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Figure 15 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 15 Penial hooks, Peronia peroniiA Mauritius [5872] (MNHN-IM-2019-1605) B Mauritius [3605] (MNHN-IM-2019-1606) C, D Guam [5840] (MNHN-IM-2019-1609). Scale bars: 6 μm (A, B, D), 40 μm (C).
Figure 16 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 16 Accessory penial gland spine, Peronia peroniiA, B, F, G Mauritius C, D, H Papua New Guinea E, I Guam A [3605] (MNHN-IM-2019-1606) B [5872] (MNHN-IM-2019-1605) C [5471] (MNHN-IM-2013-12500) D [5472] (MNHN-IM-2013-14052) E [5840] (MNHN-IM-2019-1609) F same as A; G same as B; H same as C; I same as E. Scale bars: 800 μm (A–D), 1 mm (E), 40 μm (F–I).
Figure 99 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 99 Penial hooks, Peronia verruculata (unit #3), Peninsular Malaysia A, B [2547] (USMMC 00065) C, D [975] (USMMC 00064). Scale bars: 40 μm (A, C), 20 μm (B), 6 μm (D).
Figure 22 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 22 Radula, Peronia madagascariensis, South Africa, [5841] (NMSA W7547) A right half rows of teeth B rachidian and innermost lateral teeth C rachidian and innermost lateral teeth D lateral teeth E lateral teeth, frontal view F outermost lateral teeth. Scale bars: 200 μm (A), 20 μm (B, E, F), 40 μm (C), 100 μm (D).
Figure 11 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 11 Radula, Peronia peroniiA–C Papua New Guinea [5476] (MNHN-IM-2013-16260) D–H Guam [5840] (MNHN-IM-2019-1609) A lateral teeth B rachidian and innermost lateral teeth C rachidian and innermost lateral teeth D rachidian and innermost lateral teeth E rachidian and lateral teeth F lateral teeth G lateral teeth, frontal view H outermost lateral teeth. Scale bars: 100 μm (A), 40 μm (B, D), 80 μm (C), 200 μm (E, F), 100 μm (G, H).
Figure 106 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 106 Accessory penial gland spine, Peronia verruculata (unit #3) A–C, E–I, K, L Peninsular Malaysia D, J Singapore A [975] (USMMC 00064) B [976] (USMMC 00051) C [977] (USMMC 00064) D [989] (ZRC.MOL.16070) E [2546] (USMMC 00065) F [2547] (USMMC 00065) G same as A; H same as B; I same as C; J same as D; K same as E; L same as F. Scale bars: 300 μm (A–F), 20 μm (G–L).
Figure 105 from: Dayrat B, Goulding TC, Apte D, Aslam S, Bourke A, Comendador J, Khalil M, Ngô XQ, Tan SK, Tan SH (2020) Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae). ZooKeys 972: 1-224. https://doi.org/10.3897/zookeys.972.52853
Figure 105 Accessory penial gland spine, Peronia verruculata (unit #2), Indonesia, Sumatra A, E [1797] (UMIZ 00180) B, F [1795] (UMIZ 00180) C, G [1796] (UMIZ 00180) D [1746] (UMIZ 00178). Scale bars: 300 μm (A–C), 10 μm (D–G).
Figure 9 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 9 - Habitats for Onchidium stuxbergi. A Malaysia, Kuala Sepatang, old forest with tall, old Rhizophora trees, high in the tidal zone (station 27) B Philippines, Bohol, mostly Nypa palms with Thalassina mounds (station 187) C Vietnam, Can Gio, open mangrove with large Avicennia trees, soft mud, some old logs (station 231) D Vietnam, Can Gio, hard mud with trees spread out by a small road and then a steep bank to the soft, deep mud (station 221).
Figure 6 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 6 - Radula, Onchidium typhae, Malaysia, Matang (USMMC 00005, #1). A Outermost lateral teeth, scale bar 20 µm B Lateral teeth, scale bar 20 µm C Lateral teeth, inferior view, showing the additional outer spine of each tooth, scale bar 10 µm D Rachidian and innermost lateral teeth, scale bar 20 µm.
