Fig. 2 in The systematics of Late Jurassic tyrannosauroid theropods from Europe and North America

Fig. 2. The phylogenetic relationships of tyrannosauroids, based on a revised analysis of the Brusatte et al. (2010) dataset. Details of the analysis are described in the text and the dataset is presented in SOM. The cladogram shown here is the single most parsimonious tree recovered by the analysis, with the wildcard taxon Aviatyrannis excluded (570 steps, CI = 0.640, RI = 0.835). Numbers next to nodes denote bootstrap percentages (based on 1000 replicates) and Bremer support. Note that Stokesosaurus clevelandi and "S." langhami (here referred to by its new genus name, Juratyrant) are not found as sister taxa, and therefore a monophyletic Stokesosaurus is not recovered. When Aviatyrannis is included in the analysis, the strict consensus of nine most parsimonious trees (not figured) shows identical and fully resolved relationships among Xiongguanlong and all more derived taxa. However, Stokesosaurus clevelandi, "S." langhami, and Eotyrannus, form a polytomy. This clade, in turn, is part of a large basal polytomy that also includes the Xiongguanlong + more derived clade, Dilong, Aviatyrannis, Guanlong, Kileskus, Proceratosaurus, and Sinotyrannus. On the figured cladogram, the following unambiguous synapomorphies support major clades, with character numbering following that in the character list of Brusatte et al. (2010) and SOM: all tyrannosauroids more derived than Dilong (33, 41, 49, 80, 180, 181, 196, 198, 221, 239, 241, 244, 257, 274, 281, 289, 290); the clade of S. clevelandi, Juratyrant, and Eotyrannus (258, 310, 311, 313); the clade of Juratyrant and Eotyrannus (no unambiguous synapomorphies).