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FIGURE 13 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 13. Hungerfordia triplochilus sp. nov. (A1–A5) A topotypic paratype, B0158a; (B) columellar tooth (left lateral view), topotypic paratype, B0158b; (C) axial ribs above the aperture, a subadult of topotypic paratype, B0158c; (D1) sculpture on the last 1–1.5 whorls, and (D2) sculpture on the last 1.5–2 whorls of topotypic paratypes (B0158a, d); (E1–E4) holotype, B0158T; (F) apical septum, a partially opened paratype from Loc. Gk1-2, B0145; (G) sculpture behind the aperture, holotype, B0158T; (H) a worn paratype from Loc. Gk7-7, B1025; (I) a paratype from Loc. Gk7-5, B1000; (J1, J2) outer surface, and (J3, J4) inner surface of operculum, a topotypic paratype, B0158. Scale bars, 1mm (A–E, G–I), 0.5 mm (F, J). All specimens, UMUTZ-MG.
FIGURE 12. Hungerfordia pelewensis Beddome, 1889 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 12. Hungerfordia pelewensis Beddome, 1889. (A1–A5) An adult from Loc. Ko2-2, B0046a; (B) an adult from Loc. Ko5-6, B0542; (C1–C3) an adult from Loc. Gk5-5, B0651a; (D1–D3) a juvenile, and (E1–E3) a subadult from Loc. Ub6-1 (B0752a, b); (E4) enlarged Fig. E2, showing an abrupt change of the sculpture and whorl shape on the last whorl; (F) subapical views of broken adults showing the columellar tooth and the constriction: (F1) an adult from Loc. Gk5-5, B0651b; (F2) an adult from Loc. Ko2-2, B0046b; (G) a young subadult from Loc. Gk5-5, B0752c; (H) sculpture on penultimate and the higher two whorls, a subadult from Loc. Ub6-1, B0752; (I) protoconch, an adult from Loc. Ko2-2, B0651c; (J) an adult from Loc. BA4-1, B0283; (K) an adult from Loc. Gk5-5, B0046c; (L1, L2) outer surface, and (L3, L4) inner surface of operculum, Loc. Ko2-2, B0046. Arrow heads show position of the constriction. Scale bars, 1mm (A–G, J, K), 0.5 mm (H, I, L). All specimens, UMUTZ-MG.
FIGURE 10 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 10. Opercula of Palaina (Eupalaina) and Hungerfordia species. (A) P. (E.) moussoni Crosse, 1866, B0229; (B) P. (E.) dimorpha Crosse, 1866, B0087; (C) P. (E.) sp.-18, longitudinal section, B0082; (D) P. (E.) patula Crosse, 1866, B0204; (E) H. polymorpha (Crosse, 1866), B0049; (F) H. pelewensis Beddome, 1889, B0046; (G) H. polymorpha (Crosse, 1866), longitudinal section, B0049; (H) H. alata (Crosse, 1866), B0058. Oblique view of the outer surface, oblique view of the inner surface, and vertical view of the inner surface are shown from left to right in Figs. A, B, E, and F. Oblique view of the inner surface and vertical view of the inner surface are shown from left to right in Figs. D and H. Scale bars, 0.5 mm (A–B, D, E–F, H), 0.1 mm (C, G). All specimens, UMUTZ-MG.
FIGURE 8 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 8. Radulae and buccal masses of Hungerfordia and Palaina (Eupalaina) species. (A1–A4) H. pelewensis Beddome, 1889, B0283; (B1–B4) H. polymorpha (Crosse, 1866), B0315; (C1–C4) P. (E.) sp.-20, B0595; (D1, D2) P. (E.) patula Crosse, 1866, B0320; (E1) H. pelewensis Beddome, 1889, buccal mass, B0283; (E2) H. polymorpha (Crosse, 1866), buccal mass, B0315; (E3) P. (E.) patula Crosse, 1866, buccal mass, B0320. Scale bars, 10 µm (A–D), 0.5 mm (E1–E3). All specimens, UMUTZ-MG.
