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FIGURE 24 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 24. Radulae of Clavus devexistriatus n. sp., C. hylicos n. sp. and C. maestratii n. sp. A, B. Clavus devexistriatus, New Caledonia, LAGON, Stn. 933, SL 12.8 mm (shell—Fig. 22 C); C, D. Clavus hylikos; New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1260, SL 7.5 mm (shell—Fig. 23 D). E, F. Clavus maestratii, EXPÉDITION MONTROUZIER, Koumac Stn. 1309, SL 9.8 mm (shell—Fig. 25 E).

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FIGURE 21 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 21. Clavus laetus (Hinds, 1843) (A–D) and Clavus squamiferus n. sp. (E–I). A, B. Holotype of Clavatula laeta NHMUK 1879.2.26.67, New Guinea and Straits of Makassar, 7–10 fath.[13–18 m]; C. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1446, SL 10.2 mm; D. Protoconch, New Caledonia, EXPÉDITION MONTROUZIER Touho, Stn. 1270, SL 8.5 mm; E, F. Clavus squamiferus, Holotype; G. Paratype, MNHN IM–2000–26933 SL 10.2 mm; H. Paratype, Radula voucher, MNHN IM–2000–26935, New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1273, SL 11.5 mm (radula—Fig. 18 G, H); I. Protoconch, New Caledonia, EXPÉDITION MONTROUZIER Touho, Stn. 1269, SL 3.3 mm.

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FIGURE 18 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 18. Radulae of Clavus virginieae n. sp., C. picoides n. sp. and C. squamiferus n. sp. A, B. Clavus virginieae, Lifou, LIFOU 2000, Stn. 1429, SL 10.3 mm (shell—Fig. 17 D); C, D. Clavus picoides, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1453, SL 14.0 mm (shell—Fig. 20 D); E, F. Clavus squamiferus, Paratype, Radula voucher, MNHN IM–2000–26935, New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1273, SL 11.5 mm (shell—Fig. 21 H).

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FIGURE 17. Clavus virginieae n in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 17. Clavus virginieae n. sp. A, B. Holotype; C. Largest paratype, MNHN IM–2000–26925, 12.5 mm; D. Radula voucher Loyalty Islands, Lifou, LIFOU 2000, Stn. 1429, SL 10.3 mm (radula—Fig. 18 C, D); E. Molecular voucher IM–2007–42549, Vanuatu, Aore Is., SANTO 2006, Stn. DB12, 15°36.6'S 167°10.1'E, 10–18 m, SL 11.1 mm; F, G. Protoconch, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1456, SL 2.4 mm.

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FIGURE 15 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 15. Clavus humilis (E. A. Smith, 1879). A. Drillia humilis, holotype NHMUK 1878.11.7.14, Ojica Bay, Goto, Japan, 18 m; 9.5 x 3.4 mm; B, C. New Caledonia, LAGON, Stn. 656, Canala Sector, SL 12.8 mm; D, E. New Caledonia, EXPEDITION MONTROUZIER, Touho Stn. 1261, SL 9.7 mm.

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FIGURE 20. Clavus picoides n in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 20. Clavus picoides n. sp. A, B. Holotype; C. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1432, SL 12.1 mm; D. Radula voucher, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1453, SL 14.0 mm (radula—Fig. 18 E,F).

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FIGURE 13 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 13. Clavus lamberti (Montrouzier, 1860). A, B. Holotype of Pleurotoma lamberti Montrouzier, 1860, MHNBx 2004.TY.39.1, New Caledonia, Art Is, SL 9.8 mm; C, D. New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1273, 10.5 x 4.3 mm; E. Radula voucher, same locality as previous, SL 10.4 mm; F, G. New Caledonia, EXPÉDITION MONTROUZIER Touho, Stn. 1255, SL 4.5 mm.

