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Text-fig. 6. Scanning electron micrographs of multicarpellate and apocarpous floral structures from the Early Cretaceous Puddledock locality, Virginia, USA (a, b: PP43701, Puddledock sample 001; c: PP43000x, Puddledock sample 073). a) Anacostia? sp., strongly compressed, elongated receptacle with spirally arranged carpels (red dots; not all shown); note larger size compared to the other floral structures; b) Numerous Anacostia type pollen grains in proximal view from the base of floral structure in (a); note graded reticulum over the proximal pole of the pollen grains; c) Elongated receptacle with numerous carpels in a spiral arrangement, possibly representing an earlier developmental stage of Anacostia? sp. Scale bars = 1 mm (a, c), 10 µm (b). in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal
Text-fig. 6. Scanning electron micrographs of multicarpellate and apocarpous floral structures from the Early Cretaceous Puddledock locality, Virginia, USA (a, b: PP43701, Puddledock sample 001; c: PP43000x, Puddledock sample 073). a) Anacostia? sp., strongly compressed, elongated receptacle with spirally arranged carpels (red dots; not all shown); note larger size compared to the other floral structures; b) Numerous Anacostia type pollen grains in proximal view from the base of floral structure in (a); note graded reticulum over the proximal pole of the pollen grains; c) Elongated receptacle with numerous carpels in a spiral arrangement, possibly representing an earlier developmental stage of Anacostia? sp. Scale bars = 1 mm (a, c), 10 µm (b).
Text-fig. 5. Scanning electron micrographs (d–g) and synchrotron radiation X-ray tomographic microscopy orthoslices (a–c) of flowers of Atlantocarpus virginiensis gen. et sp. nov. (a–d: holotype, PP43780, Puddledock sample 156), Atlantocarpus sp. from the Early Cretaceous Buarcos locality (e, f: S105025, Buarcos sample 244) and receptacle of Atlantocarpus? from the Early Cretaceous Vale de Água locality (g: S101300, Vale de Água sample 141). a) Flower in lateral view showing scar from a single bract (br), attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels; b) Flower in lateral view showing expanded basal portion of the elongated receptacle and young carpels; c) Flower in longitudinal section showing expanded basal portion of the elongated receptacle and young carpels; note the irregular, possibly expanded stigmatic region (arrow heads), (orthoslice yz0340); d) Flower in lateral view showing attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels with possible grooved stigmatic regions (arrow heads); e) Flower in lateral view showing expanded basal portion of elongated receptacle and young carpels; f) Detail of flower in (e) showing in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal
Text-fig. 5. Scanning electron micrographs (d–g) and synchrotron radiation X-ray tomographic microscopy orthoslices (a–c) of flowers of Atlantocarpus virginiensis gen. et sp. nov. (a–d: holotype, PP43780, Puddledock sample 156), Atlantocarpus sp. from the Early Cretaceous Buarcos locality (e, f: S105025, Buarcos sample 244) and receptacle of Atlantocarpus? from the Early Cretaceous Vale de Água locality (g: S101300, Vale de Água sample 141). a) Flower in lateral view showing scar from a single bract (br), attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels; b) Flower in lateral view showing expanded basal portion of the elongated receptacle and young carpels; c) Flower in longitudinal section showing expanded basal portion of the elongated receptacle and young carpels; note the irregular, possibly expanded stigmatic region (arrow heads), (orthoslice yz0340); d) Flower in lateral view showing attachment scars of tepals (t) and stamens (st) on the expanded basal portion of elongated receptacle and young carpels with possible grooved stigmatic regions (arrow heads); e) Flower in lateral view showing expanded basal portion of elongated receptacle and young carpels; f) Detail of flower in (e) showing
Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy orthoslices of flowers of Lambertiflora virginiense gen. et sp. nov. from the Early Cretaceous Puddledock locality, Virginia, USA (holotype, PP53796, Puddledock sample 081). White dots – tepals, yellow dots – stamens or staminodes, red dot – central conical gynoecial region. a) Flower in longitudinal section showing elongated overlapping tepals, remains of probable poorly developed stamens or staminodes and probable poorly developed carpels on the central conical gynoecial region of the receptacle (orthoslice yz0454); b) Flower in transverse section showing rhomboidal bases of 30 tepals, nine poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1160); c) Flower in transverse section at the level of the floral receptacle showing 30 tepals, nine of the poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1250). Scale bars = 250 µm (a–c). in Multiparted, Apocarpous Flowers From The Early Cretaceous Of Eastern North America And Portugal
Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy orthoslices of flowers of Lambertiflora virginiense gen. et sp. nov. from the Early Cretaceous Puddledock locality, Virginia, USA (holotype, PP53796, Puddledock sample 081). White dots – tepals, yellow dots – stamens or staminodes, red dot – central conical gynoecial region. a) Flower in longitudinal section showing elongated overlapping tepals, remains of probable poorly developed stamens or staminodes and probable poorly developed carpels on the central conical gynoecial region of the receptacle (orthoslice yz0454); b) Flower in transverse section showing rhomboidal bases of 30 tepals, nine poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1160); c) Flower in transverse section at the level of the floral receptacle showing 30 tepals, nine of the poorly developed stamens or staminodes, and the central poorly differentiated gynoecial region of the receptacle (orthoslice xy1250). Scale bars = 250 µm (a–c).
