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PLATE XIII. FIGURES 48b–51. Male genitalia. 48b in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE XIII. FIGURES 48b–51. Male genitalia. 48b. Metarctia (Hebena) haraldsulaki sp.n., paratype, Angola, Huambo Province, Chnigenge, GP RF 661.2021 (RCRF); 49a. Metarctia (Hebena) henrardi, holotype, [DRC] Terr. Kanda-Kanda, Gandajika, GP LP319 (RMCA); 49b. Paratype of Metarctia subincarnata Kiriakoff, 1954 [DRC] Lupweji, GP LP321 (RBINS); 50a. Metarctia (Hebena) lukaszi sp.n., holotype, Angola, Benguela Province, GP 28.685 (RCRF/ZSM); 50b. Idem, paratype, same data as in 50a, GP RF 659.2021 (RCRF); 51. Metarctia (Hebena) venustissima, holotype, [DRC] Katanga, Kolwezi, GP LP147 (RMCA).

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PLATE XVIII. FIGURES 73–78. Habitat photos. 73–74 in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE XVIII. FIGURES 73–78. Habitat photos. 73–74. Habitat of Metarctia (Hebena) haraldsulaki sp.n.. 73. Angola, Bié Province, ca. 5 km N Sao Lucas, 1690m; 74. Type locality, Angola, Cuanza Sul Province, ca. 80km E Quibala, 1352m; 75–76. Habitat of Metarctia (Hebena) haraldsulaki sp.n. and Metarctia (Hebena) lukaszi sp.n., Angola, Huambo Province, Serra do Dembi, ca. 5km NW Quingenge, 1400 m (73–76. © S. Naumann); 77–78. Habitat of Metarctia (Hebena) lateritia. 77. Uganda western, Semliki NP, 670 m; 78. Uganda western, Mburo Lake NP, Nshara Gate, Rwakobo Rock Lodge, 1291 m (77–78. © R. Fiebig)

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PLATE XI. FIGURES 41b–44. Male genitalia. 41b in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE XI. FIGURES 41b–44. Male genitalia. 41b. Metarctia (Hebena) rubra, RSA, Waterberg mountain, GP 36.748 (RCRF); 41c. Idem, RSA, Free Sate Province, Drakensberge, GP 36.555 (RCRF); 42. Metarctia (Hebena) manfredi sp.n., holotype, Kenya, Kibwezi, GP 36.549 (RCRF/ZSM); 43a. Metarctia (Hebena) kelleni, lectotype, [Angola] Humpata, RMNH.INS.1298160 (uncus broken) (RMNH); 43b. Idem, paratype of Metarctia titan Talbot, 1929, Angola, Upper Cubango-Cunene Watershede, GP LG 4701 (NHMUK); 44. Metarctia (Hebena) elleni sp.n., holotype, Tanzania, Rukwa Region, Mbizi Mts., GP 30.955 (RCRF/ZSM).

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PLATE X. FIGURES 40–41a. Male genitalia. 40a in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE X. FIGURES 40–41a. Male genitalia. 40a. Metarctia (Hebena) lateritia, neotype, RSA, Barberton Nature Reserve, GP 36.749 (ZSM); 40b. Idem, Kenya, Mount Kenya, GP 36.634 (RCRF); 40c. Idem, Angola, Benguela Province, GP 36.757 (RCRF); 40d. Idem, Tanzania, Province Arusha, GP 36.766 (RCRF); 40e. Idem, Uganda, Bwindi Impenetrable NP, GP 31.059 (RCRF); 41a. Metarctia (Hebena) rubra, lectotype, [South Africa], Cape G. Hope, GP BM ARCT 5581 (NHMUK).

