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Figure 7 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 7 Female reproductive tract of Theprisa spp., ventral view AT. otwayBT. convexaCT. montanaDT. australis. Abbreviations: bc, bursa copulatrix; co, common oviduct; dgd, defensive gland efferent duct; gc, gonocoxa; hg, hindgut; hs, helminthoid sclerite; sd, spermathecal duct; sg, spermathecal gland; sgd, spermathecal gland duct; sgs, spermathecal gland stem; sp, spermatheca; v, vagina. Scale bars: 0.50 mm.

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Figure 8 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 8 Left gonocoxa, ventral view of Theprisa spp., ventral view AT. otwayBT. convexaCT. montanaDT. australisET. darlingtoni. Abbreviations: afs, apical fringe seta(e) of gonocoxite 1; ans, apical nematiform setae; des, dorsal ensiform seta; gc1, basal gonocoxite 1; gc2, apical gonocoxite 2; les, lateral ensiform setae; r, ramus.

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Figure 5 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 5 Parameres of male aedeagus for Theprisa spp., outside lateral view. Right paramere shown above left paramere AT. otwayBT. convexaCT. montanaDT. australisET. darlingtoni. Abbreviations: lp, left paramere; rp, right paramere.

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Figure 4 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 4 Male aedeagus internal sac in everted condition for Theprisa spp. AT. otway, right view BT. convexaCT. montanaDT. australisET. darlingtoni. Abbreviations: fl, flagellum; mf, microtrichial field; rp, right paramere. Scale bars: 0.5 mm

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Figure 3 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 3 Male aedeagus including parameres of Theprisa spp., right, or anatomically ventral view AT. otwayBT. convexaCT. montanaDT. australisET. darlingtoni. Abbreviations: ad, apical divot; af, apical face; fl, flagellum; lp, left paramere; mf, microtrichial field; rp, right paramere.

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Figure 6 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 6 Structures associated with abdominal segments VIII and IX for Theprisa spp., dorsal view AT. otwayBT. convexaCT. montanaDT. australisET. darlingtoni. Abbreviations: VIII, antecostal apodeme of abdominal segment VIII; IX, antecostal apodeme of abdominal segment IX; sp, spiracle of segment VIII; t, tergite of segment IX. Scale bars: 0.5 mm.

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Figure 2 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 2 Dorsal habitus illustrations of Theprisa spp.; range of standardized body lengths indicated AT. australis male BT. darlingtoni, female.

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Figure 11 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 11 Female reproductive tract of Theprisa darlingtoni, ventral view; scale bar = 0.50 mm. Abbreviations: bc, bursa copulatrix; co, common oviduct; gc, gonocoxa; hg, hindgut; hs, helminthoid sclerite; sd, spermathecal duct; sg, spermathecal gland; sgd, spermathecal gland duct; sgs, spermathecal gland stem; sp, spermatheca; v, vagina.

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Figure 12 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 12 Strict consensus of two equally parsimonious, 380-step cladograms including Theprisa spp. and cladistically neighboring moriomorphine taxa of the subtribe Tropopterina (see text); consensus cladogram length 382 steps, CI = 0.43, RI = 0.66. Character numbers are shown above cladogram edges, character states below. Cladogram root placed so tree topology is compatible with the more inclusive cladogram of Liebherr (2020: fig. 1). Filled squares represent unique state transformations on cladogram, open squares indicate state transformations that occur more than once on cladogram. Clades of this analysis previously recovered in Liebherr (2020) are indicated by pointers numbered 1; the clade subtended by Pterogmus reported in Liebherr (2019) indicated by the pointer numbered 2. Decay indices (i.e., Bremer Support values) shown beneath cladogram edges for all internal edges of the cladogram. Geographic distributions of species are indicated by abbreviations following species epithets: EOZ, southeastern Australia; i.e., A. C. T., New South Wales and Victoria; NNZ, North Island, New Zealand; NZ, New Zealand; SNZ, South Island, New Zealand; QOZ, Queensland; Tas, Tasmania; WOZ, Western Australia

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Figure 10 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 10 Distributional records for Theprisa spp.: T. australis (circle);  T. darlingtoni (square).

