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FIGURES 18–27 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 18–27. Zagurya eusquamosa (Frey, 1975), new combination, appendages. 18–19, antennae (male, female); 20, labium, ventral (left side with setae omitted); 21–22, protibia–tarsus, anterior (male, female); 23–24, male mesotibia–tarsus (anterior, posterior; some setae omitted); 25, female mesotibia–tarsus, anterior; 26–27, metatibia–tarsus, anterior (male, female). Scale = 1 mm.

FIGURES 34–52 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 34–52. Macrodactylini and other phytophagous Scarabaeidae, distal metatarsomeres and claws, some setae omitted. 34–35, Zagurya eusquamosa (Frey, 1975), new combination (anterior, inner); 36–37, Plectris tomentosa LePeletier & Audinet-Serville, 1828 (anterior, inner); 38–39, Mallotarsus spadiceus Blanchard, 1850 (anterior, inner); 40–44, Ancistrosoma klugi Curtis, 1835; 40–41, anterior (with and without some setae); 42, inner; 43–44, internal section (with empodium protracted and retracted); 45–49, Alvarinus pallidipennis Blanchard, 1850; 45, anterior; 46, internal section; 47–49, inner (with empodium protracted, partially retracted, and fully retracted); 50–51, Hoplia argentea (Poda, 1761), Hopliini (anterior, inner); 52, Rutela lineola (Linnaeus, 1767), Rutelini, inner. Scale = 1 mm.

FIGURES 53–55 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 53–55. Generalized scarabaeoid beetle and study of leg position; 53, beetle lateral, all legs with anterior side faced to the anterior body side; 54, beetle dorsal, legs position as usually found in pinned specimens; 55, diagram of an insect body axis and leg axis. Sides of legs and podomeres colorful, see that the dorsoventral axis is used to the body but not to legs. aa, anterodistal articular lobe of femur; io, inner ornamentation of metatarsomere V; is, inner spur of protibia; oc1, outer transverse carina I of metatibia; ot1, outer tooth I of protibia; pa, posterodistal articular lobe of femur.

FIGURES 9–17 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 9–17. Zagurya eusquamosa (Frey, 1975), new combination, habitus. 9–13, male (dorsal, latero-dorsal, antero-dorsal, lateral, posterior); 14–17, female (dorsal, antero-dorsal, lateral, posterior). Scale = 4 mm.

FIGURES 1–8 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 1–8. Specimen and habitat of Zagurya eusquamosa (Frey, 1975), new combination. 1, Banho do Belchior, Santuário do Caraça (credits: Santuário do Caraça); 2–3, female holotype (dorsal and labels; credits: Christoph Germann); 4, head of male (left side showing posterior angle of clypeus (red) not in canthus area (green)); 5, scutellum (arrow showing the angulate contact of mesoscutum−scutellum); 6–7, male at Banho do Belchior locality (credits: Fernando Frieiro-Costa); 8, female at Refúgio do Peixe Tolo locality (credits: Frederico Falcão Salles). Scale = 1 mm.

FIGURES 56 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 56. Zagurya eusquamosa (Frey, 1975), new combination, distributional record (red dots). Minas Gerais State with regional division, rivers in dark blue, Serra do Espinhaço in brown. Scale = 300 km.

FIGURES 28–33 in Zagurya, a new genus from Brazil with notes on claw movement in Macrodactylini (Coleoptera: Scarabaeidae: Melolonthinae)

FIGURES 28–33. Zagurya eusquamosa (Frey, 1975), new combination, appendages and terminalia. 28–29, apex of metatibia (male, female); 30, genital ring; 31, aedeagus, lateral; 32, parameres, distal; 33, female terminalia, posterior (left side with setae represented by their puncture). dgc, distal piece of gonocoxite; dsp, distal piece of spiculum gastrale; pgc, proximal piece of gonocoxite; psp, proximal piece of spiculum gastrale (apodeme); scm, second connective membrane (connecting genital ring to aedeagus); st9, abdominal sternite IX; tcm, third connective membrane (connecting abdominal segment VIII to genital ring). Scale = 1 mm.

Figs. 5–8. 5 in PseudopectinosomaKatovich, New Genus, and Review ofPectinosomaArrow (Scarabaeidae: Melolonthinae: Macrodactylini)

Figs. 5–8. 5) Pseudopectinosoma mesa, male, procoxae and pro- and mesosternum, ventral view; 6) P. mesa, male, visible abdominal segments, ventral view, membranous posterior margin of sixth ventrite shaded gray; 7) Pectinosoma elongatum, male, procoxae and pro- and mesosternum, ventral view; 8) P. elongatum male, visible abdominal segments, ventral view.

