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FIGURE 12 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 12. Males of Dicharax species. A–B. Dicharax (?) alticola Páll-Gergely & Hunyadi n. sp., paratype from the type locality, coll. JUO. C–D. Dicharax cristatus (Möllendorff, 1886), 20130911B, coll. JUO. E–F. Dicharax depressus (Bavay & Dautzenberg, 1912), 20080510B, coll. JUO. G–H. Dicharax (?) immaculatus Páll-Gergely n. sp. 20110420E (2 different specimens), coll. JUO. Figures not to scale.
FIGURE 18 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 18. Dicharax (?) moellendorffi (Kobelt & Möllendorff, 1886). A. lectotype (SMF 39236). B–D. surface of R2 (NHMW 40497). E. cross section of R2 (NHMW 40497). All images: B. Páll-Gergely.
FIGURE 15 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 15. Shells of Dicharax species. A. Dicharax (?) ellipticus Páll-Gergely n. sp. (holotype, RMNH 5004014). B. Dicharax (?) cf. ellipticus Páll-Gergely n. sp., Vn11-104. C. Dicharax canaliculatus (von Moellendorff, 1894) (lectotype, SMF 109468); D: Dicharax microdiscus (Möllendorff, 1887) (lectotype, SMF 109496). E. Dicharax parvulus (Möllendorff, 1887) (lectotype, SMF 109507). Scale represents 3 mm. All images: B. Páll-Gergely.
FIGURE 9. R2 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 9. R2 ribs of Dicharax species. A–D. Dicharax (?) alticola Páll-Gergely & Hunyadi n. sp. (A–B. 20130913A, OC1, coll. JUO, C–D. 20040520A, coll. PGB). E–F. Dicharax (?) dolichodeiros (2013/6). All images: B. Páll-Gergely.
FIGURE 27 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 27. Shells of Dicharax cristatus (Möllendorff, 1886). A. holotype of Alycaeus simplicilabris Bavay & Dautzenberg, 1912 (MNHN IM-2012-27167). B. 20060109A. C. MNHN 2012-27041. D. 20111021B, coll. PGB. E. 2011/22, coll. HA. F. 2012/60, coll. HA. Scale represents 5 mm. All photos: B. Páll-Gergely.
FIGURE 8 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 8. Shells of Dicharax species. A. Dicharax (?) alticola Páll-Gergely & Hunyadi n. sp. (holotype, HNHM 99702). B. Dicharax (?) fargesianus (Heude, 1885) (lectotype, MCZ 167229). C. Dicharax (?) immaculatus Páll-Gergely n. sp. (holotype, HNHM 99707). D. Dicharax (?) dolichodeiros Heude, 1890 (2013/6). E. Dicharax dolichodeiros (2016/100). Scale represents 5 mm. All photos: B. Páll-Gergely.
FIGURE 7 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 7. Distribution of narrow range or single-site endemic Dicharax species. Filled circle. Dicharax tangmaiensis (Chen & Zhang, 2001). Filled triangle, top up. Dicharax (?) alticola Páll-Gergely & Hunyadi n. sp. Filled diamond. Dicharax (?) immaculatus Páll-Gergely n. sp. Empty diamond. Dicharax microcostatus Páll-Gergely n. sp., Dicharax micropolitus Páll- Gergely & Hunyadi, n. sp. and Dicharax bison Páll-Gergely & Hunyadi n. sp. Filled triangle, top down. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. Empty triangle, top down. Dicharax draco Páll-Gergely & Hunyadi n. sp. Empty square. Dicharax imitator Páll-Gergely & Hunyadi n. sp. Empty triangle, top up. Dicharax (?) moellendorffi (Kobelt & Möllendorff, 1886). Filled square. Dicharax (?) ellipticus n. sp. Cross. Dicharax (?) abdoui Páll-Gergely n. sp. Empty circle shows the position of Tchen K'eou (Chengkou), which is the type locality of Alycaeus pentagonus Heude, 1886 (synonym of Alycaeus anthostoma Möllendorff, 1885), Dicharax diminutus (Heude, 1885), Dicharax (?) fargesianus (Heude, 1885), Dicharax planorbulus (Heude, 1885), Metalycaeus muciferus (Heude, 1885), Alycaeus dolomiticus Heude, 1890 (synonym of Alycaeus rathouisianus Heude, 1882), Dicharax (?) dolichodeiros (Heude, 1890), Metalycaeus (?) elevatus (Heude, 1886), Metalycaeus (?) expansus (Heude, 1890).
FIGURE 6 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 6. Dicharax (?) abdoui n. sp. (holotype, MNHN IM-2012-27329). A–D. whole shell. E–F. protoconch. G–H. R2 ribs. All images: B. Páll-Gergely.
FIGURE 32 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 32. Distribution of Dicharax species. Triangle: Dicharax fraterculus (Bavay & Dautzenberg, 1900); Circle: Dicharax depressus (Bavay & Dautzenberg, 1912).
FIGURE 4 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 4. Representatives of the genus Dioryx from China and Vietnam. A. Dioryx dautzenbergi Páll-Gergely nom. nov., Phi-Mi, Haut Tonkin, leg. Messager (holotype, MNHN-IM-2000-31788). B. Dioryx messageri (Bavay & Dautzenberg, 1900), That-Khé, leg. Messager (holotype, MNHN-IM-2000-31785). C. Dioryx compactus (Bavay & Dautzenberg, 1900), Tonkin, Bac-Kan, leg. Messager (holotype, MNHN-IM-2000-31796). D. Dioryx globuloides Zilch, 1957, Patung, Hupei: China, coll. Möllendorff (lectotype, SMF 39221). E. Dioryx kobeltianus (Moellendorff, 1875), China, Kiukiang, coll. Möllendorff (holotype, SMF 39298). Scales represent 5 mm; upper scale refers to figs A–B, lower scale refers to figs C–E. Photos: M. Caballer Gutierrez (A–C); B. Páll-Gergely (D–E).
