Figure 2 from: Shi H, Liang H (2015) The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini). ZooKeys 536: 1-92. https://doi.org/10.3897/zookeys.536.5982

Figure 2 - Possible character evolution of the subgenus Circinatus based on one of the most parsimonious trees (tree I) obtained from the SW analysis. Solid circles represent apomorphies; empty circles represent parallelisms and reversals; reversals are marked with a slash; twice-reversals are marked with a double slash; multistate evolution is marked under the characters. The species groups are shown in different colors, as in Figure 1.