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Figure 4 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 4 Labial cartilages. A, detail of labial cartilages in Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, detail of labial cartilages in Asymbolus rubiginosus, AMS I.30393-004, male, 527 mm TL. ucl, upper labial cartilage; lcl, lower labial cartilage.
Figure 30 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 30 Strict consensus cladogram of the three equally most-parsimonious trees (L = 233, CI = 0.37, RI = 0.69). Number of clades shown above branches. Values of relative Bremer support shown below branches.
Figure 3 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 3 Nasoral region with lifted anterior nasal flap and exposed posterior nasal flap. A, Scyliorhinus canicula, USNM 221470, female, 438 mm TL; B, Atelomycterus fasciatus, CSIRO H1298-7, male, 370 mm TL; C, Haploblepharus edwardsii, AMNH 40988, male, 480 mm TL. anf, anterior nasal flap; ful, flap on the upper lip margin; llf, lower labial furrow; ng, nasoral groove; pnf, posterior nasal flap; pog, postoral groove; ulf, upper labial furrow; umf, upper mesonarial flap. Scale bar: 20 mm.
Figure 5 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 5 Pelvic apron. A, detail of pelvic apron in Scyliorhinus comoroensis, MNHN 1984–0701, male, 457.2 mm TL; B, detail of pelvic apron in S. capensis, SAIAB 27577, male, 863 mm TL. imp, inner margins of pelvic fins; pa, pelvic apron. Modified from Soares and de Carvalho (2019).
Figure 29 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 29 Color patterns. A, Scyliorhinus canicula, MNHN 1999–1732, female, 418.5 mm TL; B, Scyliorhinus retifer, UF 36359, male, 372 mm TL; C, Scyliorhinus torazame, NSMT 50632, male, 427.9 mm TL; D, Scyliorhinus torrei, USNM 157852, male, 285 mm TL; E, Atelomycterus fasciatus, CSIRO H1298-7, male, 370 mm TL; F, Parmaturus angelae, MZUSP 124001, female, 425 mm TL; G, Poroderma pantherinum, SAIAB 34577, male, 640 mm TL; H, Poroderma africanum, SAIAB 25343, male, 920 mm TL. Scale bar: 20 mm.
Figure 28 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 28 Detail of the clasper siphon. A, Apristurus longicephalus, HUMZ 170382, 475 mm TL; B, Holohalaelurus regani, SAIAB 25717, 610 mm TL; C, Poroderma africanum, SAIAB 25343, 920 mm TL; D, Asymbolus rubiginosus, AMS I.30393-004, 527 mm TL; E, Halaelurus natalensis, SAIAB 26951, 400 mm TL; F, Haploblepharus edwardsii, AMNH 40988, 480 mm TL. Scale bar: 20 mm.
Figure 9 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 9 Detail of the postorbital musculature. A, Schroederichthys maculatus, UF 65846, male, 313 mm TL; B, Poroderma africanum, AMNH 43134, male, 505 mm TL. mdp, muscle depressor palpebrae nictitantis; mlp, m. levator palpebrae nictitantis; mrp, m. retractor palpebrae nictitantis; msp, m. espiracularis, ob, orbit; sp, spiracle.
Figure 31 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 31 Phylogenetic hypotheses of the subfamily Scyliorhininae. A, morphology- based hypothesis (present work; Compagno, 1988a); B, molecular-based hypothesis (Naylor et al., 2012a, 2012b).
Figure 27 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 27 Clasper; skeleton. A, Scyliorhinus boa, USNM 221563, 348 mm TL; B, Scyliorhinus canicula, BMNH 1983.8.3.1–4, 585.7 mm TL; C, Scyliorhinus capensis, SAIAB 27577, 863 mm TL; D, Scyliorhinus retifer, UF 36359, 372 mm TL; E, Scyliorhinus stellaris, BMNH 1976.7.30.10, 476 mm TL; F, Scyliorhinus torazame, NSMT 50632, 427.9 mm TL. end, endstyle; rd, dorsal marginal cartilage; rv, ventral marginal cartilage; t3, accessory terminal cartilage; td, dorsal terminal cartilage; td2, dorsal terminal 2 cartilage; tv, ventral terminal cartilage; tv2, ventral terminal 2 cartilage. Modified from Soares and de Carvalho (2019).