Figs. 9–12. 9 in PseudopectinosomaKatovich, New Genus, and Review ofPectinosomaArrow (Scarabaeidae: Melolonthinae: Macrodactylini)

Figs. 9–12. 9) Pseudopectinosoma mesa, male, head and pronotum, dorsal view; 10) P. mesa, male, scutellum, elytra, and pygidium, dorsal view; 11) Pectinosoma elongatum, male, head and pronotum, dorsal view; 12) P. elongatum, male, scutellum, elytra, and pygidium, dorsal view.

Figs. 13–16. 13 in PseudopectinosomaKatovich, New Genus, and Review ofPectinosomaArrow (Scarabaeidae: Melolonthinae: Macrodactylini)

Figs. 13–16. 13) Pseudopectinosoma mesa, female, head and pronotum, dorsal view; 14) P. mesa, female, scutellum, elytra, and pygidium, dorsal view; 15) Pectinosoma elongatum, female, head and pronotum, dorsal view; 16) P. elongatum, female, scutellum, elytra, and pygidium, dorsal view.

Figs. 1– 4. 1–2 in PseudopectinosomaKatovich, New Genus, and Review ofPectinosomaArrow (Scarabaeidae: Melolonthinae: Macrodactylini)

Figs. 1– 4. 1–2) Pseudopectinosoma mesa, male, dorsal and lateral views; 3–4) Pectinosoma elongatum, male, dorsal and lateral views.

Fig. 15. Schizochelus Blanchard, 1850 in Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification

Fig. 15. Schizochelus Blanchard, 1850, ♂, habitus, lateral (without some appendages). A. Schizochelus flavescens Blanchard, 1850. B. Schizochelus bicoloripes Blanchard, 1850. C. Schizochelus mirabilis (Moser, 1921) comb. nov. Scale bar = 1 mm.

Fig. 22. A–D in Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification

Fig. 22. A–D. Holotype, ♂ (dorsal, labels). A–B. Paulosawaya ornatissima Martínez & d'Andretta, 1956. C–F. Pseudoleuretra bokermanni Martínez & d'Andretta, 1956. E–F. Non-type material. E. Junkia ceylanica Nonfried, 1894. F. Plectris tomentosa LePeletier de Saint-Fargeau & Audinet-Serville, 1828. Scale bars = 2 mm.

Fig. 13 in Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification

Fig. 13. ♂♂. A. Head–prothorax dorsal, tarsus rotated laterally to apex. B–C. Mesotibia−tarsus (C, with details of tarsomere I and IV). D−G. Aedeagus (lateral, parameres apex). H. Metafemur. A–B, D–E. Pseudoserica marmorea Guérin-Méneville, 1831. C, F−H. Mallotarsus spadiceus Blanchard, 1850. Scale bars = 1 mm.

Fig. 12 in Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification

Fig. 12. ♂♂. A. Head–prothorax dorsal, tarsus rotated laterally to apex. B. Antenna. C. Protibia−tarsus. D−I. Aedeagus (lateral, parameres apex). J. Metatibia. K. Head−prothorax, ventral. A, F−G. Euryaspis obesa (Burmeister, 1855) comb. nov. B−E. Euryapsis gaudichaudi Blanchard, 1850. H−J. Philochloenia amata nom. nov. K. Philochloenia castaneipennis (Guérin-Méneville, 1831) comb. nov. Scale bars = 1 mm.

Fig. 9. A in Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification

Fig. 9. A. Lateroventral detail of male abdomen. B, F, H. Aedeagus, lateral. C. Left paramere. D, G, I. Parameres, apex. E. Parameres, ventral. A−E. Chariodema virescens Blanchard, 1842. F−G. Ceraspis bivulnerata (Germar, 1824). H−I. Faula cornuta Blanchard, 1850. Scale bars = 1 mm.

Fig. 1. Agaocnemis pruina Moser, 1918. A in Macrodactylini (Coleoptera, Scarabaeidae, Melolonthinae): primary types of type species and taxonomic changes to the generic classification

Fig. 1. Agaocnemis pruina Moser, 1918. A. Male habitus, lateral (without some appendages). B. Female abdomen lateral detail. C–D. Aedeagus (lateral, parameres apex). Scale bars = 1 mm.