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Figure 1 from: Liebherr JK, Porch N, Shaw M, Sinclair BE, Maddison DR (2021) Systematic revision of the trans-Bassian moriomorphine genus Theprisa Moore (Coleoptera, Carabidae). In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 339-373. https://doi.org/10.3897/zookeys.1044.62335

Figure 1 Dorsal habitus illustrations of Theprisa spp.; range of standardized body lengths indicated AT. otway female BT. convexa female CT. montana female.

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Supplementary material 1 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Tables S1–S3

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Figure 4 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figure 4 Maximum likelihood tree for concatenated matrix of all genes. Scale bar: 0.1 units, as estimated by RAXML.

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Chart 1 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Chart 1 Support for or against various clades. All columns provide maximum likelihood bootstrap values for or against a particular clade, except for column "8G B," which shows the Bayesian posterior probability estimates for the eight-gene matrix. "8GML" shows the bootstrap values for the eight-gene concatenated matrix, "Nuc G" for the concatenated nuclear genes, "NPC G" for the concatenated nuclear protein-coding genes, and "Mito G" for the concatenated mitochondrial genes. The remaining eight columns provide values for the single gene analyses. All values are expressed as percentages, with positive numbers indicating support for a clade and negative numbers indicating support for a contradictory clade having the highest support. Specific contradictory clades from alternative trees are highlighted in medium grey. Cells with bootstrap values ≥ 90 are shown in black, with values between 75 and 89 in dark grey, and values from 50 to 74 in light grey. Cells in white indicate clades present in the ML tree, but with bootstrap values < 50. Cells in red have bootstrap values for a contradictory clade ≥ 50. Cells in pink have bootstrap values for or against a clade < 50, and the clade is not present in the ML tree. A "-" in a cell indicates that taxon sampling for that gene was not sufficient to assess monophyly of that clade. "#g" shows the number of single-gene analyses (maximum of eight) that support a clade with bootstrap values of 50 or more.

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Supplementary material 2 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figures S1–S13

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Figure 3 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figure 3 Habitus images of NebriaAN. (Eonebria) djakonovi Semenov & Znojko BN. (Orientonebria) coreica Solsky CN. (Spelaeonebria) nudicollis Peyerimhoff DN. (Psilonebria) superna Andrewes EN. (Reductonebria) ochotica Sahlberg FN. (Catonebria) banksii Crotch. Scale bars: 1.0 mm. Photograph credits: A, B, F Kiril Makarov; C, D David Maddison; E Alexander Anischenko.

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Figure 2 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figure 2 Habitus images of NebriiniALeistus (Nebrileistus) nubivagus Wollaston BL. (Leistus) ferruginosus Mannerheim CArchastes solitarius (Ledoux & Roux) DNippononebria (Vancouveria) virescens (Horn) ENebria (Oreonebria) castanea Bonelli FN. (Eurynebria) complanata (Linnaeus). Scale bars: 1.0 mm. Photograph credits: A, D–F David Maddison; B, C Alexander Anischenko.

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Figure 6 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figure 6 Summary tree of nebriite phylogeny illustrating the revised classification; clade representation in Europe (including North Africa and the Middle East), Asia, and North America is indicated in the three-box bar.

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Figure 5 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figure 5 Majority rule consensus tree of trees from bootstrap replicates. The first number under a branch is the percentage of bootstrap replicates with that clade, the second number is the estimate of the Bayesian posterior probability of that clade expressed as a percentage.

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Figure 1 from: Kavanaugh DH, Maddison DR, Simison WB, Schoville SD, Schmidt J, Faille A, Moore W, Pflug JM, Archambeault SL, Hoang T, Chen J-Y (2021) Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data. In: Spence J, Casale A, Assmann T, Liebherr JК, Penev L (Eds) Systematic Zoology and Biodiversity Science: A tribute to Terry Erwin (1940-2020). ZooKeys 1044: 41-152. https://doi.org/10.3897/zookeys.1044.62245

Figure 1 Habitus images of NebriitaeANotiokasis chaudoiri Kavanaugh & Nègre BPelophila borealis (Paykull) COpisthius richardsoni Kirby DParopisthius indicus chinensis Bousquet & Smetana ENotiophilus palustris Duftschmid FArchileistobrius hwangtienyuni Shilenkov & Kryzhanovskij. Scale bars: 1.0 mm. Photograph credits: A, C David Maddison; B, E Kiril Makarov; D, F Alexander Anischenko.

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