FIGURE 9 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 9. Radulae of Hungerfordia species. (A) H. pelewensis Beddome, 1889, B0283; (B) H. triplochilus sp. nov., topotypic paratype, B0158; (C) H. expansilabris sp. nov., topotypic paratype, B1299; (D) H. nudicollum sp. nov., topotypic paratype, B0364; (E) H. echinata echinata sp. et subsp. nov., paratype, B1084; (F) H. echinata tubulispina subsp. nov., topotypic paratype, B0169; (G) H. elegantissima sp. nov., topotypic paratype, B0375; (H) H. goniobasis goniobasis sp. et subsp. nov., topotypic paratype, B0235; (I) H. goniobasis dmasechensis subsp. nov., topotypic paratype, B0579; (J) H. lamellata (Crosse, 1866), B0157; (K) H. subalata sp. nov., paratype, B0021; (L) H. pteropurpuroides sp. nov., topotypic paratype, B1116; (M) H. alata (Crosse, 1866), B1041; (N) H. papilio papilio sp. et subsp. nov., paratype, B0374; (O) H. papilio stenoptera subsp. nov., paratype, B0761. Scale bars, 10 µm. All specimens, UMUTZ-MG.
FIGURE 6 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 6. Terminology used in this study. (A) Shell dimensions. Abbreviations: D, shell diameter; H, shell height; PH, peristome height; SW, suture width. (B) An example showing the axial protrusions of shell wall along the axial ribs. Umbilical view of a shell of Hungerfordia pyramis (Crosse, 1866), only the last whorl shown. (C) Apical septa of some Hungerfordia species: (C1) apical whorls of a shell (live condition) of H. pteropurpuroides sp. nov.; (C2) apical whorls of a partially opened shell of H. alata (Crosse, 1866), which shows construction of an internal septum; (C3) subapical view of a shell of H. triplochilus sp. nov. after removal of the empty apical whorls above the septum.
FIGURE 7 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 7. Male soft parts of Palaina (Eupalaina) and Hungerfordia species. (A1, A2) P. (E.) patula Crosse, 1866, B0753; (B) P. (E.) sp.-20, live condition, B0732; (C) H. inflatula (Crosse, 1866), B0217; (D) H. pelewensis Beddome, 1889, live condition, B0651; (E) H. polymorpha (Crosse, 1866), live condition, B0659. Arrow heads indicate penes. Scale bars, 1mm. All specimens, UMUTZ-MG.
FIGURES 4–5 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURES 4–5. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 4), map of Ngerkutabel island and its neighboring islands or islets; bottom (Fig. 5), map of Mecherchar island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2.
FIGURE 1 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 1. Map of Palau islands showing collecting sites. Small red circles show collecting sites. Areas encircled by blue rectangles are shown as enlarged maps in Figs. 2–5. Abbreviations of the collecting sites are listed in Table 2.
FIGURE 26 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 26. Hungerfordia alata (Crosse, 1866). (A1–A4) Syntype of "Palaina" alata, MNHN 26601; (B1, B2) original labels of the syntype; (C) a fresh specimen with a narrow interspace between the inner and outer peristomes, Loc. Ch5-1, UMUTZ- MG-B0523b. Scale bar, 1mm.
FIGURE 25 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 25. Hungerfordia alata (Crosse, 1866). (A1–A4) Loc. Ko2-2, B1396a; (B) columellar tooth (dorsal view), Loc. Ko2- 2, B1396b; (C) axial ribs above the aperture (after removal of the inner peristome), Loc. Ko2-2, B1396a; (D) sculpture on the penultimate whorl, Loc. Ko2-2, B1396c; (E) sculpture above the suture of the last whorl, Loc. Ch8-2, B1396; (F1–F3) Loc. Ko2-2, B1396d; (G) apical septum, a partially opened specimen, Loc. Ko2-2, B1396; (H) umbilical view of a specimen after removal of the winged ribs, only the last whorl shown, Loc. Ko2-2, B1396e; (I1, I2) Loc. BA5-2, B0446; (J) Loc. Ch5-1, B0523a; (K) a reddish specimen from Loc. Ub7-2, B1062; (L1, L2) outer surface, and (L3, L4) inner surface of operculum, Loc. Ko2-2, B1396. Scale bars, 1mm (A–K), 0.5 mm (L). All specimens, UMUTZ-MG.
