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Figures 115-120 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 115-120 Megacraspedus adults in dorsal view. 115M.lagopellus Herrich-Schäffer, 1860 – female, Hungary (NHMW) 116M.coleophorodes (Li & Zheng, 1995) – Holotype, male, China (NKU) 117M.coleophorodes (Li & Zheng, 1995) – Paratype, female, China, (NKU) 118M.feminensis sp. n. – Paratype, female, Kazakhstan (RCKN) 119M.kirgizicus sp. n. – Paratype, male, Afghanistan (NHMW) 120M.kirgizicus sp. n. – Paratype, female, Afghanistan (NHMW).
Figures 103-108 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 103-108 Megacraspedus adults in dorsal view. 103M.multispinella Junnilainen & Nupponen, 2010 – male, Russia (MZH) 104M.multispinella Junnilainen & Nupponen, 2010 – female, Russia (MZH) 105M.nupponeni sp. n. – Paratype, male, Russia (RCKN) 106M.nupponeni sp. n. – Paratype, female, Russia (RCKN) 107M.cerussatellus Rebel, 1930 – Lectotype, male, Bulgaria (NHMW) 108M.attritellus Staudinger, 1871 – male, Russia (NMPC).
Figure 6 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figure 6 Maximum likelihood tree (built with MEGA7) of cytochrome c oxidase subunit I (COI) barcode fragments. Values on branches represent bootstrap values of ≥ 50 % inferred from 500 replicates, scale bar represents substitutions per site. Note: the scale bar only applies to internal branches between species. The width of the triangles represents the sample size, and the depth the relative genetic variation within the cluster (2× scale bar). Source: DNA Barcode data from BOLD (Barcode of Life Database, cf. Ratnasingham and Hebert 2007).
Figures 192-194 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 192-194 Megacraspedus male genitalia. 192M.granadensis sp. n. – Holotype, Spain, GU 16/1458 P.H. (ZMUC) 193M.heckfordi sp. n. – Paratype, Spain, GEL 1205 P.H. (TLMF) 194M.tenuiuncus sp. n. – Paratype, Spain, GU 15/1402 P.H. (TLMF).
Figure 5 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figure 5 Terminology of important characters in female genitalia of Megacraspedusniphorrhoa (Meyrick, 1926).
Figures 159-161 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 159-161 Megacraspedus male genitalia. 159M.homochroa Le Cerf, 1932 – Morocco, GU 16/1453 P.H. (ZMUC) 160M.monolorellus Rebel, 1905 – Lectotype, Turkey, gen. slide 16.649 (NHMW) 161M.junnilaineni sp. n. – Holotype, Turkey, GU 16/1448 P.H. (RCJJ).
Figures 201-203 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 201-203 Megacraspedus male genitalia. 201M.barcodiellus sp. n. – Holotype, Macedonia, GEL 1184 P.H. (TLMF) 202M.bilineatella Huemer & Karsholt, 1996 – Paratype, Italy, GEL 561 P.H. (TLMF) 203M.andreneli Varenne & Nel, 2014 – Paratype, France, gen. slide 24592 J. Nel (TLMF).
Figures 177-179 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 177-179 Megacraspedus male genitalia. 177M.cuencellus Caradja, 1920 – Spain, GU 15/1401 P.H. (ZMUC) 178M.bidentatus sp. n. – Holotype, Spain, GU 16/1433 P.H. (ZMUC) 179M.fuscus sp. n. – Holotype, Spain, GU 17/1478 P.H. (ZMUC).
Figure 3 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figure 3 Wing venation of Megacraspedus. aM.lagopellus Herrich-Schäffer, 1860 – male, Armenia, (NMPC), forewing 8.5 mm bM.peyerimhoffi Le Cerf, 1925 – male, Spain (NMPC), forewing 9 mm cM.peyerimhoffi Le Cerf, 1925 – female, Spain (NMPC), forewing 10 mm dM.binotella (Duponchel, 1843) – female, Slovakia (NMPC), forewing 5.5 mm eM.fallax (Mann, 1867) – male, Russia (NMPC), forewing 8.5 mm.
Figures 225-227 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 225-227 Megacraspedus male genitalia. 225M.tenuignathos sp. n. – Holotype, Morocco, GU 16/1423 P.H. (RCAW) 226M.glaberipalpus sp. n. – Paratype, Morocco, GEL 1245 P.H. (TLMF) 227M.glaberipalpus sp. n. – Paratype, Morocco, gen. slide 16.665 (NHMW).
