FIGURE 4 in Mata Atlântica enchytraeids (Paraná, Brazil): A new genus, Xetadrilus gen. nov., with three new species, and four new species of Guaranidrilus Černosvitov (Enchytraeidae, Oligochaeta)

FIGURE 4. Guaranidrilus hoeferi sp. nov. A. Anterior 13 segments, lateral view. Shaded grey: Spermatheca (left), spermfilled seminal vesicle (right). Specimen submature with small egg. The seminal vesicle can extend over 2 more segments. B. Segments IV–VIII, dorsal view. C–F and H, K show details of A. C. Spermatheca, ectal duct and ectal dilatation of ampulla. D,E. Spermatheca, connecting tube. In D the epithelium is cylindrical; sperm is present as a central strand in the lumen. In E sperm is embedded in epithelium with undulating inner walls. F. Histological section (optical longitudinal section) from body surface to inner lumen of spermatheca; cl, coelom; ct, cuticle; eg, epidermal gland cells; lm, longitudinal muscles; rm, ring muscles; se, epithelium of spermathecal ental dilatation with inclusions of irregular outline; sp, sperm in lumen of ental dilatation. G. Coelomocytes. H. Detail of clitellum, lateral view. I. Preclitellar nephridium. J. Postclitellar nephridium. K. Detail of copulatory pad in XIII, optical longitudinal section. G from living specimens, rest from whole mounts.