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FIGURE 2 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 2. Alpheus microrhynchus De Man, 1897: syntype, incomplete male (cl indet., tl 40.0 mm) from Pontianak, Kalimantan, Indonesia (after De Man 1897, as Alpheus sp. and A. microrhynchus, specimen lost) [A–C]; female (cl indet., tl 43.0 mm) from Bangkok, Thailand (after De Man 1898) [D, E]; female (tl 85.0 mm) from Makassar, Sulawesi, Indonesia (after De Man 1911), specimen tentatively assigned to A. takla sp. nov. (see text) [F, G]: A—frontal region, dorsal view; B—male minor (left) chela, dorsal view; C—same, lateral view; D—female minor (right) cheliped, mesial view; E—female major (left) cheliped, lateral view; F—frontal area of carapace, dorsal view; G—telson, dorsal view. Not to scale.

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FIGURE 4 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 4. Alpheus euphrosyne De Man, 1897: chelipeds of male (cl indet.) from Sumatra, Indonesia (ZMA Crus.D.202726) showing main features and terminology used in the present study; A—major (right) cheliped, lateral view; B—same, mesial view; C—minor (left) cheliped, mesial view; D—same, lateral view. Abbreviations (in alphabetical order): ad—adhesive disks (plaques) on dactylus and palm (small on major chela, highly reduced in minor chela); be—balaeniceps expansion of minor chela dactylus; ddr—dorsal ridge of major chela dactylus (dactylar dorsal ridge); dla—dorsolateral angle of major chela pollex; dlt—dorsolateral tooth of major / minor chela palm; dma—dorsomesial angle of major chela pollex; dmt—distomesial tooth of major / minor chela palm; dp—plunger of major chela dactylus (dactylar plunger); ds—dorsal shoulder of major / minor chela palm; dtg—dorsal transverse groove of major / minor chela palm (also known as dorsal notch); lbd—lateral balaeniceps ridge of minor chela dactylus; lbp—lateral balaeniceps ridge of minor chela pollex; li—linea impressa of major / minor chela palm; lld—lateral longitudinal depression (groove) of major / minor chela palm; ltr—smooth lateral transverse ridge of major chela palm (often poorly developed); mbd—mesial balaeniceps ridge of minor chela dactylus; mbp—mesial balaeniceps ridge of minor chela pollex; mld—mesial longitudinal depression (groove) of major / minor chela palm; msr—mesial subdistal ridge on major chela pollex; mtr—smooth mesial transverse ridge of major chela palm (often poorly developed); pf—fossa (socket) of major chela pollex; vs—ventral shoulder of major / minor chela palm; vtg—ventral transverse groove of major / minor chela palm (also known as ventral notch).

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FIGURE 7 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 7. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) [A–G] and female (cl 18.0 mm) [H] from Angsila, Thailand (MNHN-IU-2018-5593); A—major (right) cheliped, chela, carpus and distal part of merus, mesial view; B—same, chela, lateral view; C—same, coxa to carpus, lateral view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela, carpus and distal part of merus, mesial view; F—same, chela, lateral view; G—same, coxa to carpus, lateral view; H—female minor chela, lateral view.

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FIGURE 5 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 5. Alpheus euphrosyne De Man, 1897: neotype, male (cl 16.6 mm) from Surabaya, Java, Indonesia (RMNH Crus. D.49669); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) chela in situ, mesial view; D—same, lateral view; E—minor (right) chela in situ, dorsal view. Refer to A for scale.

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FIGURE 19 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 19. Alpheus microrhynchus De Man, 1897: neotype, male (cl 16.3 mm) from Kabupaten Sambas, Kalimantan Barat, Indonesia (ZRC 2000.2193); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (right) cheliped, lateral view; D—same, mesial view; E—minor (left) cheliped, lateral view; F—same, mesial view. Refer to A for scale.

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FIGURE 13 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 13. Alpheus eurydactylus De Man, 1920: female (cl 21.3 mm) from Surabaya, Java, Indonesia (RMNH Crus.D.49664): A—habitus, lateral view; B—minor (left) chela in situ, mesial view; C—major (right) cheliped in situ, mesial view; D—same, lateral view. Refer to A for scale.

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FIGURE 25 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 25. Alpheus nomurai sp. nov.: holotype, male (cl 8.8 mm) from Tanabe, Wakayama, Japan (MNHN-IU-2018-5590); A—habitus, dorsal view; B—frontal region, dorsal view; C—major (left) cheliped, lateral view; D—same, mesial view; E— minor (right) cheliped, mesial view; F—same, dorsolateral view; G—same, lateral view. Refer to A for scale.

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FIGURE 12 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 12. Alpheus eurydactylus De Man, 1920: lectotype, male (cl 13.8 mm) [A–E] and paralectotype, ovigerous female (cl 15.0 mm) [F, G], both from Surabaya, Java, Indonesia (RMNH Crus.D.49670 = ZMA Crus.D.102536); A—major (left) cheliped, lateral view; B—same, chela, latreral view; C—same, fingers open, mesial view (see also Fig. 10C); D—minor chela in situ, dorsal view (cf. Fig. 3H); E—same, lateral view; F—frontal region, lateral view; G—frontal region and chelipeds in situ, dorsal view. Refer to Fig. 11A for scale in A–E.