FIGURE 35 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 35. Shells of Dicharax fimbriatus (Bavay & Dautzenberg, 1912). A. DDS7. B. holotype (MNHN IM-2012-27166). C. 20060108D, coll. PGB. D. 20041113A, coll. PGB. E. 20111025B, coll. PGB. Scale represents 5 mm. All photos: B. Páll- Gergely.
FIGURE 29 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 29. Protoconchs of Dicharax species. A–B. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (20000319, spec.1, HNHM 99726, paratype from the type locality). C–D. Dicharax cristatus (Möllendorff, 1886) (20100508A, coll. PGB). E–F. Dicharax depressus (Bavay & Dautzenberg, 1912), MAA1, Specimen2. All images: B. Páll-Gergely.
FIGURE 3 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 3. Representatives of the genus Alycaeus (A–C, E) and Pincerna (D, F) from China, Laos and Vietnam. A. Alycaeus eydouxi Venmans, 1956, Annam, Touranne, leg. Fruhstorfer, coll. Möllendorff, SMF 109290). B. Alycaeus rolfbrandti Maassen, 2006 Laos, Kalkberge ca. 20 km östl. Takek, leg. Brandt 08.09.1963, SMF 262541. C. Alycaeus vanbuensis Bavay & Dautzenberg, 1900, Tonkin, Van-Bu, leg. Dr. R. Bavay (holotype, MNHN-IM-2000-31798). D. Pincerna liratula (Preston, 1907), Ke-lan-tan (syntype, NHMUK 1907.5.20.191). E. Alycaeus anceyi Mabille, 1887, Tonkin, coll. Mabille (syntype, MNHN-IM-2000-31797). F. Pincerna costulosa (Bavay & Dautzenberg, 1912), Tonkin, Phong Tho, leg. Messager (holotype, MNHN-IM-2000-31786). Scales represent 5 mm; upper scale refers to figs A–B, lower scale refers to figs E–F. Photos: M. Caballer Gutierrez (E–F) and B. Páll-Gergely (A–B).
FIGURE 2 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 2. Characters states of the shell and the operculum. Shell shapes from dorsal view (A. oval, B. rounded). Shape of the body whorl (C. slightly keeled, D. rounded). Protoconch (E–F. low, smooth, G–H. elevated, spirally striated). Outer surface of the operculum (I. multispiral without elevated lamella. J. with low lamella. K. with elevated lamella. L. lamella forms a funnelshaped structure). Inner surface of the operculum (M. no nipple. N. low nipple. O. elevated nipple). P–R. swelling on R3 (P. no swelling. Q. blunt swelling. R. sharp swelling). All images by B. Páll-Gergely.
FIGURE 1 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 1. Nomenclature of alycaeid shell characters. A–B. regions of the alycaeid shell. Dark area: Protoconch (embryonic whorls); Red area: R1; Green area: R2, Blue area: R3. Arrow indicates the approximate position of the operculum when the snail is retracted into the shell. C. Schematic diagram of a cross section of R2 (mainly after Páll-Gergely et al. 2016). Arrow indicates the place where the anterior crust folds over the posterior crust. The aperture is situated leftwards. D. aperture. E. peristome. F–G. Character states of the edge of the peristome (F. not crenulated. G. aperture internally crenulated. H. aperture externally crenulated). Abbreviations: A: anterior crust; AP: anterior projection; H: hollow of the anterior crust; IL: inner lip; LI: lower incision; OL: outer lip; P: posterior crust; PP: posterior projection; T: microtunnel; UI: upper incision. All photos: B. Páll-Gergely.
FIGURE 17. R2 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 17. R2 ribs of Dicharax species. A–B. Dicharax (?) fargesianus (Heude, 1885). C–F. Dicharax (?) immaculatus Páll- Gergely n. sp. (20110420E sp1, coll. PGB). All images: B. Páll-Gergely.
FIGURE 10 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 10. Protoconchs of Dicharax species. A–B. Dicharax (?) alticola Páll-Gergely & Hunyadi n. sp. (A. 20130913A, OC1, coll. JUO, B. 20040520A, coll. PGB). C–D. Dicharax (?) fargesianus (Heude, 1885) (lectotype). E–F. Dicharax (?) dolichodeiros (2013/6). All images: B. Páll-Gergely.
FIGURE 28. R2 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 28. R2 ribs of Dicharax species. A–B. Dicharax cristatus (Möllendorff, 1886) (20130911B, coll. JUO). C–D. Dicharax cristatus 20111020A, coll. JUO. E–H. Dicharax depressus (Bavay & Dautzenberg, 1912) (MAA1, HNHM 99727). All images: B. Páll-Gergely.
FIGURE 26 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 26. Shells of Dicharax cristatus (Möllendorff, 1886). A. 2016/103. B. 2016/114. C. SMF 39234. Scale represents 5 mm. All photos: B. Páll-Gergely.
FIGURE 19 in Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
FIGURE 19. Shells of Dicharax species. A. Dicharax anthostoma (Möllendorff, 1885) (lectotype, SMF 39225). B. Dicharax anthostoma (syntype of Alycaeus pentagonus, MCZ 167215). C. Dicharax robustus Páll-Gergely & Hunyadi, n. sp. (holotype, HNHM 99704). Scale represents 5 mm. All photos: B. Páll-Gergely.
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