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Figure 34 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 34 Mangrove habitats, Wallaconchis ater. A–B Indonesia, Bali, Pemuteran, Labuhan Lalang Harbor, coral rubble, rocks and a few Avicennia (st 157) C–D Indonesia, Lombok, Don Don, old Avicennia forest, with coral rubble (st 149) E–F Philippines, Bohol, Mabini, open Avicennia and Sonneratia mangrove with sand and coral rubble (st 196).
Figure 18 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 18 Penis, Wallaconchis uncinus, Indonesia. A Two most distal loops of fully evaginated penis (the loop on the right is proximal to the loop on the left), Halmahera, scale bar 0.5 mm [5079] (UMIZ 00011) B Two most distal loops of (nearly fully evaginated) penis (the loop on the right is proximal to the loop on the left), Ambon, scale bar 300 μm [2843] (UMIZ 00007) C Proximal region of penis, detail view of curved hooks, Sulawesi, scale bar 100 μm [2268] (UMIZ 00005) D Distal region of penis which is not fully evaginated, scale bar 300 μm, same as C. Abbreviations: ch curved hooks ih immature hooks rh rounded hooks.
Figure 33 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 33 Penis, Wallaconchis nangkauriense, Philippines. A Luzon, scale bar 200 μm [3208] (PNM 041205) B Bohol, scale bar 200 μm [3268] (PNM 041206).
Figure 17 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 17 Reproductive system, Wallaconchis uncinus, Indonesia. A Hermaphroditic (female), posterior parts, Sulawesi, scale bar 2 mm [2268] (UMIZ 00005) B Anterior, male copulatory parts, Halmahera, scale bar 3 mm [5079] (UMIZ 00011) C Anterior, male copulatory parts, Bali, scale bar 1 mm [3138] (UMIZ 00008) D Penis (fully evaginated), Halmahera, scale bar 0.5 mm [5079] (UMIZ 00011). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca v vestibule.
Figure 15 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 15 External morphology and digestive system, Wallaconchis uncinus, Indonesia. A Anterior region, ventral view, Sulawesi, scale bar 2 mm [2268] (UMIZ 00005) B Posterior region, ventral view, Sulawesi, scale bar 2 mm [2268] (UMIZ 00005) C Dorsal notum (papillae with dorsal eyes), Halmahera, scale bar 2.6 mm [5056] (UMIZ 00009) D Digestive system, dorsal view, Sulawesi, scale bar 1 mm [2268] (UMIZ 00005). Abbreviations: a anus ddg dorsal lobe of digestive gland f foot fo female opening hn hyponotum i intestine mo male opening ol oral lobe ot ocular tentacle pdg posterior lobe of the digestive gland pn pneumostome ppg peripodial groove st2 stomach chamber 2.
Figure 22 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 22 Reproductive system, Wallaconchis buetschlii. A Hermaphroditic (female), posterior parts, Indonesia, Sulawesi, scale bar 2 mm [2120] (UMIZ 00021) B Anterior, male copulatory parts, Australia, Queensland, scale bar 1 mm [2555] (MTQ st100) C Anterior, male copulatory parts, Indonesia, Kei, scale bar 1 mm [2933] (UMIZ 00029). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland of oviduct fibers attaching to the inner wall of the visceral cavity ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
Figure 14 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 14 Live specimens, Wallaconchis uncinus, Indonesia. A Dorsal view, 25 mm long [5070], Halmahera (UMIZ 00010) B Holotype, dorsal view, 22 mm long [2751], Ambon (UMIZ 00004) C Dorsal view, 30 mm long [2752], Ambon (UMIZ 00006) D Dorsal view, 28 mm long [2843], Ambon (UMIZ 00008) E Dorsal view, 15 mm long [3138], Bali (UMIZ 00005) F Dorsal view, 25 mm long [5080], Halmahera (UMIZ 00011) G Ventral view, same as F H Ventral view, same as B.
Figure 27 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 27 Reproductive system, Wallaconchis gracile, Indonesia. A Hermaphroditic (female), posterior parts, Sulawesi, scale bar 2 mm [2280] (UMIZ 00055) B Anterior, male copulatory parts, Bali, scale bar 2 mm [3106] (UMIZ 00056). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland of oviduct fibers attaching to the inner wall of the visceral cavity ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
Figure 12 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 12 Penis, Wallaconchis sinanui, Indonesia, Ambon. A Penis evaginated outside the male opening, scale bar 200 μm [5846] (UMIZ 00059) B Penis removed from vestibule, scale bar 20 μm [5844] (UMIZ 00059).
Figure 24 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 24 Habitats, Wallaconchis gracile, Indonesia. A–B Bali, Gilimanuk, muddy-sandy beach near a mangrove (st 155) C–D Ternate, sandy mudflat adjacent to a rocky beach (st 220) E–F Timor, sandy area of mangrove with ripples in the sand and near crab pellets (st254).
Figure 13 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 13 Habitats, Wallaconchis uncinus, Indonesia. A, B Sulawesi, Wori, tall mangrove forest of Sonneratia and some Avicennia with a rocky area and dead logs (st 90) C Ambon, Lateri, muddy mangrove with Rhizophora and Avicennia (st 131) D Type locality, Ambon, Lateri, mudflat adjacent to a shallow river through a mangrove (st 128) E Halmahera, Sofifi, mangrove with Sonneratia trees, dense roots and hard mud (st 205) F Ternate, Bastiong, muddy rocks nearby a mangrove patch (st 203).