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FIGURE 14 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 14. Clavus obliquicostatus (Reeve, 1845). A, B. Pleurotoma obliquicostata, syntype NHMUK1991048; type locality unknown; C, D. Possible syntype of Drillia leforestieri (Hervier, 1896), MNHN IM–2000–3067, Loyalty Islands, Lifou, 15.7 x 6.4 mm; E. LAGON, Stn. 542, Grand Récif Sud, SL 15.3 mm; F. PLOUVEAL Stn. 1221, SL 15.1 mm; G, H. New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1242, 12.2 x 5.1 mm; I. Radula voucher, New Caledonia, EXPÉDITION MONTROUZIER, Koumac Stn. 1283, SL 12.6 mm (radula—Fig. 12 E,F); J. Protoconch, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1424.

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FIGURE 12 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 12. Radulae of Clavus bilineatus, C. lamberti; C. obliquicostatus and C. formosus. A, B. Clavus bilineatus, LAGON, Stn. 68, SL 13.8 mm (shell—Fig. 11H); C, D. Clavus lamberti, New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1273, SL 10.4 mm (shell—Fig. 13 E); E, F. Clavus obliquicostatus, New Caledonia, EXPÉDITION MONTROUZIER, Koumac Stn. 1283, SL 12.6 mm (shell—Fig. 14 I); G, H. Clavus formosus (shell—Fig. 16 F).

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FIGURE 9. Clavus protentus Hervier, 1896. A, B in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 9. Clavus protentus Hervier, 1896. A, B. Probable syntype of Clavus protentus, MNHN IM–2000–3195, Loyalty Islands, Lifou, 11.5 x 4.4 mm; C. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1419, 12.0 x 4.7 mm; D, E. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1451, 11.2 x 4.7 mm; F. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1466, SL 12.1 mm; G. Radula voucher, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1433, SL 13.5 mm (radula—Fig. 7 E,F); H. Protoconch, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1434, SL 3.6 mm.

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FIGURE 11 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 11. Clavus bilineatus (Reeve, 1845). A, B. Syntype of Pleurotoma bilineatus, NHMUK 1963473, Philippines, "Capul and Mindoro Islands", 15 fath. [27 m], 13.3 x 5.2 mm; C, D. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1422, 16.9 x 7.0 mm; E. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1439, 17.0 x 6.8 mm; F. New Caledonia, LAGON Stn. 713, Canala Sector, SL 15.1 mm; G. Philippines, Balicasag Is, lumun-lumun net, 60–80 m, SL 14.2 mm; H. Radula voucher, LAGON, Stn. 68, Ile Ouen-Baie du Prony, SL 13.8 mm (radula—Fig. 12 A,B); I. Protoconch, New Caledonia, EXPÉDITION MONTROUZIER Touho, Stn. 1242, SL 3.3 mm.

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FIGURE 19 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 19. Clavus moquiniana (Montrouzier in Souverbie & Montrouzier, 1874) and Clavus pica (Reeve, 1845). A, B. Holotype of Pleurotoma moquiniana Montrouzier in Souverbie & Montrouzier, 1874, MHNBx 2004.TY.30, New Caledonia, Art Is, SL 12.2 mm, SW 4.5 mm; C, D. Clavus pica, LAGON, Stn. 1354, SL 20.9 mm.

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FIGURE 10. Clavus delphineae n in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 10. Clavus delphineae n. sp. A, B. Holotype; C. Paratype, MNHN IM–2000–26921, 12.3 x 5.2 mm; D. Protoconch, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1422, 3.3 mm; E. Radula voucher, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1422, SL 10.1 mm (radula—Fig. 7 G, H).

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FIGURE 23. Clavus hylikos n in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 23. Clavus hylikos n. sp. A. B. Holotype; C. Paratype, MNHN IM–2000–26941, SL 8.7 mm; D. Radula voucher, New Caledonia, EXPÉDITION MONTROUZIER, Touho Stn. 1260, SL 7.5 mm (radula—Fig. 24 C, D).