FIGURE 24 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 24. Hungerfordia pteropurpuroides sp. nov. (A1–A4) A topotypic paratype, B0281a; (B) columella (left lateral view), (C) axial ribs above the aperture, (D) apical septum, and (E) sculpture on the penultimate whorl of topotypic paratypes (B0281a–d); (F1) left lateral view, and (F2) oblique apical view of sculpture on the penultimate whorl of the holotype; (G) umbilical view of a topotypic paratype after removal of the winged ribs, only the last whorl shown, B0281e; (H) oblique umbilical view of a paratype from Loc. BA8-3, a specimen with a narrowly opened umbilicus, B1311; (I1–I4) holotype, B0281T; (J) a paratype from Loc. BA5-2, B0447; (K) a paratype from Loc. BA6-3, B0684; (L) a paratype from Loc. BA4-2, B0295; (M1, M2) outer surface, and (M3, M4) inner surface of operculum, a topotypic paratype, B0281. Scale bars, 1mm (A–L), 0.5 mm (M). All specimens, UMUTZ-MG.
FIGURE 23 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 23. Hungerfordia subalata sp. nov. (A1–A4) A topotypic paratype, B0385a; (B) columellar tooth (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the penultimate whorl of topotypic paratypes (B0385a, b); (E) sculpture above the suture of the last whorl, holotype, B0385T; (F1–F4) holotype, B0385T; (G1–G3) a paratype from Loc. Gk6-2, untypical morph with developed spiny ribs above the aperture and with a narrowly opened umbilicus, B0738; (H) a paratype from Loc. Gk5-5, B0655; (I) a paratype from Loc. Gk4-4, B0425; (J1, J2) outer surface, and (J3, J4) inner surface of operculum, a topotypic paratype, B0385c. Scale bars, 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG.
FIGURE 22 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 22. Hungerfordia lamellata (Crosse, 1866). (A1–A6) Syntype of "Palaina" lamellata, MNHN 26604; (A7) sculpture on apical whorls of the syntype; (B1, B2) original labels of the syntype; (C) a fresh specimen with circum-apertural sculpture similar to the syntype, Loc. Ko5-6, UMUTZ-MG-B0541. Arrow head shows an abrupt whorl swelling after the constriction. Scale bars, 1mm.
FIGURE 21 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 21. Hungerfordia lamellata (Crosse, 1866). (A1–A4) Loc. Ub3-1, B0199a; (B) sculpture on penultimate whorl, Loc. Ub3-1, B0199a; (C) axial ribs above the aperture, Loc. Ko8-2, B1419; (D1) inner surface of constriction (apical view), Loc. Ub3-1, B0199b; (D2) inner surface of constriction (apical view), Loc. BA4-1, B0282a; (E) sculpture on apical whorls, Loc. BA4-1, B0282b; (F1–F4) Loc. Ub3-1, B0199; (G1–G3) Loc. BA4-1, B0282b–c; (H) Loc. Ko2-2, B0059a; (I, J) Loc. Ko5-6, B0541; (K1) columella (dorsal view), Loc. Ub3-1, B0199c; (K2) columellar tooth (dorsal view), Loc. Ko2-2, B0059b; (L1, L2) outer surface, and (L3, L4) inner surface of operculum, Loc. Ub3-1, B0199. Arrow head shows position of the constriction. Scale bars, 1mm (A–C, F–K), 0.5 mm (D–E, L). All specimens, UMUTZ-MG.