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PLATE IX. FIGURES 39a–c in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE IX. FIGURES 39a–c. Terminology of male and female genitalia morphology. 39a. Male genitalia capsule; 39b. Phallus; 39c. Female genitalia. Abbreviations after Klots (1970): unc—uncus; tg—tegumen; gn—gnathos; val—valva; sl—sacculus; sa—saccus; vin—vinculum; jux—juxta; set—setae; t.pr—terminal process of valva; co—costa; c.pr—costal process of valva; ves—vesica; coe—coecum penis; du.ej—ductus ejaculatorius; pa.a—papilla analis; apo.po—apophysis posteriores; sin.v—sinus vaginalis; o.bu—ostium bursae; du.bu—ductus bursae; du.sml—ductus seminalis; sig.bu—signum; crp.bu—corpus bursae

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PLATE VIII. Images of live adults and larvae. FIGURES 33–34. Metarctia (Hebena) lateritia, adult male, Uganda, Semliki NP (33–34. © Steffen Schellhorn). 35–36. Metarctia (Hebena) rubra, larva, last instar, RSA, Waterberg Mts., reared; 37. Idem, adult male, same data as in 35; 38. Idem, adult female, same data as in 35 (35–38. © Ralf Fiebig). in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE VIII. Images of live adults and larvae. FIGURES 33–34. Metarctia (Hebena) lateritia, adult male, Uganda, Semliki NP (33–34. © Steffen Schellhorn). 35–36. Metarctia (Hebena) rubra, larva, last instar, RSA, Waterberg Mts., reared; 37. Idem, adult male, same data as in 35; 38. Idem, adult female, same data as in 35 (35–38. © Ralf Fiebig).

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PLATE VI. FIGURES 10a–f in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE VI. FIGURES 10a–f. Metarctia (Hebena) henrardi. Adults. 10a. Holotype ♂, [DRC] Terr. Kanda-Kanda, Gandajika, GP P319 (RMCA); 10b. Underside of 10a; 10c. Paratype of Metarctia subincarnata Kiriakoff, 1954 ♂, [DRC] Congo belge, Lupweji (RBINS); 10d. ♂, Zambia, Mwinilunga, GP 26.897 (MWW); 10e. ♂, Zambia, Hillwood, Ikelenge, BC ZSM Lep 113369, GP AV 4365 (ANHRT); 10f. ♀, same data as in 10e, BC ZSM Lep 113333, GP AV6410 (ANHRT); FIGURES 11a–f. Metarctia (Hebena) lukaszi sp.n. Adults. 11a. Holotype ♂, Angola, Benguela Province, rd. Ganda—Ndende, BC ZSM Lep 111901, GP 28.685 (RCRF/MWW); 11b. Underside of 11a; 11c. Paratype ♀, same data as in 11a, GP 28.690 (MWW); 11d. Paratype ♂, Angola, Huambo Province, GP 28.686 (MWW); 11e. Paratype ♂, same data as 11a, GP RF 659.2021 (RCRF); 11f. Paratype ♀, same data as in 11a; BC ZSM Lep 111902, GP 36.770 (RCRF); FIGURES 12a–b. Metarctia (Hebena) venustissima. Adult. 12a. Holotype ♂, [DRC] Katanga, Kolwezi (RMCA); 12b. Underside of 12a. Scale bar 10 mm.

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PLATE XIX. FIGURES 79–84. Habitat photos. Figs 79–82 in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE XIX. FIGURES 79–84. Habitat photos. Figs 79–82. Habitat of Metarctia (Hebena) lateritia. 79–80. Uganda western, Bwindi Impenetrable NP, Cuckooland Lodge, 1700 m; 81. Uganda, Kibale Forest, a broad-leafed rainforest at a medium altitude; 82. Kenya, Mount Kenya (W), Naro Moru River Lodge, 1950 m, with the Mt. Kenya in the background (79–82. © R. Fiebig); 83. Habitat of Metarctia (Hebena) smithi sp.n., type locality, Zambia, Jiwundu Swamp; 84. Habitat of Metarctia (Hebena) smithi sp.n. and Metarctia (Hebena) henrardi, Zambia, near Kapishya Hot Springs (83–84. © ANHRT).

