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Figure 26 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 26. Trachyphyllia Milne Edwards & Haime, 1848a, is flabello-meandroid and free-living, with large (> 15 mm) and high-relief (> 6 mm) calices, septa in ≥ four cycles (≥ 48 septa) spaced
Figure 15. Hydnophora Fischer von Waldheim, 1807 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 15. Hydnophora Fischer von Waldheim, 1807, has uniserial corallites, monticules and fused walls, septa in
Figure 16 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 16. Leptoria Milne Edwards & Haime, 1848a, has uniserial corallites, fused walls, small (<4 mm) and low-relief (<3 mm) calices, septa in
Figure 14 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 14. Goniastrea Milne Edwards & Haime, 1848a, generally has discrete corallites, small to medium (≤ 15 mm) calices, equally thick costosepta, and well-developed paliform (uniaxial) and/or septal (multiaxial) lobes. Septal teeth are often low (<0.3 mm) and narrowly spaced (<0.3 mm). Walls formed by strong abortive septa and partial septotheca; trabeculothecal elements may be present. A–C, Goniastrea retiformis (Lamarck, 1816), type species of Goniastrea; macromorphology, holotype MNHN IK-2010-693, unknown locality (A); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype USNM 1013047 (FA1030), Saipan, Mariana Islands. D–F, Goniastrea edwardsi Chevalier, 1971; macromorphology, holotype MNHN IK-2010-654, Seychelles (D); micromorphology (E) and microstructure (F), hypotype RMNH 11194, Lizard Island, Australia. G–I, Goniastrea favulus (Dana, 1846); macromorphology (G), micromorphology (H), and microstructure (I), syntype USNM 66, Fiji. J–L, Goniastrea stelligera (Dana, 1846); macromorphology (J), micromorphology (K), and microstructure (L), syntype USNM 55, Fiji.
Figure 13. Favites Link, 1807 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 13. Favites Link, 1807, has discrete corallites with double or fused walls, septa generally in ≥ four cycles (≥ 48 septa), weak to well-developed paliform (uniaxial) lobes, and spongy columellae. Septal teeth with medium height (0.3– 0.6 mm) and spacing (0.3–1 mm). Walls formed by dominant paratheca and partial septotheca, with strong costa centre clusters and transverse septal crosses. A–C, Favites abdita (Ellis & Solander, 1786), type species of Favites; macromorphology, holotype GLAHM 104005, unknown locality (A; photo by K. G. Johnson); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype RMNH 10760, Pulau Air, Indonesia. D–F, Favites flexuosa (Dana, 1846); macromorphology, syntype USNM 27, Fiji (D); micromorphology (E) and microstructure (F), hypotype RMNH 14165, New Caledonia. G–I, Favites rotundata Veron, Pichon & Wijsman-Best, 1977; macromorphology (G) and micromorphology (H; photo by N. Santodomingo), holotype NHMUK 1977.1.1.6, southwest Swain Reefs, Australia; microstructure, hypotype MTQ G61874, Pelorus Island, Australia (I). J–L, Favites valenciennesi (Milne Edwards & Haime, 1849b); macromorphology, holotype MNHN IK-2010-696, unknown locality (J); micromorphology (K) and microstructure (L), hypotype UP P1L02131, Batangas, the Philippines.
Figure 5 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 5. Australogyra Veron & Pichon, 1982, is ramose and has uniserial corallites with few centres, fused walls, septa in
Figure 11. Echinopora Lamarck, 1816 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 11. Echinopora Lamarck, 1816, has discrete corallites that bud extracalicularly, extensive coenosteum (≥ corallite diameter), medium-size (4–15 mm) and low-relief (<3 mm) calices, large (≥ 1/4 of calice width) spongy columellae, and weak/moderate paliform (uniaxial) lobes. Septal teeth are low (<0.3 mm) with medium spacing (0.3–1 mm) and multiaxial tips. Walls formed by partial septotheca and weak abortive septa. A–C, Echinopora lamellosa (Esper, 1795), type species of Echinopora; macromorphology, Echinopora rosularia Lamarck, 1816, holotype of Echinopora MNHN IK-2010-635, unknown locality (A); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype USNM 89851, Alofi, Niue. D–F, Echinopora horrida Dana, 1846; macromorphology, syntype USNM 162, Fiji (D); micromorphology (E) and microstructure (F), hypotype RMNH 33956, Kepulauan Seribu, Indonesia. G, Echinopora mammiformis (Nemenzo, 1959); macromorphology, holotype UP C-99, Puerto Galera, the Philippines. H, I, Echinopora gemmacea (Lamarck, 1816); micromorphology, hypotype RMNH 17270, Watamu, Kenya (H); microstructure, hypotype USNM 1113168, Red Sea (I).