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FIGURE 18. Alpheus richardsoni Yaldwyn, 1971 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 18. Alpheus richardsoni Yaldwyn, 1971: male (cl indet.) from Brunswisk River, New South Wales, Australia [A, B]; male from Parengarenga (Paua), New Zealand [C]; male from Whangarei, New Zealand [D]; A, C—habitus, dorsal view; B—same, lateral view; D—tail fan, dorsal view. Specimens not deposited / deposition unknown. Photographs courtesy of Denis Riek (A, B) and Richard B. Taylor (C, D).

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FIGURE 24 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 24. Alpheus cyanoteles Yeo & Ng, 1996: male (cl 9.5 mm) from Sungei Tementang, southern Johor, Malaysia (OUMNH.ZC. 2013.05.6); A—habitus, dorsal view; B—frontal region, dorsal view; C—tailfan, dorsal view; D—chelipeds (major chela widely opened), mesial view; E—H.H. Tan and S. De Grave collecting A. cyanoteles in its natural habitat, a small stream in southern Johor. Photographs by the author.

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FIGURE 29 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 29. Alpheus takla sp. nov.: holotype, male (cl 25.5 mm) from Bohol, Philippines (NMCR 99110) [A–G]; two paratypes, males (cl 24.0, 25.0 mm) from the same locality (MNHN-IU-2022-2939) [H–K]; A—habitus, dorsal view; B—same, lateral view; C—frontal region, dorsal view; D—major (left) cheliped, lateral view; E—same, mesial view; F—minor (right) chela, lateral view; G—same, mesial view; H—minor (left) cheliped, lateral view; I—same, chela and carpus, dorsal view; J—major (right) cheliped, mesial view; K—same, dorsal view. Refer to A, B, H, J for scale.

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FIGURE 28 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 28. Alpheus nomurai sp. nov.: male (cl not determined) from Yabu River, Nago, Okinawa-jima, Japan (not deposited) [A, B]; male paratype (cl 9.6 mm) from Tanabe, Wakayama, Japan (OUMNH.ZC. 2011.06-006) [C]; four specimens, including male holotype (cl 8.8 mm), from Tanabe, Wakayama, Japan (holotype: MNHN-IU-2018-5590) [D]; A—habitus, lateral view; B–D, same, dorsal view. Photographs courtesy of Tadafumi Maenosono (A, B) and Keiichi Nomura (C, D).

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FIGURE 23 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 23. Alpheus cyanoteles Yeo & Ng, 1996: ovigerous female (cl indet.) [A, B] and female (cl indet.) [C, D] from an unknown locality in southern Johor, Malaysia; A—habitus, lateral view; B—same, dorsal view; C—same, lateral view; D—detail of frontal margin of carapace and left antenna, lateral view. Aquarium-kept specimens, probably not deposited. Photographs courtesy of C.H. Wah (A, B) and Heok-Hui Tan (C, D).

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FIGURE 1 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 1. Alpheus euphrosyne De Man, 1897: syntype, female (cl 46.0 mm) from Java Sea, Indonesia (after De Man 1897, specimen lost) [A–E]; male (cl indet., tl 34.0 mm) from Bangkok, Thailand (after De Man 1898) [F, G]; A—frontal region, dorsal view; B—female minor (left) cheliped, lateral view; C—second pereiopod, lateral view; D—third pereiopod, lateral view; E—sixth pleonite, telson and uropods, dorsal view; F—male minor (right) cheliped, dorsolateral view; G—same, minor chela, lateral view. Not to scale.

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FIGURE 16 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 16. Alpheus eurydactylus De Man, 1920: male (cl indet.) from Can Gio, Vietnam (ZMMU) [A]; female (cl indet.) from the same locality (ZMMU) [B, C]; male (cl indet.) from near Bangkok, Thailand (deposition unknown) [D]: A—habitus, dorsal view; B, D—same, lateral view; C—major cheliped, mesial view. Photographs courtesy of Ivan N. Marin (A–C) and Sukkrit Nimitkul (D).

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FIGURE 3 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 3. Three species of Alpheus described by Kemp (1915), De Man (1920) and Schmitt (1926). Alpheus paludicola Kemp, 1915: syntype, male (cl indet., tl 22.0 mm) from Chilika Lake, India [A–G]; A—frontal region, dorsal view; B—scaphocerite of left antenna, dorsal view; C—(left) major cheliped, mesial view; D—same, lateral view; E—minor (right) cheliped, distal portion, mesial view; F—second pereiopod, lateral view; G—telson, dorsal view (after Kemp 1915). Alpheus eurydactylus De Man, 1920: syntypes, male (cl 13.8 mm) [H] and ovigerous female (cl 15.0 mm) [I] from Java, Indonesia (see text for lectotype / paralectotype designation); H—male minor (right) chela, distal portion of palm and fingers, dorsal view; I—female (right) minor chela, lateral view (after De Man 1924). Alpheus langi (Schmitt, 1926), originally described as Crangon langi, presenly junior synonym of Alpheus pontederiae de Rochebrune, 1883: holotype, male (cl 11.0 mm, tl 34.0 mm) from Congo [J–M]: J—frontal region, dorsal view; K—major (left) chela, lateral view; L—minor (right) chela, lateral view; M—second pereiopod, lateral view (after Schmitt 1926). Not to scale.

