Fig. 21 in The Basal Penguin (Aves: Sphenisciformes) Perudyptes Devriesi And A Phylogenetic Evaluation Of The Penguin Fossil Record

Fig. 21. Phylogeny of Sphenisciformes calibrated to the stratigraphic record. Ghost lineages (Norell, 1992) are minimized. Dotted lines extending the range of clade 7 are based on identification of fragmentary fossils of late Eocene age to this clade. We utilize age ranges for Spheniscus urbinai and Spheniscus megaramphus provided by Stucchi (2007). Most penguins from the La Meseta Formation occur in Telm 7 (34.2–36.1 Ma), but a few species have been reported from lower units (see Myrcha et al., 2002; Jadwiszczak, 2006a), accounting for the extended ranges of Delphinornis larseni, Anthropornis grandis, and Palaeeudyptes gunnari. A specimen from the Chilcatay Formation (see fig. 1) referred to Palaeospheniscus sp. by Acosta Hospitaleche and Stucchi (2005) is provisionally referred to Palaeospheniscus patagonicus (Ksepka, 2007), accounting for the longer range of this taxon. For taxa that are poorly constrained in age, the midpoint of possible ages is used (e.g., Marplesornis is placed at the late Miocene). Geographical locality for fossil taxa and breeding range of extant taxa are provided in parentheses following taxa names. Asterisks indicate taxa with extensive breeding ranges including oceanic islands (see Bertelli and Giannini [2005] for more data on these species). Abbreviations: AF, Africa; AN, Antarctica; AU, Australia; NZ, New Zealand and surrounding islands; SA, South America.