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Fig. 7 in Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

Fig. 7. Stereopairs of Uchkudukodon nessovi (McKenna, Kielan−Jaworowska, and Meng, 2000); Dzharakuduk, Uzbekistan, Late Cretaceous (Turonian). A. URBAC 04−181, m2 from right dentary with m2 and alveoli for p5, m1, 3; in lingual (A1), occlusal (A2), and labial (A3) views. B. URBAC 04−42, right denary with alveoli for p2–5, m1–3 and angular process, in occlusal view. C. ZIN 84967, m2 from right dentary with worn m2 and alveoli for m3; occlusal. D. URBAC 98−122, m3 from right dentary with worn m3, in occlusal view.

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Fig. 10 in Bone microstructure and growth patterns of early mammals

Fig. 10. Heterocephalus glaber Rüppell, 1842. Recent, ZOO/UCT no number; femur, section catalogue number, ZOO/UCT/192, showing the wellvascularized compacta and the narrow outer region of parallel fibred bone (OCL).

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Fig. 6 in Revision of a pretribosphenic mammal Arguimus from the Early Cretaceous of Mongolia

Fig. 6. Known specimens of Arguimus khosbajari Dashzeveg, 1979 (Höövör, Mongolia; Early Cretaceous) drawn at the same scale in labial view, with interpretation of preserved dentition. Reversed images are marked by asterisk.

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Fig. 2. Arguimus khosbajari Dashzeveg, 1979. PIN 3101 in Revision of a pretribosphenic mammal Arguimus from the Early Cretaceous of Mongolia

Fig. 2. Arguimus khosbajari Dashzeveg, 1979. PIN 3101/400, left dentary fragment with m1 and alveoli for p2–5 and m2–3. Höövör, Mongolia; Early Cretaceous. A–C. The m1 in labial (A), lingual (B), and occlusal (C, stereopair) views. D–F. The whole specimen in lingual (D), occlusal (E), and labial (F) views. Scale bars 1 mm.

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Fig. 9 in Bone microstructure and growth patterns of early mammals

Fig. 9. Barunlestes butleri Kielan−Jaworowska, 1975,?late Campanian, Baruungoyot Formation, Khulsan, Nemegt Basin, Gobi Desert, Mongolia. ZPAL MgM−I/77, section catalogue number, ZOO/UCT/189. A. Transverse section of a rib, showing the poorly vascularized compacta and the parallel fibred nature of the compacta, as well as several lines of arrested growth (arrows). B. The femur from the same ZPAL specimen, section catalogue number ZOO/UCT/189 (on same slide as rib). B1. Partial transverse section of the femur on the same slide showing the similarly poorly vascularized compacta of parallel fibred bone in the bone wall. B2. Higher magnification of the femur in B1. Small arrows in B2 indicate lines of arrested growth; large arrow indicates the more globular haphazardly oriented osteocyte lacunae.

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Fig. 8 in Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

Fig. 8. Stereopairs of Bulaklestes kezbe Nessov, 1985; Dzharakuduk, Uzbekistan, Late Cretaceous (Turonian). A. 1985. CCMGE 19/12176, right P4; in labial (A1) and occlusal (A2) views. B. ZIN 82591, left M1 lacking paracone; in occlusal (B1), labial (B2), and anterior (B3) views. C. CCMGE 12/12176, type, right M3; in occlusal (C1), labial (C2), and anterior (C3) views.

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Fig. 8 in Bone microstructure and growth patterns of early mammals

Fig. 8. Zalambdalestes lechei Gregory and Simpson, 1926;?early Campanian, Djadokhta Formation, Bayan Zak, Gobi Desert, Mongolia, ZPAL MgM−I/43, transverse section of a femur (section catalogue number, ZOO/ UCT/186). A. Low magnification showing the relatively thin bone wall and the relatively poorly vascularized parallel−fibred compacta. B. Higher magnification of the same, showing the parallel organization of the osteocyte lacunae and the vascularization of the bone. Arrow indicates a single LAG located near the peripheral margin of the bone wall. C. Still higher magnification of the same section, showing some simple vascular channels and primary osteons (arrows).