Text-fig. 54. Scanning electron microscope (SEM; a–e, g, h) and synchrotron x-ray microscopy (SRXTM; f, i) images of unnamed angiosperm fruits and seeds; Torres Vedras locality, Portugal. a) Follicle sp. 1, narrow elongate follicle with sessile and decurrent stigma; b) Follicle sp. 2, broad, dehisced follicle with elongate and transverse fibers lining the locule; c) Angiosperm seed sp. 1 with thin, smooth seed coat; d) Angiosperm seed sp. 2 with verrucate seed coat; e) Angiosperm seed sp. 3 with exotestal and foveolate seed coat; f) Angiosperm seed sp. 4 with smooth seed coat; g, h) Angiosperm seed sp. 5 with raised epidermal cells forming a reticulate pattern; i) Wedge-shaped angiosperm fruit with remains of floral parts near apex. Specimens, TV43-S136726 (a), TV44-S148144 (b), TV43-S170074 (c), TV43-S136747 (d), TV43-S170073 (e), TV38-S174615 (f), TV44-S148003 (g), TV44-S148004 (h), TV43-S174685 (i). Scale bars 300 Μm (a–i). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 54. Scanning electron microscope (SEM; a–e, g, h) and synchrotron x-ray microscopy (SRXTM; f, i) images of unnamed angiosperm fruits and seeds; Torres Vedras locality, Portugal. a) Follicle sp. 1, narrow elongate follicle with sessile and decurrent stigma; b) Follicle sp. 2, broad, dehisced follicle with elongate and transverse fibers lining the locule; c) Angiosperm seed sp. 1 with thin, smooth seed coat; d) Angiosperm seed sp. 2 with verrucate seed coat; e) Angiosperm seed sp. 3 with exotestal and foveolate seed coat; f) Angiosperm seed sp. 4 with smooth seed coat; g, h) Angiosperm seed sp. 5 with raised epidermal cells forming a reticulate pattern; i) Wedge-shaped angiosperm fruit with remains of floral parts near apex. Specimens, TV43-S136726 (a), TV44-S148144 (b), TV43-S170074 (c), TV43-S136747 (d), TV43-S170073 (e), TV38-S174615 (f), TV44-S148003 (g), TV44-S148004 (h), TV43-S174685 (i). Scale bars 300 Μm (a–i).
Text-fig. 45. Scanning electron microscope (SEM) images of monocolpate pollen of Dinisia portugallica gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Two pollen grains showing poorly defined distal aperture (arrowhead) and distinctive vermiform reticulum forming luminae of variable shapes and sizes; note especially the irregularly and incomplete reticulum in the grain on the left; c) Reticulum showing smooth, vermiform muri attached to the smooth surface of the foot layer by long columellae; note that columellae often terminate segments of muri that are not closed; d, e) Pollen grains showing proximal surface (d), poorly defined distal aperture (e, arrowhead) and distinctive vermiform reticulum supported by long columellae; note dense covering of small, spherical orbicules on the inner surface of the anther wall. Specimen, TV44-S148216 (holotype). Scale bars 300 Μm (a), 6 Μm (b, d, e), 3 Μm (c). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 45. Scanning electron microscope (SEM) images of monocolpate pollen of Dinisia portugallica gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Two pollen grains showing poorly defined distal aperture (arrowhead) and distinctive vermiform reticulum forming luminae of variable shapes and sizes; note especially the irregularly and incomplete reticulum in the grain on the left; c) Reticulum showing smooth, vermiform muri attached to the smooth surface of the foot layer by long columellae; note that columellae often terminate segments of muri that are not closed; d, e) Pollen grains showing proximal surface (d), poorly defined distal aperture (e, arrowhead) and distinctive vermiform reticulum supported by long columellae; note dense covering of small, spherical orbicules on the inner surface of the anther wall. Specimen, TV44-S148216 (holotype). Scale bars 300 Μm (a), 6 Μm (b, d, e), 3 Μm (c).
