype; d, lateral view of rolled up paratype; Synarmostes fortis sp. nov., e, dorsal view of distended holotype; f, ventral view of rolled up paratype; g, dorsal view of rolled up paratype; h, lateral view of rolled up paratype.

opencc-by-4.0Nov 2021View details →
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Fig. 7 – a in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 7 – a, Synarmostes benaraensis sp. nov., apex of female mesotibia; b, Goudotostes mayottensis sp. nov. (Mont Benara) ventral side of male protibia; c, Goudotostes mayottensis sp. nov. (Mont Benara) detail of fist protarsomere; d, Synarmostes benaraensis sp. nov. apex of male mesotibia; e, Philharmostes scapulatus sp. nov. detail of apex of female mesotibia, showing the lateral setae of first mesotarsomere; f, Goudotostes mayottensis sp. nov. detail of pronotal setation and punctation; g, Goudotostes mayottensis sp. nov. (Mont Benara) detail of pronotal setation and punctation (transverse impressed lines); h, Cryptosphaeroides hystrix, detail of elytral setation and punctation.

opencc-by-4.0Nov 2021View details →
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Fig. 4 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 4 – Heads of Synarmostes in ventral, lateral and dorsal views (not to scale) in order to show the extent of ocular area, a–c, Synarmostes tibialis (example of large–sized ventral ocular area); d–f, Synarmostes annamariae sp. nov. (example of medium–sized ventral ocular area); g–i, Synarmostes fortis sp. nov. (example of small–sized ventral ocular area and vestigial dorsal ocular area); j–l, Synarmostes benaraensis sp. nov. (example of small–sized ventral ocular area).

opencc-by-4.0Nov 2021View details →
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Fig. 36 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 36 – Map showing the distribution of the genus Synarmostes in Mayotte island (blank map source: © Rémi Kaupp, CC–BY–SA, Wiki- media Commons).

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Fig. 10 – Wings. a in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 10 – Wings. a, Synarmostes tibialis; b, Synarmostes occidentalis sp. nov.; c, Synarmostes sp. C; d, Philharmostes metallicus sp. nov.; e, Synarmostes makirovanae sp. nov.; f, Goudotostes laevis sp. nov..

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Fig. 21 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 21 – Synarmostes humilis (Mont Combani), a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, lateral view of rolled up specimen; Synarmostes benaraensis sp. nov., e, dorsal view of distended paratype; f, ventral view of rolled up paratype; g, dorsal view of rolled up paratype; h, lateral view of rolled up paratype.

opencc-by-4.0Nov 2021View details →
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Fig. 5 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 5 – Heads of Goudotostes, a–c, female of Goudotostes orangeanus sp. nov. (ventral, lateral and dorsal views) (cap: clypeal apical projection); d–e, Goudotostes electrimontis sp. nov., circled area showing alleged vestigial dorsal ocular area; f–g, Goudotostes litoralis sp. nov., circled area showing alleged vestigial dorsal ocular area.

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Fig. 1 – a in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 1 – a, Synarmostes tibialis, ventral side, abdomen removed (cm: connecting membrane between prothorax and mesothorax); b, Goudotostes ramamonjisoae sp. nov. ventral view of rolled up specimen (bac: bilobate anterior carina; ppt: pronotal paradiscal tubercle; pltc: pronotal lateral transverse carina; ft: fronto-clypeal tubercle); c, Goudotostes sp. C, dorsal view of rolled up specimen (bbc: bilobate basal carina; pt: parasutural tubercle; efi: elytral first interstria; eflc: elytral first longitudinal carina; ht: humeral tubercle); d, Goudotostes lapidisilvae sp. nov., dorsal view of rolled up specimen (mbc: monolobate basal carina; pt: parasutural tubercle; ht: humeral tubercle; obt: other basal tubercles; ps: parascutellar furrow); e, Goudotostes lokobensis sp. nov., ventral view of rolled up specimen (ltc: lateral transverse carina; mllt: medio-lateral large tubercle; pt: paradiscal tubercle; blac: bilobate anterior carina; ct: clypeal tubercle); f, Goudotostes electrimontis sp. nov., dorsal view of rolled up specimen (efi: elytral first interstria; pt: parasutural tubercle; ht: humeral tubercle; obt: other basal tubercles); g, Philharmostes nelsoni sp. nov., lateral view of rolled up specimen (eds: elytral dorsal surface; elc: elytral lateral carina; iss: inferior sutural stria; p: pseudoepipleure; saa: striated articular area; ma: marginal area) (photos not to scale).