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Fig. 7 in Bone microstructure and growth patterns of early mammals

Fig. 7. Nemegtbaatar gobiensis Kielan−Jaworowska, 1974,?late Campanian, red beds of Khermeen Tsav, Khermeen Tsav II, Gobi Desert, Mongolia. ZPAL MgM−I/81, section catalogue number, ZOO/UCT/183. A. Higher magnification of framed region in Fig. 6B4 showing the complex nature of the compacta. Region 1 is a periosteal deposition of parallel−fibred bone, region 2 is a compaction of fine cancellous bone. Arrow indicates the tide line. B. Higher magnification of framed region in Fig. 6B2 (section catalogue number, ZOO/UCT/181), illustrating the true woven bone. Arrow indicates the tide line.

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Fig. 4 in Bone microstructure and growth patterns of early mammals

Fig. 4. Ornithorhynchus anatinus (Shaw, 1799); Recent, ZOO/UCT no number, femur, section catalogue number, ZOO/UCT/194. Transverse section showing the extensive fibro−lamellar bone and the episodically formed lamellar deposits of bone.

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Fig. 5 in Bone microstructure and growth patterns of early mammals

Fig. 5. Kryptobaatar dashzevegi Kielan−Jaworowska, 1971?early Campanian, Djadokhta Formation, Bayan Zak, Gobi Desert, Mongolia, femur (ZPAL MgM−I/18; Section catalogue number, ZOO/UCT/184). A. Low magnification of the femur showing the overall structure of the cross section. Note the woven bone nature of the thin bone wall. B. A higher magnification of the same section, illustrating the channels in the bone, and the irregular peripheral and medullary margins of the bone wall.

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Fig. 13 in Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

Fig. 13. Strict consensus tree of three equally parsimonious trees produced by reweighting the 33 characters listed in the text using the maximum value of the rescaled consistency indices, resulting in more internal resolution of Asioryctitheria. The single numbers on each tree are bootstrap values greater than 50 that were each produced from 10,000 replicate runs. The characters and state changes are those listed and discussed in the text.

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Fig. 3 in Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

Fig. 3. Stereopairs of Daulestes inobservabilis (Nessov, 1982); Dzharakuduk, Uzbekistan, Late Cretaceous (Turonian). A. URBAC 98−146, right P4 missing small part of anterior margin, in labial (A1) and occlusal (A2) views. B. URBAC 02−91, P5 and M1 from right maxilla with P5, M1, and alveoli for P4 and M2–3, in labial (B1), occlusal (B2), and lingual (B3) views. C. URBAC 98−140, left M2, in occlusal (C1), labial (C2), and anterior (C3) views.

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Fig. 3 in Bone microstructure and growth patterns of early mammals

Fig. 3. Sections through bones of a lepidosaurian Gephyrosaurus sp. (A) and morganucodontan Morganucodon watsoni Kühne, 1949 (B, C), both from Early Jurassic, Pant Quarry, Wales. A. Gephyrosaurus sp. section of a femur (UCL.ADB no number; section catalogue number ZOO/UCT/193), showing the parallel−fibred nature of the compacta, and several lines of arrested growth (LAGs). Arrow indicates the embryonic bone. B, C. Morganucodon watsoni. B. Ulna, UCL.ADB no number; section catalogue number ZOO/UCT/132. B1. Transverse section showing the predominantly woven nature of the compacta, and the general organization of the compacta. Arrow indicates the parallel−fibred bone in the localized area. B2. A higher magnification of the framed area in B1, showing the large erosion cavities, and the irregular orientation of the globular osteocyte lacunae. C. Femur (UCL.ADB no number; section catalogue number ZOO/UCT/164); transverse section showing the altered nature of most of the bone.

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Fig. 1 in Bone microstructure and growth patterns of early mammals

Fig. 1. Cladogram showing the phylogenetic relationship of the taxa studied (simplified from Kielan−Jaworowska et al. 2004).