Text-fig. 43. Scanning electron microscope (SEM) images of monocolpate pollen of Kempia longicolpites gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Pollen grains showing the very long distal aperture and loosely attached reticulum; c, d) Pollen grains showing the very long, clearly defined distal aperture that extends around the ends of the grain; e) Reticulum in nonapertural region showing the smooth muri loosely attached to the smooth surface of the foot layer by short columellae; f) Internal view of reticulum showing the short columellae loosened from the foot layer. Specimen, TV44-S105018 (holotype). Scale bars 300 Μm (a), 30 Μm (b), 6 Μm (c, d), 3 Μm (e, f). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 43. Scanning electron microscope (SEM) images of monocolpate pollen of Kempia longicolpites gen. et sp. nov. from a fragmentary stamen; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure; b) Pollen grains showing the very long distal aperture and loosely attached reticulum; c, d) Pollen grains showing the very long, clearly defined distal aperture that extends around the ends of the grain; e) Reticulum in nonapertural region showing the smooth muri loosely attached to the smooth surface of the foot layer by short columellae; f) Internal view of reticulum showing the short columellae loosened from the foot layer. Specimen, TV44-S105018 (holotype). Scale bars 300 Μm (a), 30 Μm (b), 6 Μm (c, d), 3 Μm (e, f).
Text-fig. 46. Scanning electron microscope (SEM) images of monocolpate pollen of Teebacia hughesii gen. et sp. nov. pollen from a stamen fragment; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure (d, e); b) Detail of detached reticulum showing inner surface of muri and scattered columellae; note the finely granular covering of the muri and columellae; c, d) Detail of reticulum showing the outer surface of muri with supratectal ornamentation of narrow ridges; note orbicules attached to the reticulum (d); e, f) Pollen grains showing loose, beaded, reticulum with long columellae; note continuous muri bordering the apertures and densely spaced minute orbicules lining the inner surface of the anther wall (e, arrowheads). Specimens, TV44-S136666 (holotype; a, d, e), TV44-S149207 (b, c, f). Scale bars 300 Μm (a), 6 Μm (e, f), 3 Μm (b), 1.5 Μm (c), 1.2 Μm (d). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 46. Scanning electron microscope (SEM) images of monocolpate pollen of Teebacia hughesii gen. et sp. nov. pollen from a stamen fragment; Torres Vedras locality, Portugal. a) Holotype; stamen fragment showing elongated pollen sacs that yielded the pollen in this Text-figure (d, e); b) Detail of detached reticulum showing inner surface of muri and scattered columellae; note the finely granular covering of the muri and columellae; c, d) Detail of reticulum showing the outer surface of muri with supratectal ornamentation of narrow ridges; note orbicules attached to the reticulum (d); e, f) Pollen grains showing loose, beaded, reticulum with long columellae; note continuous muri bordering the apertures and densely spaced minute orbicules lining the inner surface of the anther wall (e, arrowheads). Specimens, TV44-S136666 (holotype; a, d, e), TV44-S149207 (b, c, f). Scale bars 300 Μm (a), 6 Μm (e, f), 3 Μm (b), 1.5 Μm (c), 1.2 Μm (d).
Text-fig. 42. Scanning electron microscope (SEM) images of monocolpate pollen of Piercipollis sp. 1 (a–c) from an isolated anther and Piercipollis sp. 2 (d–f) from a coprolite (not shown); Torres Vedras locality, Portugal. a) Narrow elongate anther that yielded the pollen in (b) and (c); b) Distal view of pollen grain showing well-developed reticulum that is only loosely attached to the foot layer; note the smooth foot layer (arrowhead) of a grain from which the reticulum has become detached; c) Reticulum showing the smooth muri and long columellae that are mostly detached from the foot layer; d–f) Pollen grains in distal (d, e), and lateral views (f) showing the very long colpus and the well-developed reticulum only loosely attached to the smooth surface of the foot layer; note the main body of the grains (foot layer) does not fill out the whole space of the reticulum (f). Specimens, TV44-S148218 (a–c), TV142-S170216 (d–f). Scale bars 300 Μm (a), 6 Μm (b, d–f), 3 Μm (c). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 42. Scanning electron microscope (SEM) images of monocolpate pollen of Piercipollis sp. 1 (a–c) from an isolated anther and Piercipollis sp. 2 (d–f) from a coprolite (not shown); Torres Vedras locality, Portugal. a) Narrow elongate anther that yielded the pollen in (b) and (c); b) Distal view of pollen grain showing well-developed reticulum that is only loosely attached to the foot layer; note the smooth foot layer (arrowhead) of a grain from which the reticulum has become detached; c) Reticulum showing the smooth muri and long columellae that are mostly detached from the foot layer; d–f) Pollen grains in distal (d, e), and lateral views (f) showing the very long colpus and the well-developed reticulum only loosely attached to the smooth surface of the foot layer; note the main body of the grains (foot layer) does not fill out the whole space of the reticulum (f). Specimens, TV44-S148218 (a–c), TV142-S170216 (d–f). Scale bars 300 Μm (a), 6 Μm (b, d–f), 3 Μm (c).