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FIGURE 6 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 6. Alpheus euphrosyne De Man, 1897: male (cl 16.4 mm) from Angsila, Thailand (MNHN-IU-2018-5593); A— frontal region, dorsal view; B—telson, dorsal view; C—ventromesial carina of first article of antennular peduncle, lateral view; D—antennal basicerite, lateral view; E—third maxilliped, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H, I—same, dactylus, mesial (H) and dorsal (I) views; J—fifth pereiopod, lateral view; K—same, propodus and dactylus, mesial view; L—second pleopod, appendices masculina and interna, anterior view; M—left uropod, dorsal view; N—right uropod, detail of distolateral angle.

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FIGURE 9 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 9. Alpheus euphrosyne De Man, 1897: male (cl 16.8 mm) from Pattani, Thailand (ZRC 2014.0672), in dorsal view (A), with details of the frontal area and chelipeds (B). Photographs courtesy of Sukree Hajisamae.

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FIGURE 15 in Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae)

FIGURE 15. Alpheus eurydactylus De Man, 1920: male (cl 16.4 mm) from Pattani, Thailand (ZRC 2014.0672) [A–G]; ovigerous female (cl indet.) from Johor, Malaysia (MNHN-IU-2018-5598) [H]; A—major (right) cheliped, chela, carpus and distal part of merus, lateral view; B—same, ischium to carpus, lateral view; C—same, chela and carpus, mesial view; D—same, dactylus, lateral view; E—male minor (left) cheliped, chela and carpus, mesial view; F—same, chela, lateral view; G—same, ischium to carpus, lateral view; H—female minor chela, lateral view.

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Highly variable (no clear pattern). All portions of the dorsal views were equally used. In head images the area around the eye, the top of the head, the snout and the throat were all used in similar proportions. P. carbonelli Variable for both views. Snout and middle of the dorsum used in dorsal view. Top of the head most frequently (but not strictly) used in lateral view. P. guadarramae Whole body used for dorsal view (but variable); either throat (most common) or ear region used in head lateral views. P. hispanicus Variable. Anterior portion of snout used more frequently than in other species for both dorsal and head lateral views. P. liolepis Highly variable. Whole body used in most dorsal images, area around the eye and throat used in head lateral views, but other patterns common. P. lusitanicus Highly variable. All parts of the dorsum used (but frequently the most posterior part); area around the ear frequently used in head lateral images. P. tunesiacus Highly variable. Dorsal area near the insertion of the posterior limbs used more frequently than in other species; different regions of the head used, often simultaneously. P. Ʋaucheri Highly variable. Different regions of dorsum (from head to the posterior region) used in dorsal images, all portions of the head, but most frequently the throat, used in lateral images. P. Ʋirescens Highly variable. All parts of both images used. Head and anterior part of the dorsum more used than in other species. in Identification of morphologically cryptic species with computer vision models: wall lizards (Squamata: Lacertidae: Podarcis) as a case study

Highly variable (no clear pattern). All portions of the dorsal views were equally used. In head images the area around the eye, the top of the head, the snout and the throat were all used in similar proportions. P. carbonelli Variable for both views. Snout and middle of the dorsum used in dorsal view. Top of the head most frequently (but not strictly) used in lateral view. P. guadarramae Whole body used for dorsal view (but variable); either throat (most common) or ear region used in head lateral views. P. hispanicus Variable. Anterior portion of snout used more frequently than in other species for both dorsal and head lateral views. P. liolepis Highly variable. Whole body used in most dorsal images, area around the eye and throat used in head lateral views, but other patterns common. P. lusitanicus Highly variable. All parts of the dorsum used (but frequently the most posterior part); area around the ear frequently used in head lateral images. P. tunesiacus Highly variable. Dorsal area near the insertion of the posterior limbs used more frequently than in other species; different regions of the head used, often simultaneously. P. Ʋaucheri Highly variable. Different regions of dorsum (from head to the posterior region) used in dorsal images, all portions of the head, but most frequently the throat, used in lateral images. P. Ʋirescens Highly variable. All parts of both images used. Head and anterior part of the dorsum more used than in other species.

opennotspecifiedApr 2023View details →

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Allen Brain Atlas

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Last verified 2026-04-30Open record

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electrophysiologyopenPublished Dandiset metadata and archive endpoints are available through the production DANDI API.
Last verified 2026-04-30Open record

International Brain Laboratory public data

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behavioral-neuroscienceopenPublic sessions can be searched and loaded from the IBL public data server through ONE.
Last verified 2026-04-29Open record

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neuroscienceopenPublished datasets are available on demand over the internet.
Last verified 2026-04-29Open record