Figures 189-191 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 189-191 Megacraspedus male genitalia. 189M.spinophallus sp. n. – Holotype, Spain, GEL 1208 P.H. (TLMF) 190M.spinophallus sp. n. – Paratype, Spain, gen. slide 33659 (BMNH) 191M.occidentellus sp. n. – Holotype, Portugal, GU 13/1352 P.H. (ZMUC).
Figure 1 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figure 1 Labial palpus of Megacraspedus. aM.squalida Meyrick, 1926 bM.longipalpella Junnilainen, 2010.
Figures 198-200 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 198-200 Megacraspedus male genitalia. 198M.binotella (Duponchel, 1843) – Austria, GEL 1187 P.H. (TLMF) 199M.brachypteris sp. n. – Paratype, Greece, GU 16/1460 P.H. (ZMUC) 200M.brachypteris sp. n. – Paratype, Albania, gen. slide 16.660 (NHMW).
Figures 219-221 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 219-221 Megacraspedus male genitalia. 219M.balneariellus (Chrétien, 1907) – Croatia, GU 16/1457 P.H. (RCJJ) 220M.podolicus (Toll, 1942) – Austria, GEL 1202 P.H. (TLMF) 221M.kazakhstanicus sp. n. – Holotype, Kazakhstan, GU 17/1496 P.H. (RCKN).
Figures 127-132 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 127-132 Megacraspedus adults in dorsal view. 127M.violacellum (Chrétien, 1915) – Lectotype, female (halfella), Tunisia (MNHN) 128M.squalida Meyrick, 1926 – male, Spain (NMPC) 129M.squalida Meyrick, 1926 – female, Spain (NMPC) 130M.pentheres Walsingham, 1920 – male, France (TLMF) 131M.steineri sp. n. – Holotype, male, Morocco (ZMUC) 132M.gibeauxi sp. n. – Holotype, male, Tunisia (ZMUC).
Figures 210-212 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 210-212 Megacraspedus male genitalia. 210M.numidellus (Chrétien, 1915) – Syntype (mareotidellus), Libya, gen. slide 33664 (BMNH) 211M.numidellus (Chrétien, 1915) – Morocco, GU 16/1417 P.H. (ZMUC) 212M.numidellus (Chrétien, 1915) – Spain (Fuerteventura), GU 16/1424 P.H. (ZMUC).
Figures 168-170 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 168-170 Megacraspedus male genitalia. 168M.dolosellus (Zeller, 1839) – Bulgaria, GEL 1216 P.H. (TLMF) 169M.dolosellus (Zeller, 1839) – Greece, GEL 1242 P.H. (TLMF) 170M.dolosellus (Zeller, 1839) – Russia, GEL 1246 P.H. (TLMF).
Figures 109-114 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 109-114 Megacraspedus adults in dorsal view. 109M.consortiella Caradja, 1920 – male, Kyrgyzstan (RCKN) 110M.pototskii sp. n. – Holotype, male, Kyrgyzstan (RCKN) 111M.leuca (Filipjev, 1929) – male, Russia (NMPC) 112M.leuca (Filipjev, 1929) – Paratype (kaszabianus), female, Mongolia (HNHM) 113M.orenburgensis Junnilainen & Nupponen, 2010 – Holotype, male, Russia (RCKN) 114M.lagopellus Herrich-Schäffer, 1860 – male, Hungary (NHMW).
Figures 207-209 from: Huemer P, Karsholt O (2018) Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae). ZooKeys 800: 1-278. https://doi.org/10.3897/zookeys.800.26292
Figures 207-209 Megacraspedus male genitalia. 207M.gallicus sp. n. – Paratype, France, GEL 1203 P.H. (TLMF) 208M.ribbeella (Caradja, 1920) – Spain, gen. slide 16.650 (NHMW) 209M.libycus sp. n. – Holotype, Libya, GU 16/1435 P.H. (ZMUC).
Supplementary material 1 from: Singh G, Aptroot A, Rico VJ, Otte J, Divakar PK, Crespo A, Cáceres ME da S, Lumbsch HT, Schmitt I (2018) Neoprotoparmelia gen. nov. and Maronina (Lecanorales, Protoparmelioideae): species description and generic delimitation using DNA barcodes and phenotypical characters. MycoKeys 44: 19-50. https://doi.org/10.3897/mycokeys.44.29904
Table S1: List of primers used in this study :
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