Text-fig. 40. Scanning electron microscope (SEM) images of monocolpate pollen of Eckhartiopsis parva gen. et sp. nov. from an anther; Torres Vedras locality, Portugal. a) Holotype; elongated anther that yielded the pollen in this Text-figure; b–e) Pollen grains showing dense reticulate tectum supported by short columellae that are only loosely attached to the smooth surface of the foot layer: note dentate orbicules on inner surface of anther wall (b, arrowhead); f) Reticulate tectum showing smooth muri with small adhering orbicules (arrowhead); g) Pollen grains showing one in which the reticulum has been lost exposing the smooth surface of the foot layer; h) Fractured pollen wall showing reticulum, short columellae and thick foot layer; note granules on the inner surface of the apertural region (arrow). Specimen, TV39-S170217 (holotype). Scale bars 300 Μm (a), 6 Μm (b–e, g), 3 Μm (f, h). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 40. Scanning electron microscope (SEM) images of monocolpate pollen of Eckhartiopsis parva gen. et sp. nov. from an anther; Torres Vedras locality, Portugal. a) Holotype; elongated anther that yielded the pollen in this Text-figure; b–e) Pollen grains showing dense reticulate tectum supported by short columellae that are only loosely attached to the smooth surface of the foot layer: note dentate orbicules on inner surface of anther wall (b, arrowhead); f) Reticulate tectum showing smooth muri with small adhering orbicules (arrowhead); g) Pollen grains showing one in which the reticulum has been lost exposing the smooth surface of the foot layer; h) Fractured pollen wall showing reticulum, short columellae and thick foot layer; note granules on the inner surface of the apertural region (arrow). Specimen, TV39-S170217 (holotype). Scale bars 300 Μm (a), 6 Μm (b–e, g), 3 Μm (f, h).
Text-fig. 37. Scanning electron microscope (SEM) images of anthers and monocolpate pollen of Eckhartia brevicolumella gen. et sp. nov. from stamen fragment; Torres Vedras locality, Portugal. a) Dithecate, tetrasporangiate anther partially split along the elongated sporangia; b) Dithecate, tetrasporangiate anther; c) Holotype; distal view of pollen grain from holotype showing the long colpus with a distinct margin and a heterobrochate reticulum; d) Distal view of pollen grain from a stamen fragment showing long colpus, colpus margin and heterobrochate reticulum; e) Inner surface of anther wall from stamen fragment showing densely spaced dentate orbicules; f) Lateral view of pollen grains from stamen showing the psilate to foveolate-microreticulate colpus margin; g) Detail of pollen grain from holotype showing the slightly rounded profile of the muri, the short, densely spaced columellae and the dentate orbicules. Specimens, TV44-S136662 (a), TV44-S149204 (b), TV44-S136760 (holotype; c, g), TV44-S148021 (d, e), TV44-S136752 (f). Scale bars 300 Μm (a, b), 6 Μm (c, d, f), 3 Μm (e, g). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 37. Scanning electron microscope (SEM) images of anthers and monocolpate pollen of Eckhartia brevicolumella gen. et sp. nov. from stamen fragment; Torres Vedras locality, Portugal. a) Dithecate, tetrasporangiate anther partially split along the elongated sporangia; b) Dithecate, tetrasporangiate anther; c) Holotype; distal view of pollen grain from holotype showing the long colpus with a distinct margin and a heterobrochate reticulum; d) Distal view of pollen grain from a stamen fragment showing long colpus, colpus margin and heterobrochate reticulum; e) Inner surface of anther wall from stamen fragment showing densely spaced dentate orbicules; f) Lateral view of pollen grains from stamen showing the psilate to foveolate-microreticulate colpus margin; g) Detail of pollen grain from holotype showing the slightly rounded profile of the muri, the short, densely spaced columellae and the dentate orbicules. Specimens, TV44-S136662 (a), TV44-S149204 (b), TV44-S136760 (holotype; c, g), TV44-S148021 (d, e), TV44-S136752 (f). Scale bars 300 Μm (a, b), 6 Μm (c, d, f), 3 Μm (e, g).
