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Fig. 46. A in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 46. A. Canthon cobosi (Pereira & Martínez, 1960) stat. et comb. nov., holotype. B–C. Canthon machadoi (Martínez & Pereira, 1967) comb. nov. B. Paratype 2. C. Paratype 4.

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Fig. 43 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 43. The three Amazonian species of the furvus subgroup. A, D. Sylvicanthon furvus (Schmidt, 1920). A. Dorsal view. D. Ventral view. B, E. S. monnei sp. nov. B. Dorsal view. E. Ventral view. C, F. S. mayri sp. nov. C. Dorsal view. F. Ventral view.

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Fig. 44 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 44. Differences on the parameres of Sylvicanthon mayri sp. nov. and S. monnei sp. nov. (grayish zones represent original membraneous areas). A–B. S. mayri sp. nov. C–D. S. monnei sp. nov. Note that both branches of the apical bifurcation of the parameres of S. mayri sp. nov. are much more divergent than those of S. monnei sp. nov., which makes the internal angle between them more open in the first species (~110º) than in the second (~78º). In the same way, as the inferior branch is much more projected in S. mayri sp. nov. than in S. monnei sp. nov., the angle between the posterior region of that branch and the rest of the paramere is more open in S. monnei sp. nov. (~147º) than in S. mayri sp. nov. (~137º), species that seems to have a strong excavation at this point of the paramere (indicated by red arrow in A).

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Fig. 42 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 42. Sylvicanthon obscurus (Schmidt, 1920). A. Dorsal view of the purplish form. B. Dorsal view of the yellowish form. C. Ventral view.

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Fig. 41 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 41. Distribution of Sylvicanthon securus (Schmidt, 1920) comb. nov. and the four species of the furvus subgroup.

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Fig. 35 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 35. Variation on the pilosity at the centre of the hypomeral cavity among members of the bridarollii subgroup. A. Sylvicanthon seag sp. nov. B. S. attenboroughi sp. nov. C–D. S. bridarollii (Martínez, 1949). Note the first two species have the hypomeral cavity entirely glabrous or with setae limited to its periphery (centre always glabrous), whereas S. bridarollii possesses long and dense setae throughout the tegument of the hypomeral cavity.

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Fig. 32 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 32. Sylvicanthon bridarollii (Martínez, 1949). A–B. Individual from southern Bolivia. A. Dorsal view. B. Ventral view. C–D. Individual from Ecuador. C. Dorsal view. D. Ventral view.

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Fig. 36 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 36. Clinal morphological variation in S. bridarollii (Martínez, 1949). Note that, from south to north, the colouration of metafemora gradually fades from dark brown to reddish brown and orange with a light-brown base in populations of northern Peru, Ecuador, and Colombia, and that the internal margin of protibiae (indicated by the red arrows) becomes progressively much subtler. Other features varying along this north-south cline are the presence of coarse elongate punctures at the base of metafemora (present in southern populations and absent in the northern ones) and of a fine transverse line on the posterior edge of pronotum (absent in the southern populations). See the text for more details.

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Fig. 21 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 21. Sylvicanthon enkerlini (Martínez et al., 1964) comb. nov. A. Dorsal view. B. Ventral view. C. Holotype and its labels.

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Fig. 16 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 16. Abdominal lateral foveae of females (arrows). A. Sylvicanthon furvus (Schmidt, 1920). B. S. mayri sp. nov. C. S. obscurus (Schmidt, 1920). D. S. foveiventris (Schmidt, 1920).

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Fig. 15 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 15. Morphological variation and sexual dimorphism in the protibial spur of Sylvicanthon. A. S. enkerlini (Martínez et al., 1964) comb. nov., ♂. B. S. foveiventris (Schmidt, 1920), ♂. C. S. candezei (Harold, 1869), ♂. D. S. genieri sp. nov., ♂. E. S. aequinoctialis (Harold, 1868), ♂. F. S. aequinoctialis comb. nov., ♀. G. S. proseni (Martínez, 1949) stat. et comb. nov., ♂. H. S. proseni stat. et comb. nov., ♀. I. S. bridarollii (Martínez, 1949), ♂. J. S. seag sp. nov., ♂. K. S. edmondsi sp. nov., ♂. L. S. attenboroughi sp. nov., ♂. M. S. obscurus (Schmidt, 1920), ♂. N. S. furvus (Schmidt, 1920), ♂. O. S. securus (Schmidt, 1920) comb. nov., ♂.