Figure 7 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 7 - Nervous and reproductive systems, Onchidium typhae. A Nervous system, Malaysia, Langkawi Island, scale bar 3.8 mm (USMMC 00002, #1) B Hermaphroditic (female), posterior parts, Singapore, scale bar 3.8 mm (ZRC.MOL.6397, #1) C Male, anterior, copulatory parts, Malaysia, Matang, scale bar 5 mm (USMMC 00005, #1) D Male, anterior, copulatory parts, Malaysia, Matang, scale bar 2.7 mm (USMMC 00005, #2). Abbreviations: agf, accessory gland flagellum; dd, deferent duct; fgm, female gland mass; hd, hermaphroditic duct; hg, hermaphroditic gland; lcg, left cerebral ganglion; lpg, left pedal ganglion; lplg, left pleural ganglion; ov, oviduct; ps, penial sheath; rcg, right cerebral ganglion; rm, retractor muscle; rpg, right pedal ganglion; rplg, right pleural ganglion; rs, receptaculum seminis; sp, spermatheca; spv, spermoviduct; v, vestibule; vg, visceral ganglion.
Figure 8 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 8 - Male, anterior, copulatory parts, Onchidium typhae. A Penial hooks, Malaysia, Matang, scale bar 30 µm (USMMC 00005, #1) B Flat disc at distal end of flagellum of penial accessory gland, Malaysia, Langkawi Island, scale bar 100 µm (USMMC 00002, #1) C Tip of hollow spine, penial accessory gland, Singapore, scale bar 4 µm (ZRC.MOL.6397, #1) D Detail of B, scale bar 20 µm (USMMC 00002, #1) E Hollow spine, penial accessory gland, Singapore, scale bar 100 µm (ZRC.MOL.6397, #1) F Hollow spine, penial accessory gland, Malaysia, Langkawi Island, scale bar 100 µm (USMMC 00002, #1).
Figure 4 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 4 - Live specimens, Onchidium typhae. A Dorsal view, 45 mm long, India, West Bengal, station 48 (BNHS) B Dorsal view, 55 mm long, India, Andaman Islands, station 57 (BNHS) C Dorsal view, 40 mm long, Singapore (ZRC.MOL.6396) D Frontal view, 25 mm wide, India, Andaman Islands, station 57 (BNHS) E Ventral view, 50 mm long, India, Andaman Islands, station 59 (BNHS) F Ventral view, 40 mm long [DNA 1064], India, West Bengal, station 49 (BNHS) G Dorso-lateral view, 40 mm long, India, Andaman Islands, station 59 (BNHS) H Dorsal view, 42 mm long, Malaysia, Matang (USMMC 00005, #1).
Figure 5 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 5 - External morphology and digestive system, Onchidium typhae. A Anterior region, ventral view, Singapore, scale bar 8 mm (ZRC.MOL.6396) B Posterior region, ventral view, Singapore, scale bar 7 mm (ZRC.MOL.6396) C Dorsal papillae (preserved), Singapore, station 8, scale bar 3.8 mm (ZRC.MOL.6396) D Buccal gland, Malaysia, Langkawi Island, scale bar 3 mm (USMMC 00002, #1) E Digestive system, dorsal view, Singapore, scale bar 5 mm (ZRC.MOL.6397, #1) F Digestive system, ventral view, Singapore, scale bar 5 mm (ZRC.MOL.6397, #1) G Stomach (digestive gland removed), ventral view, Singapore, scale bar 5 mm (ZRC.MOL.6397, #1). Abbreviations: ddg, dorsal lobe of digestive gland; e, esophagus; i, intestine; ldg, lateral lobe of the digestive gland; pdg, posterior lobe of the digestive gland; rg, rectal gland; st1, stomach chamber 1; st2, stomach chamber 2; st3, stomach chamber 3; st4, stomach chamber 4.
Figure 14 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 14 - Internal anatomy, holotype, Onchidium reevesii (NHMUK 20160036). A Digestive system, dorsal view, scale bar 7.5 mm B Reproductive system, anterior, penial accessory gland, scale bar 3 mm C Reproductive system, anterior, penial sheath, scale bar 5 mm. Abbreviations: agf, accessory gland flagellum; dd, deferent duct; ddg, dorsal lobe of digestive gland; i, intestine; pdg, posterior lobe of the digestive gland; ps, penial sheath; rg, rectal gland; rm, retractor muscle; st2, stomach chamber 2.