Figure 23 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 23 Detail of the gill pickax. A, Scyliorhinus haeckelii, UERJ 1691, male, 522 mm TL; B, Cephaloscyllium sufflans, SAIAB 6242, male, 800 mm TL.
Figure 22 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 22 Detail of the oropharyngeal denticles in Cephaloscyllium sufflans, SAIAB 6242, male, 800 mm TL. cb I–V, ceratobranchials I–V.
Figure 19 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 19 Detail of the basihyal cartilage. A, Scyliorhinus haeckelii, UERJ 1691, male, 522 mm TL; B, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL. thf, thyroid foramen.
Figure 21 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 21 Detail of the lateral processi rastriformis (ras) in Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL.
Figure 17 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 17 Detail of jaws; lighter portion of Meckel's cartilage is less calcified portion. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. lrq, labial ridge of the quadrate process; mck, Meckel's cartilage; opq, orbital process of the palatoquadrate; pq, palatoquadrate.
Figure 2 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 2 Nasoral region with lifted anterior nasal flap and exposed posterior nasal flap. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Cephaloscyllium isabella, USNM 320594, female, 390 mm TL; Poroderma pantherinum, SAIAB 34577, male, 640 mm TL. anf, anterior nasal flap; ful, flap on the upper lip margin; llf, lower labial furrow; mrd, mesonarial crest; nb, nasal barbel; pnf, posterior nasal flap; pog, postoral groove; ulf, upper labial furrow. Scale bar: 20 mm.
Figure 16 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 16 Neurocranium; lateral view. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Figaro boardmani, CSIRO H989-5, female, 465 mm TL; C, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. enc, external nasal cartilage; hf, hyomandibular facet; lrc, lateral rostral cartilage; mrc, medial rostral cartilage; nc, nasal capsule; pog, postorbital groove.
Figure 18 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 18 Detail of the articular region of the quadratomandibular joint of Meckel's cartilage. A, Cephaloscyllium sufflans, SAIAB 6242, male, 800 mm TL; Galeus antillensis, UF 77853, female, 370 mm TL; C, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. lac, labial condyle; lic, lingual condyle; mfa, mandibular facet.
Figure 15 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 15 Neurocranium; ventral view. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Cephaloscyllium variegatum, AMS I.43762-001, female, 670 mm TL; C, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL; D, Atelomycterus fasciatus, CSIRO H1298-7, male, 370 mm TL; E, Asymbolus rubiginosus, AMS I.30393-004, male, 527 mm TL; F, Figaro boardmani, CSIRO H989-5, female, 465 mm TL; G, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. bp, basal plate; enc, external nasal cartilage; exc, excurrent aperture; hf, hyomandibular facet; icf, internal carotid foramen; inc, incurrent aperture; lrc, lateral rostral cartilage; mrc, medial rostral cartilage; nf, nasal fontanelle; sbp, subnasal plate; sf, stapedial foramen; ss, suborbital shelf.
Figure 24 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 24 Coracoid bar; ventral view. A, Scyliorhinus haeckelii, UERJ 1691, male, 522 mm TL; B, Halaelurus natalensis, SAIAB 26951, male, 400 mm TL; C, Holohalaelurus regani, SAIAB 25717, male, 610 mm TL; D, Cephalurus cephalus, USNM 221527, female, 285 mm TL. mpc, medial projection of the coracoid bar; lpc, lateral processes on pectoral girdle.
Figure 8 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420
Figure 8 Detail of the caudal crest in Galeus antillensis, UF 77853, female, 370 mm TL. A, general view, B, closed view. Scale bar:10 mm (A); 200 µm (B).
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