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PLATE IV. FIGURES 4c–e in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE IV. FIGURES 4c–e. Metarctia (Hebena) kelleni. Adults. 4c. Holotype of Metarctia titan Talbot, 1929 ♂, [Angola] Upper Cubango-Cunene Watershed (NHMUK); 4d. Paratype of Metarctia titan ♂, same data as in 4c, GP LG 4701 (NHMUK); 4e. ♂, Gabun, Atembe, GP RF 616.2021 (SMNS); FIGURES 5a–c. Metarctia (Hebena) elleni sp.n. Adults. 5a. Holotype ♂, Tanzania, Rukwa Region, Mbizi Mts., BC ZSM Lep 104704, GP 30.955 (RCRF/ZSM); 5b. Paratype ♂, same data as in 5a, BC ZSM Lep 104716 (RCRF); 5c. Underside of 5a; FIGURES 6a–c. Metarctia (Hebena) brigitta sp.n. Adults. 6a. Holotype ♂, Zimbabwe, Nyanga Nat.Park, BC ZSM Lep 111876, GP RF 294.2019 (RCRF/MfN); 6b. Underside of 6a; 6c. Paratype ♀, same data as in 6a, GP RF 283.2019, BC ZSM Lep 111873 (RCRF). Scale bar 10 mm.

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PLATE VII. FIGURES 13–22. Adults male head. 13a in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE VII. FIGURES 13–22. Adults male head. 13a. Metarctia (Hebena) lateritia, RSA; 13b. Idem, Tanzania; 13c. Idem, Angola; 14. M. (H.) rubra; 15. M. (H.) manfredi sp.n.; 16. M. (H.). kelleni; 17. M. (H.) elleni sp.n.; 18. M. (H.) brigitta sp.n.; 19. M. (H.) smithi sp.n.; 20. M. (H.) smithi transvallesiana ssp.n.; 21. M. (H.) haraldsulaki sp.n.; 22. M. (H.) lukaszi sp.n.; FIGURES 23–32. Adults male midleg. 23a. Metarctia (Hebena) lateritia, RSA; 23b. Idem, Tanzania; 23c. Idem, Angola; 24. M. (H.) rubra; 25. M. (H.) manfredi sp.n.; 26. M. (H.) kelleni; 27. M. (H.) elleni sp.n.; 28. M. (H.) brigitta sp.n.; 29. M. (H.) smithi sp.n.; 30. M. (H.) smithi transvallesiana ssp.n.; 31. M. (H.) haraldsulaki sp.n.; 32. M. (H.) lukaszi sp.n.

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PLATE III. Figures 2i–l in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE III. Figures 2i–l. Metarctia (Hebena) rubra. Adults. 2i. ♂, RSA, Prov. Free State, vic. Clarens, reared, BC ZSM Lep 111915 (RCMS); 2j. ♀, RSA, Provinc Limpopo, Waterberg Mts., reared (RCRF); 2k. ♂, same data as in 2j (RCRF); 2l. ♂, same data as in 2j (RCRF); FIGURES 3a–c. Metarctia (Hebena) manfredi sp.n. Adults. 3a. Holotype ♂, Kenya, Kibwezi, GP 36.549 (RCRF/ZSM); 3b. Underside of 3a; 3c. Paratype ♀, Kenya, Kilimandscharo, GP 36.548 (RCRF); FIGURES 4a–b. Metarctia (Hebena) kelleni. Adults. 4a. Lectotype ♂, [Angola] Humpata (RMNH); 4b. Paralectotype ♂, same data as in 4a (RMNH). Scale bar 10 mm.

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PLATE II. FIGURES 2a–h in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE II. FIGURES 2a–h. Metarctia (Hebena) rubra. Adults. 2a. Lectotype ♂, [South Africa], Cape G. Hope, GP BM Arct. 5581 (NHMUK); 2b. Holotype of Hexaneura cinnamomea Wallengren, 1896 ♂, [South Africa], Caffraria (SNHM); 2c. ♂, RSA, Provinc Limpopo, Waterberg mountain, reared (RCRF); 2d. Holotype of Metarctia lateritia ab.lateritiola Strand, 1917 ♂, [RSA] Natal (NHMUK); 2e. Undersite of 2c; 2f. ♀, same data as in 2c (RCRF); 2g. ♂, RSA, Drakensberg Mts., GP 36.555, BC ZSM Lep104705 (RCRF); 2h. Underside of 2g. Scale bar 10 mm.