FIGURE 20 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 20. Hungerfordia goniobasis dmasechensis subsp. nov. (A1–A5) A topotypic paratype, B0579a; (B) aperture, (C) columellar tooth (dorsal view), (D) axial ribs above the aperture, and (E) sculpture on the penultimate whorl (left lateral view) of topotypic paratypes (B0579a, b); (F) sculpture on the penultimate whorl of the holotype; (G1–G5) holotype, B0579T; (H) a topotypic paratype, B0579c; (I1, I2) outer surface, and (I3, I4) inner surface of operculum, a topotypic paratype, B0579. Scale bars 1mm (A, G–H), 0.5 mm (B–F, I). All specimens, UMUTZ-MG.
FIGURE 19 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 19. Hungerfordia goniobasis goniobasis sp. et subsp. nov. (A1–A5) A topotypic paratype, B0235a; (B) aperture, (C) columella (dorsal view), (D) axial ribs above the aperture (umbilical view), and (E) sculpture on the penultimate whorl of a topotypic paratype (B0235a); (F1–F5) holotype, B0235T; (G) a topotypic paratype, B0235b; (H1, H2) a paratype with short spiny ribs from Loc. Mc6-1, B0763; (I1, I2) outer surface, and (I3, I4) inner surface of operculum, a topotypic paratype, B0235. Scale bars, 1mm (A, D–H), 0.5 mm (B, C, I). All specimens, UMUTZ-MG.
FIGURE 17 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 17. Hungerfordia echinata tubulispina subsp. nov. (A1–A4) A topotypic paratype, B0169a; (B) columella (dorsal view), (C) axial ribs above the aperture (after removal of parietal margin of the outer peristome), (D1) sculpture on the last 1.25–1.5 whorls, and (D2) sculpture on the 1.5–1.25 whorls of topotypic paratypes (B0169a, b); (E1–E4) holotype, B0169T; (F) umbilical view of last whorl, and (G) front view of entire shell of a topotypic paratype after removal of the spiny ribs (B0169c); (H, I) topotypic paratypes, B0169d, e; (J1, J2) outer surface, and (J3, J4) inner surface of operculum, a topotypic paratype, B0169. Scale bars, 1mm (A–K), 0.5 mm (J). All specimens, UMUTZ-MG.
FIGURE 16 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 16. Hungerfordia echinata echinata sp. et subsp. nov. (A1–A4) A paratype from Loc. BA5-1, B0437a; (B) columellar tooth (left lateral view), and (C) axial ribs above the aperture of paratypes from Loc. BA5-1 (B0437a, b); (D1) sculpture on the last 2–1.5 whorls, and (D2) sculpture on the last 1–1.5 whorls of a paratype from Loc. BA5-1, B0437a; (E1–E4) holotype, B1441T; (F) umbilical view of a paratype after removal of the spiny ribs, only the last whorl shown, Loc. BA7-1, B1084b; (G) a paratype from Loc. BA6-1, B0662; (H) a paratype with broken spines from Loc. BA5-1, B0437c; (I) a paratype from Loc. BA8-2, B1300; (J) apical septum, a partially opened paratype from Loc. BA5-1, B0437b; (K1, K2) outer surface, and (K3, K4) inner surface of operculum, a paratype from Loc. BA5-1, B0437. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG.
FIGURE 15 in <p><strong>Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em>Hungerfordia</em> </strong><br /><strong>species with highly developed axial ribs</strong></p>
FIGURE 15. Hungerfordia nudicollum sp. nov. (A1–A4) A paratype from Loc. Gk4-2, B0364a; (B) columellar tooth (dorsal view), a paratype from Loc. Gk4-2, B0364b; (C) axial ribs above the aperture, a subadult of topotypic paratype, B0146a; (D1) sculpture on the last 1–1.5 whorls, and (D2) sculpture on the last 1.5–2 whorls of a paratype from Loc. Gk4-2 (B0364a); (E) sculpture behind the aperture, a topotypic paratype, B0364c; (F) a paratype from Loc. Gk7-2, B0912; (G1–G4) holotype, B0364T; (H) a topotypic paratype, B0146b; (I) a paratype after removal of the spiny ribs, Loc. Gk4-2, B0364d; (J1, J2) outer surface, and (J3, J4) inner surface of operculum, a topotypic paratype, B0364. Scale bars, 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG.
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