Text-fig. 39. Scanning electron microscope (SEM) images of monocolpate pollen of Eckhartia intermedia sp. nov. from a pollen clump, probably fragment of a stamen (a–e) and anthers and pollen of Eckhartia sp. (f–k); Torres Vedras locality, Portugal. a) Pollen clump that yielded the pollen in (b–e); b–d) Pollen grains viewed from distal (d) and proximal surfaces (b, c) showing the long colpus and well-developed reticulum with large and small luminae; e) Reticulum showing smooth muri supported by short columellae firmly attached to the smooth surface of the foot layer; note the occasional small luminae; f, g) Fragments of narrow elongate anthers with same kind of pollen as in (i); h) Inner surface of anther wall showing small spherical orbicules; in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 39. Scanning electron microscope (SEM) images of monocolpate pollen of Eckhartia intermedia sp. nov. from a pollen clump, probably fragment of a stamen (a–e) and anthers and pollen of Eckhartia sp. (f–k); Torres Vedras locality, Portugal. a) Pollen clump that yielded the pollen in (b–e); b–d) Pollen grains viewed from distal (d) and proximal surfaces (b, c) showing the long colpus and well-developed reticulum with large and small luminae; e) Reticulum showing smooth muri supported by short columellae firmly attached to the smooth surface of the foot layer; note the occasional small luminae; f, g) Fragments of narrow elongate anthers with same kind of pollen as in (i); h) Inner surface of anther wall showing small spherical orbicules;
Text-fig. 38. Scanning electron microscope (SEM) images of anthers and monocolpate pollen of Eckhartia longicolumella sp. nov. from stamen fragment; Torres Vedras locality, Portugal. a) Dithecate, tetrasporangiate anther that yielded the same kind of pollen as the holotype (b); b) Holotype; stamen fragment that yielded the pollen shown in this Text-figure; c–e) Pollen grains showing the elongated colpi, reticulum with large and small luminae and continuous muri along the aperture margins; f, g) Pollen grains in in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 38. Scanning electron microscope (SEM) images of anthers and monocolpate pollen of Eckhartia longicolumella sp. nov. from stamen fragment; Torres Vedras locality, Portugal. a) Dithecate, tetrasporangiate anther that yielded the same kind of pollen as the holotype (b); b) Holotype; stamen fragment that yielded the pollen shown in this Text-figure; c–e) Pollen grains showing the elongated colpi, reticulum with large and small luminae and continuous muri along the aperture margins; f, g) Pollen grains in
Text-fig. 8. Scanning electron microscope (SEM) images of spores from clumps of spores and sporangia with affinities to polypodiopsids (a–c) and of uncertain affinity (d–k); Torres Vedras locality, Portugal. a) Folded Cyathidites minor spores in proximal view showing trilete mark, from clump of spores; b) Cyathidites minor spores in proximal view showing trilete mark, from group of sporangia; c) Cyathidites australis spores in proximal view showing trilete mark, from group of sporangia; d–f) Linear group of spore masses (d; probable sporangial contents) composed of Taurocusporites segmentatus spores showing distal surface (e, middle) with concentric regions and proximal surface with segmented laesurae of elongated granules (e, right; f); in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 8. Scanning electron microscope (SEM) images of spores from clumps of spores and sporangia with affinities to polypodiopsids (a–c) and of uncertain affinity (d–k); Torres Vedras locality, Portugal. a) Folded Cyathidites minor spores in proximal view showing trilete mark, from clump of spores; b) Cyathidites minor spores in proximal view showing trilete mark, from group of sporangia; c) Cyathidites australis spores in proximal view showing trilete mark, from group of sporangia; d–f) Linear group of spore masses (d; probable sporangial contents) composed of Taurocusporites segmentatus spores showing distal surface (e, middle) with concentric regions and proximal surface with segmented laesurae of elongated granules (e, right; f);
Text-fig. 23. Scanning electron microscope (SEM) images of Clavatipollenites sp. 2 (a–c) from a fragmentary stamen, and stamen fragments with in situ pollen of Clavatipollenites sp. 3 (d–j); Torres Vedras locality, Portugal. a, b) Distal (a) and proximal (b) view of pollen showing simple, elongate colpus on distal surface and semitectate-reticulate pollen wall; c) Detail of pollen wall showing muri with finely verrucate supratectal ornamentation and long scattered columellae; d) Fragment of tetrasporangiate stamen; e, f) Distal views of pollen from stamen fragment showing poorly defined aperture, coarse reticulum and long scattered columellae; g) Fragment of stamen; h) Distal view of pollen showing poorly defined aperture covered by irregular verrucae; i, j) Pollen wall showing rounded orbicules (i) and fractured pollen wall showing long scattered columellae (j). Specimens, TV43-S136728 (a–c), TV44-S149201 (d–f), TV44-S149220 (g–j). Scale bars 300 Μm (d, g), 6 Μm (a, b, e, f, h), 3 Μm (c), 1.5 Μm (i, j). in The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community
Text-fig. 23. Scanning electron microscope (SEM) images of Clavatipollenites sp. 2 (a–c) from a fragmentary stamen, and stamen fragments with in situ pollen of Clavatipollenites sp. 3 (d–j); Torres Vedras locality, Portugal. a, b) Distal (a) and proximal (b) view of pollen showing simple, elongate colpus on distal surface and semitectate-reticulate pollen wall; c) Detail of pollen wall showing muri with finely verrucate supratectal ornamentation and long scattered columellae; d) Fragment of tetrasporangiate stamen; e, f) Distal views of pollen from stamen fragment showing poorly defined aperture, coarse reticulum and long scattered columellae; g) Fragment of stamen; h) Distal view of pollen showing poorly defined aperture covered by irregular verrucae; i, j) Pollen wall showing rounded orbicules (i) and fractured pollen wall showing long scattered columellae (j). Specimens, TV43-S136728 (a–c), TV44-S149201 (d–f), TV44-S149220 (g–j). Scale bars 300 Μm (d, g), 6 Μm (a, b, e, f, h), 3 Μm (c), 1.5 Μm (i, j).