Figure 12 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 12. Erythrastrea Pichon, Scheer & Pillai in Scheer & Pillai, 1983, is phaceloid or flabello-meandroid, with equally thick costosepta spaced
Figure 8. Coelastrea Verrill, 1866 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 8. Coelastrea Verrill, 1866, has discrete corallites with double or fused walls, septa in ≥ four cycles (≥ 48 septa), regular free septa, and well-developed septal (multiaxial) lobes. Septal teeth with medium height (0.3–0.6 mm) and spacing (0.3–1 mm). Walls formed by dominant paratheca and partial septotheca, with strong costal and septal medial lines. A, Coelastrea tenuis Verrill, 1866, type species of Coelastrea; macromorphology, holotype YPM IZ 476, unknown locality. B–F, Coelastrea aspera (Verrill, 1866); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), syntype USNM 402, Ryukyu Islands, Japan; macromorphology, syntype USNM 403, Ryukyu Islands, Japan (D); micromorphology (E) and microstructure (F), hypotype RMNH 13844, Gulf of Aqaba, Israel. G–I, Coelastrea palauensis (Yabe & Sugiyama, 1936); macromorphology, holotype TIU 56631, Palau (G); micromorphology (H) and microstructure (I), hypotype RMNH 14084, New Caledonia.
Figure 1 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 1. Comparisons amongst recent classifications of reef corals examined in this study. Family level taxonomy follows Budd et al. (2012). See Stolarski & Roniewicz (2001) for comparisons with Vaughan & Wells (1943), Wells (1956), Alloiteau (1952), and Chevalier & Beauvais (1987).
Figure 9 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 9. Cyphastrea Milne Edwards & Haime, 1848a has discrete corallites that bud extracalicularly, small (<4 mm) and low-relief (<3 mm) calices, regular free septa, and compact columellae. Septal teeth are low (<0.3 mm) and narrowly spaced (<0.3 mm), with multiaxial tips; strong (pointed) granules scattered on septal face. Walls formed by dominant septotheca. A–C, Cyphastrea microphthalma (Lamarck, 1816), type species of Cyphastrea; macromorphology, holotype MNHN IK-2012–14002, unknown locality (A); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype RMNH 15111 (FA1035), southwest Sulawesi, Indonesia. D–F, Cyphastrea chalcidicum (Forskål, 1775); macromorphology, hypotype MTQ G61902, Orpheus Island, Australia (D); micromorphology (E) and microstructure (F), hypotype RMNH 12787, Gulf of Aqaba, Israel. G–I, Cyphastrea serailia (Forskål, 1775); macromorphology, syntype ZMUC ANT-000367, Red Sea (G; photo by M. A. Krag); micromorphology (H) and microstructure (I), hypotype RMNH 15106, southwest Sulawesi, Indonesia.
Figure 4. Astrea Lamarck, 1801 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 4. Astrea Lamarck, 1801, has discrete corallites that bud extracalicularly, medium-size (4–15 mm) and mediumrelief (3–6 mm) calices, septa in three cycles (24–36 septa), well-developed paliform (uniaxial) lobes, and spongy columellae. Septal teeth with low−medium height (≤ 0.6 mm) and medium spacing (0.3–1 mm). Walls formed by dominant paratheca, partial septotheca, and weak abortive septa. A, Astrea rotulosa (Ellis & Solander, 1786), type species of Astrea; macromorphology, holotype GLAHM 104014, unknown locality (photo by KG Johnson). B–F, Astrea curta Dana, 1846; micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype MTQ G61882, Orpheus Island, Australia; macromorphology (D), micromorphology (E), and microstructure (F), syntype USNM 14, Fiji. G–I, Astrea annuligera Milne Edwards & Haime, 1849b; macromorphology, holotype MNHN IK-2010-699, Australia (G); micromorphology (H) and microstructure (I), hypotype RMNH 10718, Heron Island, Australia.