opencc-by-4.0Nov 2021View details →
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Fig. 2 – a in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 2 – a, Philharmostes indri sp. nov., aedeagus (bap: basal apophysis of parameres; e: endophallites; ep: endophallus; te: temones; p: paramere); b, Goudotostes lokobensis sp. nov., spiculum gastrale (sg: spiculum; a: arm of spiculum; nh: ninth hemisternite); c, Synarmostes benaraensis sp. nov., head (cap: clypeal apical projection; cl: clypeus; ge: gena; doa: dorsal ocular area; gc: genal canthus; fr: frons; tlv: transverse line of vertex; v: vertex; m: mandible). White semicircle delimitates the clypeal area (photos not to scale).

opencc-by-4.0Nov 2021View details →
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Fig. 35 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 35 – Distribution map of the genus Synarmostes in Madagascar and the Comoro archipelago (for Mayotte see separate map).

opencc-by-4.0Nov 2021View details →
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Fig. 8 – a in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 8 – a, Cryptosphaeroides hystrix, antenna (ped: pedicellus); b, Synarmostes benaraensis sp. nov., antenna; c, Pseudosynarmostes mitsinjo, antenna, antenna; d, Goudotostes mayottensis sp. nov.; e, Philharmostes scapulatus sp. nov., antenna; f, Goudotostes rajemisonae sp. nov., aedeagus showing invaginated left paramere (pi: paramere invagination); g, Goudotostes rajemisonae sp. nov., detail of circled area of Fig. f showing sensilla.

opencc-by-4.0Nov 2021View details →
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Fig. 9– a in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 9– a, Synarmostes benaraensis sp. nov. female protibia (at: apical teeth); b, Synarmostes benaraensis sp. nov., male protibia; c, Philharmostes scapulatus, protibia; d, Pseudosynarmostes mitsinjo, female protibia; e, Pseudosynarmostes mistinjo, male protibial; f, Goudotostes orangeanus sp. nov., female protibia; g, Goudotostes orangeanus sp. nov., male protibia; h, Philharmostes scapulatus, female mesotibia; i, Synarmostes benaraensis sp. nov., male mesotibia (mias: male inner apical spur); j, Synarmostes benaraensis sp. nov., female mesotibia; k, Synarmostes annamariae sp. nov., male mesotibia l, Goudotostes orangeanus sp. nov. male mesotibia (mias: male inner apical spur); m, Goudotostes orangeanus sp. nov., metatibia; n, Synarmostes benaraensis sp. nov., metatibia; o, Synarmostes annamariae sp. nov., metatibia; p, Pseudosynarmostes mitinjo, metatibia.

opencc-by-4.0Nov 2021View details →
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Fig. 16 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 16 – Synarmostes antsingyi (Tsingy de Bemaraha), a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, lateral view of rolled up specimen; Synarmostes beankae sp. nov., e, dorsal view of distended holotype; f, ventral view of rolled up paratype; g, dorsal view of rolled up paratype; h, lateral view of rolled up paratype.

opencc-by-4.0Nov 2021View details →
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Fig. 27 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 27 – Synarmostes tibialis (Andasibe, Mitsinjo forest), a, dorsal view of distended specimen; b, ventral view of rolled up specimen; c, dorsal view of rolled up specimen; d, lateral view of rolled up specimen; Synarmostes bekaraokae sp. nov., holotype, e, dorsal view of distended specimen; f, lateral view of distended specimen.

opencc-by-4.0Nov 2021View details →
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Fig. 25 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 25 – Synarmostes niger, a, holotype in dorsal view (photo by MNHN); b, holotype in lateral view (photo by MNHN); c, labels of holotype (photo by MNHN); d, paratype in dorsal view (photo by MNHN); e, paratype from Antsalova in dorsal view.

opencc-by-4.0Nov 2021View details →
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Fig. 89 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 89 – Distribution map of the flightless Philharmostes in Madagascar.

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Fig. 77 in The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)

Fig. 77 – Distribution map of the genus Goudotostes in Madagascar and the Comoro archipelago.

opencc-by-4.0Nov 2021View details →

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