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PLATE V. FIGURE 6d in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE V. FIGURE 6d. Metarctia (Hebena) brigitta sp.n. Adult. 6d. Paratype ♂, Zimbabwe, Nyanga NP, GP RF 282.2019 (RCRF); FIGURES 7a–e. Metarctia (Hebena) smithi sp.n. Adults. 7a. Holotype ♂, Zambia, Jiwundu Swamp, BC ZSM Lep 113348, GP AV 4368 (ANHRT); 7b. Paratype ♀, Zambia, Kapishya Hot Springs, BC ZSM Lep 113358, GP AV 6405 (ANHRT); 7c. Paratype ♂, same data as in 7b; BC ZSM Lep 113327 (ANHRT); 7d. Paratype ♂, Zambia, Mayukuyuku Kafue NP, BC ZSM Lep 111914, GP RF 565.2020 (RCMS); 7e. Paratype ♀, Zambia, Mwinilunga, GP 32.195 (MWW); FIGURES 8a–c. Metarctia (Hebena) smithi transvallesiana ssp.n. Adults. 8a. Holotype ♂, Tanzania, Katavi Region, BC ZSM Lep 111887, GP 36.754 (RCRF/ZSM); 8b. Undersite of 8a; 8c. Paratype ♀, Tanzania, Katavi Region, BC ZSM Lep 104707, GP 30.960 (RCRF); FIGURES 9a–f. Metarctia (Hebena) haraldsulaki sp.n. Adults. 9a. Holotype ♂, Angola, Cuanza Sul Province, BC ZSM Lep 111921, GP 36.789 (RCRF/ ZSM); 9b. Underside of 9a; 9c. Paratype ♀, Angola, Huambo Province, BC ZSM Lep 111922, GP 36.792 (RCRF); 9d. Paratype ♂, Angola, Bié Province, BC ZSM Lep 111890, GP RF 36.762 (RCRF); 9e. Paratype ♂, Angola, same site as 9c, BC ZSM Lep 111906, GP 28.683 (RCRF); 9f. Paratype ♂, same data as in 9c, GP RF 650.2021 (RCRF). Scale bar 10 mm.

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MAP 3 in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

MAP 3. Distribution of Metarctia (Hebena) smithi smithi, M. (H.) smithi transvallesiana and M. (H.) haraldsulaki

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PLATE I. FIGURES 1a–o in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE I. FIGURES 1a–o. Metarctia (Hebena) lateritia. Adults. 1a. figure of Automolis lateritia Herrich-Schäffer, 1855 (pl. 74 Fig. 274 in Herrich-Schäffer, (1855)), [South Africa], Cape; 1b. Neotype ♂, RSA, Prov. Mpumalanga, Barberton Nat. Res., BC ZSM Lep 111882, GP 36.749 (ZSM); 1c. Holotype of Hebena venosa Walker, 1856 ♀, [South Africa], Natal, GP LG 5496 (OUMNH); 1d. ♂, Uganda w., Semliki NP, BC ZSM Lep 104717, GP 36.673, reared (RCRF); 1e. Underside of 1b; 1f. ♀, same data as 1b, GP HL 1-2016 (RCRF); 1g. ♂, Kenya centr., Mount Kenya W, BC ZSM Lep 104720, GP 36.634 (RCRF); 1h. ♂, Uganda w., Bwindi Imepenetrable NP, BC ZSM Lep 111881, GP 31.059 (RCRF); 1i. ♀, Uganda w., Mburo Lake NP, BC ZSM Lep 104727, GP 36.599 (RCRF); 1j. ♂, Angola, Huila, GP 28.687 (MWW); 1k. ♂, Zambia, Kapishya Hot Springs, BC ZSM Lep 113322 (ANHRT); 1l. ♂, Angola, Benguela Prov., BC ZSM Lep 111896, GP 36.757 (RCRF); 1m. ♂, Tanzania, Katavi Region, BC ZSM Lep 104723, GP 30.956 (RCRF); 1n. ♂, Tanzania, Tanga Region, BC ZSM Lep 104724, GP 30.958 (RCRF); 1o. ♀, Tanzania, Uluguru Mts., BC ZSM Lep 114247, GP RF 2021.876 (RCRF). Scale bar 10 mm.