Figure 3. Merulina Ehrenberg, 1834 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 3. Merulina Ehrenberg, 1834, has uniserial corallites with few centres, fused walls, small (<4 mm) and lowrelief (<3 mm) calices, septa in
Figure 7. Caulastraea Dana, 1846 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 7. Caulastraea Dana, 1846, is phaceloid, with discrete corallites, equally thick costosepta spaced
Figure 10 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 10. Dipsastraea de Blainville, 1830, has discrete corallites that bud intracalicularly, equally thick costosepta, and spongy columellae. Septal teeth with medium height (0.3–0.6 mm) and spacing (0.3–1 mm). Walls formed by dominant paratheca and partial septotheca, with transverse septal crosses. A–C, Dipsastraea favus (Forskål, 1775), type species of Dipsastraea; macromorphology, lectotype ZMUC ANT-000466, Red Sea (A); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype USNM 93662, Madang, Papua New Guinea. D, Dipsastraea amicorum (Milne Edwards & Haime, 1849b); macromorphology, holotype MNHN IK-2010-470, Tongatapu, Tonga (photo by P. Lozouet). E, F, Dipsastraea mirabilis (Yabe & Sugiyama, 1941); micromorphology (E) and microstructure (F), hypotype USNM 93642, Madang, Papua New Guinea. G–I, Dipsastraea laddi (Wells, 1954); macromorphology (G), micromorphology (H), and microstructure (I), holotype USNM 44942, Bikini Atoll, Marshall Islands. J–L, Dipsastraea pallida (Dana, 1846); macromorphology, syntype USNM 30, Fiji (J); micromorphology (K) and microstructure (L), hypotype USNM 44952, Bikini Atoll, Marshall Islands.
Figure 6 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 6. Boninastrea Yabe & Sugiyama, 1935, has uniserial corallites, fused walls, septa in <three cycles (<24 septa), unequally thick costosepta, rudimentary columellae, and abundant (vesicular) endotheca. A, B, Boninastrea boninensis Yabe & Sugiyama, 1935, the type and only living species of Boninastrea; macromorphology, holotype TIU 44970, Ogasawara Islands, Japan.
Figure 22 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 22. Pectinia de Blainville, 1825, has organically united corallites, extensive coenosteum (≥ corallite diameter), septa spaced
Figure 20. Paraclavarina Veron, 1985 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 20. Paraclavarina Veron, 1985: 179, is ramose and has uniserial corallites with few centres, fused walls, small (<4 mm) and low-relief (<3 mm) calices, septa in
Figure 21 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 21. Paramontastraea Huang & Budd, this study, has corallites that mostly bud extracalicularly, regular free septa, septa spaced> 11 septa per 5 mm, and well-developed paliform (uniaxial) lobes. Septal teeth are low (<0.3 mm) and narrowly spaced (<0.3 mm), with multiaxial tips. Walls formed by dominant septotheca and partial paratheca. A–C, Paramontastraea salebrosa (Nemenzo, 1959), type species of Paramontastraea; macromorphology, holotype UP C-192, Puerto Galera, the Philippines (A; photo by K. S. Luzon); micromorphology (scanning electron microscopy; B) and microstructure (transverse thin section; C), hypotype UP P1L02158), Batangas, the Philippines. D, E, Paramontastraea serageldini (Veron, 2000); macromorphology, holotype MTQ G55844, Mahé, Seychelles (D); micromorphology, hypotype SIO Co2805, Mahé, Seychelles (E). F, Paramontastraea salebrosa (Nemenzo, 1959); microstructure, hypotype UP P1L02158, Batangas, the Philippines.
Figure 19 in Taxonomic classification of the reef coral families Merulinidae, Montastraeidae, and Diploastraeidae (Cnidaria: Anthozoa: Scleractinia)
Figure 19. Oulophyllia Milne Edwards & Haime, 1848a, has fused walls, medium-size (4–15 mm) and medium-relief (3–6 mm) calices, septa spaced
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