Text-fig. 3. Pazlia hilaris gen. et sp. nov. (a–e) from the Early Cretaceous Famalicão locality (sample 025), Portugal (holotype, S175096) and Pazliopsis reyi gen. et sp. nov. (f–i) from the Early Cretaceous Torres Vedras locality, Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings a–f, i) and scanning electron microscopy (SEM, g, h). a, b) Seed in lateral (a) and oblique apical (b) views showing the truncate hilar-micropylar region; note prominent hilar scar (hi) and micropyle (mi) at the seed apex and the raphe (ra) seen as slightly raised ridge; remains of mounting media (¤). c) Cut volume rendering (cut at yz0647) showing course of raphe (ra), hilar scar (hi) and micropyle (mi); note the strongly radially elongated cells below the hilar scar. d) Seed in antiraphal view. e) Seed surface showing the raised undulate anticlinal walls of the exotestal cells. f) Seed enclosed in remains of thin-walled fruit (fr) (S174632, Torres Vedras sample 298). g) Holotype, seed enclosed in remains of fruit (fr); raphal view showing the faintly ribbed surface of the seed (S171534, Torres Vedras sample 043). h) Apical view of seed fragment showing hilar scar (hi), position of raphe (ra) and the ribbed seed surface (S136683, Torres Vedras sample 044). i) Seed surface showing the raised undulate anticlinal walls of the exotestal cells (S171534; Torres Vedras sample 043). Scale bars = 250 µm (a–d, f–h); 125 µm (e, i). in Extinct Taxa Of Exotestal Seeds Close To Austrobaileyales And Nymphaeales From The Early Cretaceous Of Portugal
Text-fig. 3. Pazlia hilaris gen. et sp. nov. (a–e) from the Early Cretaceous Famalicão locality (sample 025), Portugal (holotype, S175096) and Pazliopsis reyi gen. et sp. nov. (f–i) from the Early Cretaceous Torres Vedras locality, Portugal; Synchrotron radiation X-ray tomographic microscopy (SRXTM, volume renderings a–f, i) and scanning electron microscopy (SEM, g, h). a, b) Seed in lateral (a) and oblique apical (b) views showing the truncate hilar-micropylar region; note prominent hilar scar (hi) and micropyle (mi) at the seed apex and the raphe (ra) seen as slightly raised ridge; remains of mounting media (¤). c) Cut volume rendering (cut at yz0647) showing course of raphe (ra), hilar scar (hi) and micropyle (mi); note the strongly radially elongated cells below the hilar scar. d) Seed in antiraphal view. e) Seed surface showing the raised undulate anticlinal walls of the exotestal cells. f) Seed enclosed in remains of thin-walled fruit (fr) (S174632, Torres Vedras sample 298). g) Holotype, seed enclosed in remains of fruit (fr); raphal view showing the faintly ribbed surface of the seed (S171534, Torres Vedras sample 043). h) Apical view of seed fragment showing hilar scar (hi), position of raphe (ra) and the ribbed seed surface (S136683, Torres Vedras sample 044). i) Seed surface showing the raised undulate anticlinal walls of the exotestal cells (S171534; Torres Vedras sample 043). Scale bars = 250 µm (a–d, f–h); 125 µm (e, i).