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MAP 2 in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

MAP 2. Distribution of Metarctia (Hebena) rubra, M. (H.) manfredi, M. (H.) kelleni, M. (H.) elleni and M. (H.) brigitta.

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PLATE XII. FIGURES 45–48a. Male genitalia. 45a in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE XII. FIGURES 45–48a. Male genitalia. 45a. Metarctia (Hebena) brigitta sp.n., holotype, Zimbabwe, Nyanga NP, GP RF 294.2019 (RCRF/MfN); 45b. Idem, paratype, same data as in 45a, GP RF 282.2019 (RCRF); 46a. Metarctia (Hebena) smithi sp.n., holotype, Zambia, Jiwundu Swamp, GP AV4368 (ANHRT); 46b. Idem, paratype, Kafue NP, GP RF 565.2020 (RCMS); 47. Metarctia (Hebena) smithi transvallesiana ssp.n., holotype, Tanzania, Katavi Region, GP 36.754 (RCRF/ZSM); 48a. Metarctia (Hebena) haraldsulaki sp.n., holotype, Angola, Cuanza Sul Province, GP 36.789 (RCRF/ZSM).

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PLATE XVII. FIGURES 67–72. Habitat photos. 67–68 in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

PLATE XVII. FIGURES 67–72. Habitat photos. 67–68. Habitat of Metarctia (Hebena) lukaszi sp.n.. 67. Type locality, Angola, Benguela Province, btw. Ganda—Ndende, 1210 m; 68. Angola, Huambo Province, ca. 3 km N Camava, 1447 m; 69–71. Habitat of Metarctia (Hebena) lateritia. 69. Angola, Benguela Province, ca. 15 km E Cutembo, 1047 m; 70. Angola, Huila Province, 17 km WNW Ndongwe, 1356 m; 71. Angola, Cuanza Sul Province, Chindalala, ca.10 km SW Conda, 905 m; 72. Habitat of Metarctia (Hebena) haraldsulaki sp.n., type locality, Angola, Cuanza Sul Province, ca. 80km E Quibala, 1352m (67–72. © S. Naumann)

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TEXT FIGURE 2b in Integrative taxonomic revision of the Metarctia Walker, 1855 subgenus Hebena Walker, 1856, with descriptions of six new species and one new subspecies (Lepidoptera: Erebidae: Arctiinae: Syntomini)

TEXT FIGURE 2b. Second part of DNA barcode tree of the Hebena subgenus based on Bayesian Inference analysis, including 116 Metarctia specimens and two outgroups (Arctia caja and Balacra pulchra). Posterior probability values are indicated above the branches, BIN URIs are shown on the right.

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FIGURE 8 in Incipient speciation in allopatric Etheostoma rupestre (Percidae: Etheostomatinae) lineages, with the description of three new subspecies

FIGURE 8. Male pigmentation of A) Etheostoma rupestre rupestre—47 mm SL, Cottondale Creek, County Road 32, Tuscaloosa, Alabama (photo by K. Boone), B) E. r. piersoni—61 mm SL, Chilatchee Creek, Highway 5, 1 mile north of Alberta, Alabama (photo by K. Boone), and C) E. r. uphapeense—45 mm SL, Opintlocco Creek, County Highway 20, ~3 miles East of Tuskegee, Alabama (Photo by B. Hilburn).

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