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Figure 5. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 5. A, holotype of Hemiphyllodactyus tehtarik sp. nov. (LSUHC 10904) from Gunung Tebu, Terengganu, Malaysia; photo by L. Grismer. B, adult female Hemiphyllodactylus larutensis (LSUHC 11294) from Bukit Larut, Perak, Malaysia; photo by L. Grismer. C, adult male holotype of Hemiphyllodactylus larutensis (holotype BM 1901.3.20.2) from Bukit Larut, Perak, Malaysia; photo by G. Zug. D, adult male Hemiphylodactylus larutensis (LSUHC 11298) from Bukit Larut, Perak, Malaysia; photo by L. Grismer. E, adult male Hemiphyllodactylus harterti (LSUHC 10383) from Bukit Larut, Perak, Malaysia; photo by L. Grismer. F, adult female Hemiphyllodactylus harterti (holotype ZMB 1360) from Gunung Hijau, Perak, Peninsular Malaysia; photo by M.-O. Rödel.

Figure 7. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 7. A, adult male Hemiphyllodactylus sp. nov. 2 (LSUHC 5759) from Pulau Sibu, Johor, Malaysia; photo by L. Grismer. B, adult female Hemiphyllodactylus sp. nov. 3 (MVZ 239345) from Pulau Enggano, Sumatra, Indonesia; photo by J. McGuire.

Figure 8. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 8. A, adult male Hemiphyllodactylus sp. nov. 5 (PNM 7666) from Palaui Island; photo by R.M. Brown. B, adult male Hemiphyllodactylus insularis (KU uncatalogued) from Pasonanca, Zamboanga, Western Mindanao, Philippines. C, adult female Hemiphyllodactylus sp. nov. 4 (KU 331843) from Mount Lanoy, Municipality of Argao, Cebu Province, Philippines; photo by R.M. Brown.

Figure 2 in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 2. Distribution of the species of Hemiphyllodactylus examined (see the Appendix for a list of the Hemiphyllodactylus typus examined). Hemiphyllodactylus aurantiacus from Yercaud, Salem District (Shevaroyan Hill Range), Tamil Nadu, India; Hemiphyllodactylus ganoklonis from Palau Ngercheu; Hemiphyllodactylus harterti and Hemiphyllodactylus larutensis from Bukit Larut, Perak, Malaysia; Hemiphyllodactylus insularis from the Municipality of Pasonaca, Zamboanga City, Zamboanga City Province, Philippines; Hemiphyllodactylus tehtarik from Gunung Tebu, Terengganu, Malaysia; Hemiphyllodactylus titiwangsaensis from the Cameron Highlands, Pahang, Malaysia; Hemiphyllodactylus sp. nov. 1 from Fraser's Hill and the Genting Highlands, Pahang; Hemiphyllodactylus sp. nov. 2 from Pulau Sibu, Johor, Malaysia; Hemiphyllodactylus sp. nov. 3 from Pulau Enggano, Bengkulu Province, Sumatra, Indonesia; Hemiphyllodactylus sp. nov. 4 from Mount Lantoy, Municipality of Argao, Cebu Province, Cebu, Philippines; Hemiphyllodactylus sp. nov. 5 from Pulaui Island, Barangay Palaui, Municipality of Santa Ana, Cagayan Province, Philippines; Hemiphyllodactylus sp. nov. 6 from Barangay Kaim, Municipality of San Francisco, Agusan del Sur Province, Mindanao, Philippines; Hemiphyllodactylus sp. nov. 7 from Chiang Mai, Chiang Mai Province, Thailand; Hemiphyllodactylus sp. nov. 8 from Pyin-Oo-Lwin, Mandalay Division, Myanmar; Hemiphyllodactylus sp. nov. 9 from Pakxong District, Champasak Province, Laos; Hemiphyllodacylus zugi from Vinh Phu, Vietnam; and Hemiphyllodactylus sp. nov. 10 from Ba Na-Nui Chua Nature Reserve, Da Nang Province, Vietnam. Localities of Hemiphyllodactylus dushnanesis (closed yellow circles), Hemiphyllodactylus jinpingensis (closed blue circles), Hemiphyllodactylus longlingensis (closed orange circles), and Hemiphyllodactylus yunnanensis (closed green circles) follow Zhou et al. (1981), except for the southern specimen of H. dushanensis from Vinh Phuc, Vinh Phuc Province, Vietnam reported here.

Figure 1 in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 1. Maximum-likelihood phylogram (–lnL 22097.690183) of the genus Hemiphyllodactylus Bleeker, 1860 with Bayesian posterior probabilities and maximum-likelihood bootstrap values, respectively.

Figure 6. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 6. A, adult male Hemiphyllodactylus titiwangsaensis (LSUDPC 4506) from Cameron Highlands, Pahang, Peninsular Malaysia. B, adult female Hemiphyllodactylus sp. nov. 1 from Genting Highlands, Pahang, Peninsular Malaysia. C, adult male Hemiphyllodactylus sp. nov. 1 from Fraser's Hill, Pahang, Peninsular Malaysia.