Figure 12 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 12 - Radula, Onchidium stuxbergi. A Rachidian and innermost lateral teeth, Brunei Darussalam, scale bar 40 µm (BDMNH # . B Rachidian and innermost lateral teeth, Philippines, Bohol, scale bar 20 µm (PNM 041202 #1) C Half rows, Philippines, Bohol, scale bar 200 µm (PNM 041202 #1) D Lateral teeth, Philippines, Bohol, scale bar 30 µm (PNM 041202 #1) E Outermost lateral teeth, Philippines, Bohol, scale bar 20 µm (PNM 041202 #1).
Figure 11 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 11 - External morphology and internal anatomy, Onchidium stuxbergi. A Anterior region, dorsal view, Malaysia, Kuala Sepatang, scale bar 4.4 mm [DNA 971] (USMMC 00006) B Digestive system, dorsal view, Philippines, Bohol, scale bar 5 mm [DNA 3363] (PNM 041199, #2) C Reproductive system, anterior, male, copulatory parts, Philippines, Bohol, scale bar 6.8 mm [DNA 3363] (PNM 041199, #2) D Reproductive system, hermaphroditic (female) posterior parts, Malaysia, Kuala Sepatang, scale bar 5 mm [DNA 971] (USMMC 00006). Abbreviations: agf, accessory gland flagellum; dd, deferent duct; ddg, dorsal lobe of digestive gland; e, esophagus; fgm, female gland mass; hg, hermaphroditic gland; i, intestine; ov, oviduct; pdg, posterior lobe of the digestive gland; ps, penial sheath; rg, rectal gland; rm, retractor muscle; rs, receptaculum seminis; sp, spermatheca; spv, spermoviduct; st2, stomach chamber 2.
Figure 1 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 1 - Phylogenetic tree. Relationships within the genus Onchidium based on COI and 16S sequences. Numbers above branches are the bootstrap values (Maximum Likelihood analysis) and below are the posterior probabilities (Bayesian analysis); only significant numbers (> 80% and > 0.9) are indicated. Onchidella and Peronia sequences serve as outgroups. Numbers for each individual correspond to unique identifiers in our DNA collection. All sequences of Onchidium specimens are new, with the exception of the specimens from China which were all misidentified as Onchidium "struma" by Sun et al. (2014). Information on individually-identified specimens can be found in the additional material examined and in Table 1.
Figure 13 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 13 - Male, anterior, copulatory parts, Onchidium stuxbergi. A Penial hooks, Philippines, Bohol, scale bar 40 µm [DNA 3363] (PNM 041199, #2) B Penial hooks, Malaysia, Kuala Sepatang, scale bar 20 µm [DNA 971] (USMMC 00006) C Flat disc at distal end of flagellum of penial accessory gland, Malaysia, Kuala Sepatang, scale bar 40 µm [DNA 971] (USMMC 00006) D Tip, hollow spine, penial accessory gland, Philippines, Bohol, scale bar 3 µm [DNA 3363] (PNM 041199, #2) E Hollow spine, penial accessory gland, Philippines, Bohol, scale bar 200 µm [DNA 3363] (PNM 041199, #2).
Figure 10 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 10 - Live specimens, Onchidium stuxbergi. A Dorsal view, 33 mm long, Brunei Darussalam (BDMNH, #1) B Dorsal view, 27 mm long, Philippines, Bohol (PNM 041200) C Dorsal view, 35 mm long, Bohol, Philippines (PNM 041201) D Dorsal view, 35 mm long, Vietnam, Can Gio (ITBZC IM 00002, #1) E Dorsal view, 30 mm long [DNA 3251], Bohol, Philippines (PNM 041199) F Ventral view, 35 mm long [DNA 3363], Bohol, Philippines (PNM 041202, #2).
Figure 3 from: Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Ngô XQ, Tan SK, Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca, Gastropoda, Pulmonata, Onchidiidae). ZooKeys 636: 1-40. https://doi.org/10.3897/zookeys.636.8879
Figure 3 - Habitats for Onchidium typhae. A India, West Bengal, very soft mud on the shore with recently-planted Avicennia trees (station 49) B India, Andaman Islands, by a large river, deep mangrove with tall trees, small creeks, and plenty of old muddy logs (station 57) C Malaysia, Matang, old and open Rhizophora forest with tall trees, hard mud, creeks, and many old logs (station 28) D Malaysia, Langkawi Island, open forest with high mud lobster (Thalassina) mounds (station 26).
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