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Fig. 28. A–B in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 28. A–B. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov. A. Dorsal view. B. Ventral view. C–D. S. proseni (Martínez, 1949) stat. et comb. nov. C. Dorsal view. D. Ventral view.

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Fig. 29 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 29. Type material of the aequinoctialis subgroup. A–B. Sylvicanthon aequinoctialis (Harold, 1868) comb. nov., type series. A. Lectotype and its labels. B. Paralectotype and its labels. C–E. S. proseni (Martínez, 1949) stat. et comb. nov., type series. C. Holotype and its labels. D. Paratype 1. E. Paratype 2.

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Fig. 40 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 40. Type material of three species of Sylvicanthon described by Schmidt (1920). A. S. secures (Schmidt, 1920) comb. nov. a. Lectotype. b. Lectotype's labels. B. S. obscurus (Schmidt, 1920). a. Lectotype. b. Lectotype's labels. c. Paralectotype 1. d. Paralectotype 3. e. Paralectotype 9. C. S. furvus (Schmidt, 1920). a. Lectotype. b. Lectotype's labels. c. Paralectotype 4. d. Paralectotype 6. e. Paralectotype 3. f. Paralectotype 1.

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Fig. 13 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 13. Morphological variation in the metafemora of Sylvicanthon Halffter & Martínez, 1977. A. S. foveiventris (Schmidt, 1920). Note the presence of coarse elongate punctures at the base of the metafemur (arrow) and the tegument without microsculpture. B. S. obscurus (Schmidt, 1920). Observe the tegument entirely covered by coarse non-elongate punctures. C. S. monnei sp. nov. D. S. mayri sp. nov. E. S. furvus (Schmidt, 1920). F. S. edmondsi sp. nov. G. S. bridarollii (Martínez, 1949) (southern populations). H. S. bridarollii (Martínez, 1949) (northern populations). Note the differences in colouration and in the presence of coarse punctation at the base of metafemur in the two ends of the clinal variation observed in S. bridarollii.

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Fig. 38. A–B in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 38. A–B. Sylvicanthon edmondsi sp. nov. A. Dorsal view. B. Ventral view. C–D. Sylvicanthon attenboroughi sp. nov. C. Dorsal view. D. Ventral view.

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Fig. 18 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 18. Aedeagus morphological diversity in the bridarollii subgroup. A. S. bridarollii (Martínez, 1949). B. S. seag sp. nov. (arrow points to the ventral notch). C. S. edmondsi sp. nov. (idem). D. S. attenboroughi sp. nov. (idem).

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Fig. 11 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 11. Morphological diversity in the protibiae of Sylvicanthon Halffter & Martínez, 1977. A. S. enkerlini (Martínez et al., 1964) comb. nov. B. S. aequinoctialis (Harold, 1869) comb. nov. C. S. securus (Schmidt, 1920) comb. nov. D. S. obscurus (Schmidt, 1920). E. S. furvus (Schmidt, 1920). F. S. bridarollii (Martínez, 1949) (southern populations). G. S. bridarollii (northern populations). H. S. attenboroughi sp. nov. I. S. edmondsi sp. nov. J. S. candezei (Harold, 1869). Note the variation related to the presence and degree of development of an expansion on the internal edge of protibiae and to the number, size and position of the external teeth.

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Fig. 10. Lateral view. A in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 10. Lateral view. A. Canthon (Francmonrosia) sp. B. Sylvicanthon enkerlini (Martínez et al., 1964) comb. nov. C. S. candezei (Harold, 1869).

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Fig. 20 in A monographic revision of the Neotropical dung beetle genus Sylvicanthon Halffter & Martínez, 1977 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), including a reappraisal of the taxonomic history of 'Canthon sensu lato'

Fig. 20. Combined distribution of the species of Sylvicanthon in the Neotropical region. Note that the genus occupies the four great areas of tropical humid forest of the American continent: Central American forests, the Choco of the South American northwest, the Amazon Rainforest and the Atlantic Forest. Following the hypothesis of Martínez et al. (1964) that the Amazon Basin was the ancestral area of the Sylvicanthon of the candezei group, three lineages have independently dispersed from there (arrows): that of the ancestors of S. aequinoctialis (Harold, 1868), which, from the Choco, invaded Central America at the pace of the tropical forests' gradual northwards advance, occurred during the Plio-Pleistocene, and those of the ancestors of respectively S. obscurus (Schmidt, 1920) and S. foveiventris (Schmidt, 1920), which reached the Atlantic Forest through the corridors of humid forests that bridged that biome to the Amazon forest during the wettest periods of the Neogene (see a more detailed discussion in the text).

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