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FIGURE 16 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 16. Clavus formosus (Reeve, 1846). A. Pleurotoma formosa, syntype, NHMUK 19631862, Philippines, Capul Is., SL 12.0 mm; B, C. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1454, 13.4 x 5.0 mm; D, E. Narrow form, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1434, 9.4 x 3.5 mm; F. Radula voucher, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1457, SL 10.1 mm (radula—Fig. 12 G, H).

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FIGURE 7 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 7. Radulae of Clavus unizonalis, C. albotuberculatus, C. protentus and C. delphineae. A, B. Clavus unizonalis, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1424, SL 20.1 (shell—Fig. 6 D); C, D. Clavus albotuberculatus, LAGON, Stn. 445, SL 56.8 mm (shell—Fig. 8 C, D); E, F. Clavus protentus, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1433, SL 13.5 mm (shell—Fig. 9 G); G, H. Clavus delphineae, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1422, SL 10.1 mm (shell—Fig. 10E).

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FIGURE 6 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 6. Clavus unizonalis (Lamarck, 1822). A. Holotype of Pleurotoma unizonalis, MHNG 1097/55, locality unknown, 21.9 x 10.5 mm; B, C. Syntype of Pleurotoma vidua Reeve, 1845, NHMUK 1966435, Masbate Is., Philippines, 19.6 x 8.7 mm; D. Radula voucher, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1424, SL 20.1 (radula—Fig. 7 A–B); E, F. form vidualoides New Caledonia, EXPÉDITION MONTROUZIER Touho, Stn. 1245, 15.9 x 7.2 mm; G. Molecular voucher IM–2007–42557, Vanuatu, Belmoul lagoon, SANTO 2006 Stn. VM09, 15°35.8'S 167°06.1'E, 15.4 mm; H. Molecular voucher IM–2009–16930, Vanuatu, Malo Is, SANTO 2006 Stn. LM19, 15°38.5'S– 167°15.1'E, 22.5 mm; I. Protoconch, New Caledonia, EXPÉDITION MONTROUZIER, Touho, Stn. 1248, SL 4.1 mm.

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FIGURE 8. Clavus albotuberculatus. A, B in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 8. Clavus albotuberculatus. A, B. Holotype of Drillia albotuberculata Schepman, 1889, RMNH.MOL.122139, Locality unknown, 42.1 x 18.3 mm; C, D. Radula voucher, LAGON, Stn. 445, 18º18'S, 163º02'E, 41 m, Surprise Atoll, 56.8 x 26.3 mm (radula—Fig. 7 C, D).

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FIGURE 5. Clavus boucheti n in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 5. Clavus boucheti n. sp. A, B. Holotype; C. Paratype, MNHN IM–2000–26918; D. Paratype, MNHN IM–2000–26916; E. New Caledonia, LAGON, Stn. 290, Noumea Sector, SL 21.0 mm; F. Radula voucher, New Caledonia, LAGON, Stn. 305, Grand Récif Sud, SL 24.5 mm (radula—Fig. 2 G–H).

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FIGURE 4 in The shallow-water New Caledonia Drilliidae of genus Clavus Montfort, 1810 (Mollusca: Gastropoda: Conoidea)

FIGURE 4. Clavus exasperatus (Reeve, 1843). A, B. Probable holotype, NHMUK 1966462, Cuming colln, locality unknown. 23.7 x 11.2 mm; C. Molecular voucher IM–2007–42576, Vanuatu, Aese Is., SANTO 2006 Stn. FR30, 15°27.3'S 167°15.6'E, ca. 45 m, SL 23.7 mm; D. Molecular voucher IM–2009–16929, Vanuatu, SANTO 2006 Stn. DR52, 23.7 mm; E, F. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1422, SL 27.7 mm; G. Loyalty Islands, Lifou, LIFOU 2000, Stn. 1457, SL 24.6 mm; H. Radula voucher, Loyalty Islands, Lifou, LIFOU 2000, Stn. 1420, SL 25.9 mmv(radula –Fig. 2 E–F).

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