FIGURE 4. Polystichum lionelii. A. Rhizome scale. B. Petiole scale with denticulate margins. C in Polystichum lionelii (Dryopteridaceae), a new Andean species from northwestern Argentina

FIGURE 4. Polystichum lionelii. A. Rhizome scale. B. Petiole scale with denticulate margins. C. Linear-lanceolate scale of raquis with glandular-cell at the apex (detail). D. Filiform scales, twisted. E. Scale margin with 1–2-cellular glandular (arrowhead) and pluricellular hair. F. Detail of glandular hair on scale margin. G. Detail of surface scale with glandular hairs (arrowhead). H. Laminar tissue with filiform scales. I. Filiform scales. J. Spores. Polystichum pycnolepis. K. Laminar tissue with glandular hairs (adaxial side). L. laminar tissue with glandular hairs (abaxial side). A–J: Serapio 120 (holotype MCNS), K-L: Castillón 1496 (LIL 41248)