Fig. 6. Intermediate, elongated spermatids. A in Comparison of spermiogenesis in the externally fertilizing Hemigrammus erythrozonus and the inseminating Corynopoma riisei (Teleostei: Characiformes: Characidae)
Fig. 6. Intermediate, elongated spermatids. A - Longitudinally sectioned spermatid with elongate nucleus containing highly granular chromatin and flagellum running within cytoplasmic canal. Scale bar 0.5 µm. B - inset: Arrangement of proximal and distal centrioles with intercentriolar lamellated body. Scale bar 0.25 µm. C - Transverse sections of tip of spermatids containing proximal and distal centrioles and anterior part of flagellum. Scale bar 0.5 µm D - Transverse sections of the middle portion of nucleus with mitochondria located around the nucleus. E - Posterior portion of nucleus showing numerous mitochondria, and flagellum contained within cytoplasmic canal. F - Flagellum within the cytoplasmic canal of the mitochondrial region. Scale bar 0.5 µm. cc – cytoplasmic canal, cs – cytoplasmic sleeve, dc – distal centriole, f – flagellum, ilb – intercentriolar lamellated body, m – mitochondrion, N – nucleus, pc – proximal centriole, v – vesicles.
Text-fig. 38. Scanning electron microscope (SEM) images of "Stamen with tricolpate pollen sp. 2"; Catefica locality, Portugal. a) Stamen fragment showing the elongate, tetrasporangiate anther but with the base and apex poorly preserved; b, d, e) Pollen grains from stamen fragment in equatorial (b, e) and polar (d) views showing the long colpi with coarsely verrucate aperture membranes and semitectate-microreticulate pollen wall; note small, spherical orbicules scattered on the surface of the tectum (arrows); c) Detail of pollen wall showing smooth muri with very faint transverse striations and a granular to columellate infratectal layer. Specimen, Catefica 153-S105614 (a–e). Scale bars = 600 Μm (a), 6 Μm (b, d, e), 1 Μm (c). in The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms
Text-fig. 38. Scanning electron microscope (SEM) images of "Stamen with tricolpate pollen sp. 2"; Catefica locality, Portugal. a) Stamen fragment showing the elongate, tetrasporangiate anther but with the base and apex poorly preserved; b, d, e) Pollen grains from stamen fragment in equatorial (b, e) and polar (d) views showing the long colpi with coarsely verrucate aperture membranes and semitectate-microreticulate pollen wall; note small, spherical orbicules scattered on the surface of the tectum (arrows); c) Detail of pollen wall showing smooth muri with very faint transverse striations and a granular to columellate infratectal layer. Specimen, Catefica 153-S105614 (a–e). Scale bars = 600 Μm (a), 6 Μm (b, d, e), 1 Μm (c).
Text-fig. 37. Scanning electron microscope (SEM) images of "Stamen with tricolpate pollen sp. 1"; Catefica locality, Portugal. a) Stamen fragment showing the elongate, tetrasporangiate anther but with the base and apex poorly preserved; b–d) Pollen grains from stamen fragment in equatorial view showing the long colpi and well-developed, heterobrochate reticulum with very distinct large and small lumina; e) Detail of pollen wall showing orbicules with irregular projections; f) Detail of pollen wall showing smooth muri supported by short, densely spaced columellae. Specimen, Catefica 50-S170419 (a–f). Scale bars = 600 Μm (a), 6 Μm (b–d), 1.5 Μm (e, f). in The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms
Text-fig. 37. Scanning electron microscope (SEM) images of "Stamen with tricolpate pollen sp. 1"; Catefica locality, Portugal. a) Stamen fragment showing the elongate, tetrasporangiate anther but with the base and apex poorly preserved; b–d) Pollen grains from stamen fragment in equatorial view showing the long colpi and well-developed, heterobrochate reticulum with very distinct large and small lumina; e) Detail of pollen wall showing orbicules with irregular projections; f) Detail of pollen wall showing smooth muri supported by short, densely spaced columellae. Specimen, Catefica 50-S170419 (a–f). Scale bars = 600 Μm (a), 6 Μm (b–d), 1.5 Μm (e, f).
