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Figure 8 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 8 Female paratypes of Cnemaspis lineatubercularis sp. nov. in preservative in A dorsal view B ventral view; from left to right, top panel: ZMKU R 00826, ZMKU R 00832, THNHM 28696 and THNHM 28697; bottom panel: ZMKU R 00833, ZMKU R 00834 and ZMKU R 00835.
Figure 6 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 6 Coloration of Cnemaspis lineatubercularis sp. nov. in dorsal (above) and ventral (below) views of A male paratype ZMKU R 00830 and B female paratype ZMKU R 00835. Note yellowish ventral coloration that is present in males but absent in females.
Figure 7 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 7 Male paratypes of Cnemaspis lineatubercularis sp. nov. in preservative in A dorsal view B ventral view; from left to right, top panel: ZMKU R 00821, ZMKU R 00822, ZMKU R 00823, ZMKU R 00824 and ZMKU R 00825; bottom panel: ZMKU R 00827, ZMKU R 00829, ZMKU R 00830, ZMKU R 00831, THNHM 28694 and THNHM 28695.
Figure 5 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 5 Male holotype (ZMKU R 00828) of Cnemaspis lineatubercularis sp. nov. from Wang Mai Pak Waterfall, Lan Saka District, Nakhon Si Thammarat Province, Thailand, in preservative A lateral view of head B dorsal view of trunk C ventral view of chin D dorsal view E ventral view. Scale bar: 10 mm (D, E).
Figure 4 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 4 Male holotype (ZMKU R 00828) of Cnemaspis lineatubercularis sp. nov. from Wang Mai Pak Waterfall, Lan Saka District, Nakhon Si Thammarat Province, Thailand, in life A dorsal view B ventral view C lateral view of the head D dorsal view of trunk E precloacal region showing distribution of pore-bearing scales (red arrows) F dorsal view of tail G ventral view of tail. Scale bar: 10 mm (in dorsal and ventral views).
Figure 3 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 3 Male holotype (ZMKU R 00828) of Cnemaspis lineatubercularis sp. nov. from Wang Mai Pak Waterfall, Lan Saka District, Nakhon Si Thammarat Province, Thailand.
Figure 2 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 2 A The single best maximum likelihood tree of the mitochondrial NADH dehydrogenase subunit 2 (ND2) gene and flanking tRNAs from geckos of the genera Cnemaspis, Cyrtodactylus and Hemidactylus, shown in full view B map illustrating the localities of Cnemaspis siamensis group samples used in this study and C close-up view of the C. siamensis group. Support values at nodes are bootstrap values from a Maximum Likelihood analysis of the same dataset followed by posterior probabilities of the Bayesian Inference analysis.
Figure 1 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 1 Results of principal component analysis (PCA), and clustering by discriminant function of principal component analysis (DAPC) of 15 morphological variables for 47 individuals of five Cnemaspis species (C. lineatubercularis sp. nov., C. adangrawi, C. chanardi, C. omari, and C. siamensis) APCA scatter plot of PC1 and PC2 showing morphometric differentiation among five species in the siamensis group BDAPC ordination of all samples showing interspecific variation among five species in the siamensis group.
Figure 9 from: Ampai N, Wood Jr PL, Stuart BL, Aowphol A (2020) Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159. https://doi.org/10.3897/zookeys.932.50602
Figure 9 Habitats of Cnemaspis lineatubercularis sp. nov A Wang Mai Pak Waterfall at type locality B microhabitat of holotype in granitic rocky stream (white arrow) C microhabitat of paratypes in granitic rocky outcrops (white arrows) at Wang Mai Pak Waterfall, Lan Saka District, Nakhon Si Thammarat Province, Thailand.
Figure 4 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 4 Phylogram based on the Bayesian analysis. Number above and below branches are MP/ML bootstrap values and Bayesian posterior probabilities (> 50 %), respectively. Dashes represent values < 50 %. Sample AB769531 is from Nishikawa et al. 2013b.
Figure 5 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 5 Scatterplot between PC1 and PC2 of the morphological characters corrected to SVL and log–transformed, for A taxon 1 and taxon 2 of the Vietnamese Tylototriton cf. asperrimusB the head- and dorso- related data of taxon 3 from Lai Chau Province and T. cf. asperrimus from Vietnam sensu lato; and C the limb related data of taxon 3 from Lai Chau Province and T. cf. asperrimus from Vietnam sensu lato. In the graphics T. cf. refers to T. cf. asperrimus.