Figure 10. A in Integrative taxonomy uncovers high levels of cryptic species diversity in Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) and the description of a new species from Peninsular Malaysia

Figure 10. A, adult Hemiphyllodactylus auriantiacus (LSUDPC 7006) from Yercaud, Salem District (Shevaroyan Hill Range), Tamil Nadu, India. B, adult Hemphyllodactylus sp. nov. 10 (ITBCZ 2450). C, male Hemphyllodactylus sp. nov. 10 (LSUDPC 6675) from Ba Na-Nui Chua Nature Reserve, Hoa Vang District, Da Nang City Province, Vietnam; photos B and C by Ngo Van Tri. D, adult male Hemiphyllodactylus zugi (LSUDPC 8089) from Cao Bang Province, Vietnam; Photo by T. Q. Nguyen and T. Lehmann.

FIGURE 21. Maximum parsimony 16S rRNA phylogram for the Boophis albipunctatus group. From 485 total characters, 391 were constant and 72 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 21. Maximum parsimony 16S rRNA phylogram for the Boophis albipunctatus group. From 485 total characters, 391 were constant and 72 parsimony informative. MP searches retained 26 trees of which a strict consensus is shown. Consensus support values higher than 50, from 2000 bootstrap replicates, are shown; an asterisk indicates Bayesian posterior probabilities equal or higher than 95%. Species newly described herein are in bold.

FIGURE 27 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 27. Spectrogram and waveform of call type 1 of Boophis luciae sp. nov. from Andasibe (recorded on 1 February 1995, air temperature 22°C).

FIGURE 19 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 19. (A) Male holotype of Boophis sandrae sp. nov. (ZMA 20133) from Ranomafana; (B) female paratype of Boophis sandrae sp. nov. (ZSM 236/2006) from Ambatolahy.

FIGURE 10 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 10. Male holotype of Boophis entingae sp. nov. (ZSM 2083/2007): (A) dorsolateral view; (B) ventral view; (C) detail showing colouration of posterior surfaces of thigh.

FIGURE 8 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 8. Male of Boophis brachychir (ZSM 2157/2007) from Forêt d'Ambre Special Reserve: (A) dorsolateral view; (B) ventral view.

FIGURE 5 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 5. Spectrograms and waveforms of advertisement calls of: (A) Boophis andrangoloaka from Ambohitantely Special Reserve (recorded on 18 January 2005, air temperature app. 18°C); (B) Boophis rhodoscelis from Ranomafanakely, Ranomafana National Park (recorded on 28 January 2004, air temperature app. 21°C).

FIGURE 9 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 9. Spectrogram, corresponding waveform and expanded waveform of the advertisement call of Boophis brachychir from Manongarivo Special Reserve (recorded on 1 February 2003, air temperature 24.5°C).

FIGURE 7. Maximum parsimony 16S in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 7. Maximum parsimony 16S rRNA phylogram of species of the Boophis goudoti group. From 510 total characters, 340 were constant and 130 parsimony informative. MP searches retained 3681 trees of which a strict consensus is shown. Consensus support values higher than 50, from 2000 bootstrap replicates, are shown; an asterisk indicates Bayesian posterior probabilities equal or higher than 95%. Species newly described or resurrected herein are in bold.

FIGURE 25 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 25. Comparative waveform sections (each 2000 ms duration) of advertisement calls of species in the Boophis albipunctatus group: (A) Boophis schuboeae (Ranomafana National Park, 23°C); (B) Boophis haingana sp. nov. (Andohahela National Park, low altitude, 23.2°C); (C) Boophis haingana sp. nov. (Andohahela National Park, 1600 m a.s.l., 17.6°C); (D) Boophis ankaratra (Manjakatompo, 18°C); (E) Boophis miadana sp. nov. (Andohahela National Park, 1550 m a.s.l., 17.6°C).

FIGURE 22 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 22. Male holotype of Boophis miadana sp. nov. (ZSM 5107/2005) from Andohahela, 1550 m a.s.l.: (A) dorsolateral view; (B) ventral view.

FIGURE 23 in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 23. Male holotype of Boophis haingana sp. nov. (ZSM 5109/2005) from Andohahela: (A) dorsolateral view; (B) ventral view.

FIGURE 1. Maximum parsimony 16S in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 1. Maximum parsimony 16S rRNA phylogram for species in the Boophis majori and B. microtympanum groups. From 480 total characters, 336 were constant and 129 parsimony informative. MP searches retained 39 trees of which a strict consensus is shown. Consensus support values higher than 50, from 2000 bootstrap replicates, are shown; an asterisk indicates Bayesian posterior probabilities equal or higher than 95%. Species newly described or resurrected herein are in bold.

FIGURE 18. Maximum parsimony 16S in Integrative taxonomy of Malagasy treefrogs: combination of molecular genetics, bioacoustics and comparative morphology reveals twelve additional species of Boophis 2383

FIGURE 18. Maximum parsimony 16S rRNA phylogram for species in the Boophis luteus group. From 470 total characters, 347 were constant and 111 parsimony informative. MP searches retained 60 trees of which a strict consensus is shown. Consensus support values higher than 50, from 2000 bootstrap replicates, are shown; an asterisk indicates Bayesian posterior probabilities equal or higher than 95%. The species newly described herein is in bold.