Figure 11 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 11 Reproductive system, Wallaconchis sinanui, holotype, Indonesia, Ambon [2737] (UMIZ 00058). A Hermaphroditic (female), posterior parts, scale bar 0.8 mm B Anterior, male copulatory parts, scale bar 2 mm. Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca v vestibule.
Figure 41 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 41 Reproductive system, anterior, male copulatory parts, Wallaconchis ater. A Vestibule removed, showing the coiled and U-shaped penis, Indonesia, Sulawesi, scale bar 3 mm [2220] (UMIZ 00050) B Vestibule with coiled penis inside, Philippines, Bohol, scale bar 3 mm [3270] (PNM 041220). Abbreviations: dd deferent duct dpp distal penial projections p penis ps penial sheath rm retractor muscle v vestibule.
Figure 40 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 40 Reproductive system, Wallaconchis ater, Indonesia. A Anterior, male copulatory parts (vestibule removed, showing the coiled and U-shaped penis), Halmahera, scale bar 2 mm [5078] (UMIZ 00047) B Penis (coil of loops removed from vestibule), Sulawesi, scale bar 2 mm [2283] (UMIZ 00038) C Anterior, male copulatory parts, scale bar 5 mm, same as B D Anterior, male copulatory parts, Kei Islands, scale bar 1 mm [2939] (UMIZ 00041). Abbreviations: dd deferent duct dpp distal penial projections p penis ps penial sheath rm retractor muscle v vestibule.
Figure 10 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 10 Nervous system, Wallaconchis sinanui, holotype, Indonesia, Ambon [2737] (UMIZ 00058) scale bar 0.5 mm. Abbreviations: lcg left cerebral ganglion lpg left pedal ganglion lplg left pleural ganglion rcg right cerebral ganglion rpg right pedal ganglion rplg right pleural ganglion vg visceral ganglion.
Figure 19 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 19 Habitats, Wallaconchis buetschlii. A Indonesia, Sulawesi, Wori, tall mangrove forest of Sonneratia and some Avicennia with a rocky area and dead logs (st 84) B Australia, Queensland, Yule Point, sandy area with a few Rhizophora and Avicennia trees (st 100) C Indonesia, Ambon, Wai, narrow band of old Avicennia trees on sandy mud, with old logs on ground (st 132) D Indonesia, Lombok, Don Don, old Avicennia forest, with coral rubble (st 149) E Indonesia, Sulawesi, Mantehage Island, rocks behind a mangrove of Sonneratia and Rhizophora (st 91) F Indonesia, Bali, Gilimanuk, sandy mudflat outside Rhizophora and Avicennia mangrove (st 156).
Figure 32 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 32 Reproductive system, Wallaconchis nangkauriense. A Hermaphroditic (female), posterior parts, Philippines, Bohol, scale bar 4 mm [3276] (PNM 041206) B Anterior, male copulatory parts, Indonesia, Sulawesi, scale bar 5 mm [2192] (UMIZ 00014) C Anterior, male copulatory parts (vestibule removed around the penis), Indonesia, Sumatra, scale bar 2 mm [1788] (UMIZ 00012). Abbreviations: bc body cavity bw lateral left body wall dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland ov oviduct p penis ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
Figure 4 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 4 Maximum Likelihood phylogenetic tree based on concatenated nuclear ITS1, ITS2, and 28S sequences. Numbers by branches show bootstrap values (only numbers >80% are indicated). All sequences of Wallaconchis specimens are new. Information on individually-identified specimens can be found in the lists of material examined and in Table 1. The color used for each species is the same as the color used in the other trees (Figs 1–3, 5) and on the map of species distribution (Fig. 6).
Figure 21 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 21 Radula, Wallaconchis buetschlii (A–D) Indonesia, Sulawesi [2120] (UMIZ 00021) (E) Australia, Queensland [2554] (MTQ st100) (F) Australia, Queensland [2555] (MTQ st100). A Rachidian and innermost lateral teeth, scale bar 10 μm B Rachidian and innermost lateral teeth, scale bar 40 μm C Inner lateral teeth, scale bar 30 μm D Outermost lateral teeth, scale bar 20 μm E Outermost lateral teeth, scale bar 10 μm F Transition between inner lateral teeth and outer lateral teeth, with the basal lateral spine revealed on the underside of the inner lateral teeth at the bottom, scale bar 30 μm.
Figure 31 from: Goulding TC, Khalil M, Tan SH, Dayrat B (2018) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae). ZooKeys 763: 1-111. https://doi.org/10.3897/zookeys.763.21252
Figure 31 Radula, Wallaconchis nangkauriense, Indonesia, Bali [3129] (UMIZ 00020). A Rachidian and innermost lateral teeth, scale bar 20 μm B Innermost lateral teeth, scale bar 20 μm C Outer lateral teeth, scale bar 20 μm D Outermost lateral teeth, scale bar 20 μm.
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Allen Brain Atlas
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