Figure 3 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 3 Holotype of Tylototriton asperrimus (ZMB 34089). In sequence: dorsal view; ventral view; lateral view with detail of ovaries; and detail of dorsal view of the head. Photographs T. Ziegler.
Figure 2 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 2 Distribution map of the new populations of Tylototriton from North Vietnam, based on the following symbols: square (taxon 3, this study) the population from Sin Ho District, Lai Chau Province; diamond (taxon 1, this study) the upper one identifies the population from Van Ho District, Son La Province, and the lower one identifies the population from Thuong Xuan District, Thanh Hoa Province; triangle (taxon 2, this study) the upper one identifies the population from Tan Son District, Phu Tho Province, the middle one identifies the population from Da Bac District, Hoa Binh Province, and the lower one identifies the population from Lac Son District, Hoa Binh Province. The two populations identified by the circles represent T. asperrimus sensu stricto from China. High resolution remote sensing land cover information was extracted from "GLAD-UMD and SERVIR-Mekong, Natural annual tree canopy structure and surface water dynamics products, 2017" (lower left panel). Bioclimatic variables (right side) were extracted from remote sensing data provided by Deblauwe et al. (2016).
Figure 7 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 7 Boxplot of the most differing characters related to limb values between taxon 1, taxon 2, and taxon 3. Characters were corrected to SVL and log–transformed. For abbreviations see Materials and methods.
Figure 10 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 10 Tylototriton pasmansi obsti ssp. nov. (holotype) In sequence: dorsal view; ventral view; lateral view; and detail of dorsal view of the head. Photographs T. Ziegler.
Figure 1 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 1 Current distribution map of the genus Tylototriton, from South and Central China, to northern Vietnam, Laos, Thailand, Myanmar, India, Bhutan and Nepal (Hernandez 2016; Qian et al. 2017; Grismer et al. 2018; Wang et al. 2018; Grismer et al. 2019; Hernandez et al. 2019; Zaw et al. 2019). In red the distribution areas belonging to the subgenus Tylototriton, and in blue the distribution areas belonging to the sub–genus Yaotriton (Dubois and Raffaëlli 2009). The numerical identification corresponds to the different species and undescribed taxa as follow: 1T. himalayanus2T. cf. himalayanus (Bhutan) 3T. cf. verrucosus (Manipur) 4T. kachinorum5T. verrucosus6T. ngarsuensis7T. shanorum8T. shanjing9T. pseudoverrucosus10T. taliangensis11T. kweichowensis12T. yangi13T. pulcherrimus14T. anguliceps15T. uyenoi16T. podichthys17T. panhai18T. notialis19Tylototriton taxon 2, this study 20Tylototriton taxon 3, this study 21T. ziegleri22T. vietnamensis23T. hainanensis24T. cf. hainanensis (Baise) 25T. cf. wenxianensis (Libo) 26T. cf. wenxianensis (Guizhou) 27T. asperrimus28T. cf. asperrimus (Guangdong) 29T. lizhengchangi30T. liuyangensis31T. broadoridgus32T. cf. wenxianensis (Chongquing and Hubei) 33T. wenxianensis34T. dabienicus35T. anhuiensis and 36T. panwaensis.
Figure 9 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 9 Dorsal and ventral views of the new taxa in life. Top: Tylototriton pasmansi sp. nov. (holotype); Center: Tylototriton pasmansi obsti ssp. nov. (holotype); Bottom: Tylototriton sparreboomi sp. nov. Photographs: C. T. Pham (upper row) and A. V. Pham (central and lower rows).
Figure 11 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 11 Tylototriton sparreboomi sp. nov. (holotype). In sequence: dorsal view; ventral view; lateral view; and detail of dorsal view of the head. Photographs T. Ziegler.
Figure 6 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 6 Boxplot of the most differing characters related to head and dorsal values between taxon 1, taxon 2, and taxon 3. Characters were corrected to SVL and log–transformed. For abbreviations see Materials and methods.
Figure 8 from: Bernardes M, Le MD, Nguyen TQ, Pham CT, Pham AV, Nguyen TT, Rödder D, Bonkowski M, Ziegler T (2020) Integrative taxonomy reveals three new taxa within the Tylototriton asperrimus complex (Caudata, Salamandridae) from Vietnam. ZooKeys 935: 121-164. https://doi.org/10.3897/zookeys.935.37138
Figure 8 Tylototriton pasmansi sp. nov. (holotype). In sequence: dorsal view; ventral view; lateral view; and detail of dorsal view of the head. Photographs T. Ziegler.
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