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Fig. 6 in Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

Fig. 6. Stereopairs of Uchkudukodon nessovi (McKenna, Kielan−Jaworowska, and Meng, 2000); Dzharakuduk, Uzbekistan, Late Cretaceous (Turonian). A. URBAC 98−124, M1 from left maxilla with M1 and alveoli or partial alveoli for P5 and M2, in occlusal (A1), labial (A2), and anterior (A3) views. B. URBAC 03−213, right M2, in occlusal (B1), labial (B2), and anterior (B3) views.

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Fig. 2 in Bone microstructure and growth patterns of early mammals

Fig. 2. Tritylodon sp. Early Jurassic, Massospondylus Range Zone, Elliot Formation, South Africa. A. Humerus, BPI/5671 section Ty4B2, showing the fibro−lamellar nature of the compacta. B. Radius, BPI/5167, section Ty1BII3, showing the peripheral development of parallel fibred bone with rest lines (arrows).

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Fig. 5. Arguimus khosbajari Dashzeveg, 1979. PIN 3101 in Revision of a pretribosphenic mammal Arguimus from the Early Cretaceous of Mongolia

Fig. 5. Arguimus khosbajari Dashzeveg, 1979. PIN 3101/108, left dentary fragment with m3–4, alveoli for m2, and partial coronoid process in posterior view. Höövör, Mongolia; Early Cretaceous. Scale bar 1 mm.

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Fig. 6 in Bone microstructure and growth patterns of early mammals

Fig. 6. Nemegtbaatar gobiensis Kielan−Jaworowska, 1974;?late Campanian, red beds of Khermeen Tsav, Khermeen Tsav II, Gobi Desert, Mongolia. A. Reconstruction of the femur in anterior view. showing the region (notations B1, B2, B3, B4) from which the sections shown in B have been taken. B. Successive sections of the femur from the upper part of the shaft of the specimen ZPAL MgM−I/81. B1 section catalogue number, ZOO/UCT/ 180; B2 section catalogue number, ZOO/UCT/ 181; B3 section catalogue number, ZOO/UCT/182; and B4 section catalogue number, ZOO/UCT/183. Note the distinct changes in the nature of the compacta and shape of the medullary cavity and overall shape of the cross section from B1 to B4. Detail of framed regions in B2 and B4 are provided in Fig. 7.

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Fig. 4. Arguimus khosbajari Dashzeveg, 1979 in Revision of a pretribosphenic mammal Arguimus from the Early Cretaceous of Mongolia

Fig. 4. Arguimus khosbajari Dashzeveg, 1979. Höövör, Mongolia; Early Cretaceous. A. PIN 3101/108, left dentary fragment with m3–4, alveoli for m2, and partial coronoid process (A1, m3–4 in occlusal view, stereopair; A2–A4, the whole specimen in occlusal, labial, and lingual views). B. PIN 3101/107, right dentary fragment with worn m2 and alveoli for m3–4 (B1, in occlusal view, stereopair; B2–B4, the whole specimen in occlusal, labial, and lingual views). Scale bars 1 mm.

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Fig. 12 in Late Cretaceous asioryctitherian eutherian mammals from Uzbekistan and phylogenetic analysis of Asioryctitheria

Fig. 12. Labial views, lower anterior dentitions of Bulaklestes kezbe Nessov, 1985 (A) and of Uchkudukodon nessovi (McKenna, Kielan−Jaworowska, and Meng, 2000) (B, drawing from McKenna et al. 2000: fig. 16D). Both from Dzharakuduk, Uzbekistan, Late Cretaceous (Turonian). Our interpretations of tooth identifications for both taxa are between the dentaries and those of McKenna et al. (2000) for U. nessovi are below the lower dentary.

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International Brain Laboratory public data

The International Brain Laboratory public data releases expose standardized mouse decision-making experiments, including Neuropixels recordings, widefield calcium imaging, behavior, and session metadata accessed through the ONE API.

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Last verified 2026-04-29Open record