Text-fig. 22. Scanning electron microscope (SEM, a–c) and synchrotron radiation X-ray tomographic microscopy (SRXTM, d) images of Ibericarpus cuneiformis gen. et sp. nov.; Catefica locality, Portugal. a) Fruiting axis bearing an elongated receptacle with numerous diamond-shaped scars from detached fruitlets; note the absence of scars from bracts, tepals or stamens at the transition to the fruitlet scars and the stalk (arrow); b) Group of ten fruitlets detached from fruiting axis in (a) showing apical stigmatic region and distinctive bulging isodiametric epidermal cells; c) Detached fruitlet showing apical stigmatic region; d) Volume rendering of three adhering fruits showing apical stigmatic region and distinctive bulging isodiametric epidermal cells. Specimens, Catefica MM75-P0477 (a, b), Catefica 49-S115852 (c), Catefica 50-S174907 (d). Scale bars = 300 Μm (a–d). in The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms
Text-fig. 22. Scanning electron microscope (SEM, a–c) and synchrotron radiation X-ray tomographic microscopy (SRXTM, d) images of Ibericarpus cuneiformis gen. et sp. nov.; Catefica locality, Portugal. a) Fruiting axis bearing an elongated receptacle with numerous diamond-shaped scars from detached fruitlets; note the absence of scars from bracts, tepals or stamens at the transition to the fruitlet scars and the stalk (arrow); b) Group of ten fruitlets detached from fruiting axis in (a) showing apical stigmatic region and distinctive bulging isodiametric epidermal cells; c) Detached fruitlet showing apical stigmatic region; d) Volume rendering of three adhering fruits showing apical stigmatic region and distinctive bulging isodiametric epidermal cells. Specimens, Catefica MM75-P0477 (a, b), Catefica 49-S115852 (c), Catefica 50-S174907 (d). Scale bars = 300 Μm (a–d).
Text-fig. 3. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images of fruits of Canrightia foveolata sp. nov.; Catefica locality, Portugal. a) Volume rendering of fruit showing prominent rim around the middle of the fruit with reduced tepals (arrowheads) and partly abraded fruit wall exposing the pitted endotesta surface of one of two seeds (arrow); note two of the vascular bundles (vb) extending from the base of the fruit to the tepals; b) Voltex of fruit showing prominent rim around the fruit (arrowhead) and dense precipitation of crystals in the endothelium cells of one of the two seeds in the fruit; c) Longitudinal section of fruit (orthoslice yz0520) showing the inferred hypanthium rim (arrow head) and two seeds, one with a dense precipitation of crystals; note the prominent endothelium cells (asterisks) of the inner integument and the well-developed fruit wall above the seeds; d) Transverse section through basal part of fruit and seeds close to the micropyle (orthoslice xy0312) showing partly abraded fruit wall with five vascular bundles (vb) and details of the seed coat with endotesta (oi-end) surrounding the tegmen consisting of an outer epidermis (ii-o), middle layer (ii-m) and a distinct inner epidermis (endothelium) consisting of radially elongated cells (asterisk); e) Transverse section (orthoslice xy1680) through apical part of the fruit close to chalaza showing the tips of two seeds; note the endotesta (oi-end) surrounded by thick-walled cells of the exotesta (oi-o); f) Transverse section (orthoslice xy1485) through fruit in the region of the hypanthium rim showing sections through the two seeds close to the chalazal region; note endotesta (oi-end) surrounded by larger cells of exotesta (oi-o) and fruit wall (fr). Specimen, Catefica 49-S174249 (holotype, a–f). Scale bars = 300 Μm (a–c, e, f), 100 Μm (d). in The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms
Text-fig. 3. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images of fruits of Canrightia foveolata sp. nov.; Catefica locality, Portugal. a) Volume rendering of fruit showing prominent rim around the middle of the fruit with reduced tepals (arrowheads) and partly abraded fruit wall exposing the pitted endotesta surface of one of two seeds (arrow); note two of the vascular bundles (vb) extending from the base of the fruit to the tepals; b) Voltex of fruit showing prominent rim around the fruit (arrowhead) and dense precipitation of crystals in the endothelium cells of one of the two seeds in the fruit; c) Longitudinal section of fruit (orthoslice yz0520) showing the inferred hypanthium rim (arrow head) and two seeds, one with a dense precipitation of crystals; note the prominent endothelium cells (asterisks) of the inner integument and the well-developed fruit wall above the seeds; d) Transverse section through basal part of fruit and seeds close to the micropyle (orthoslice xy0312) showing partly abraded fruit wall with five vascular bundles (vb) and details of the seed coat with endotesta (oi-end) surrounding the tegmen consisting of an outer epidermis (ii-o), middle layer (ii-m) and a distinct inner epidermis (endothelium) consisting of radially elongated cells (asterisk); e) Transverse section (orthoslice xy1680) through apical part of the fruit close to chalaza showing the tips of two seeds; note the endotesta (oi-end) surrounded by thick-walled cells of the exotesta (oi-o); f) Transverse section (orthoslice xy1485) through fruit in the region of the hypanthium rim showing sections through the two seeds close to the chalazal region; note endotesta (oi-end) surrounded by larger cells of exotesta (oi-o) and fruit wall (fr). Specimen, Catefica 49-S174249 (holotype, a–f). Scale bars = 300 Μm (a–c, e, f), 100 Μm (d).
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