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Fig. 6 in Giant theropod dinosaurs from Asia and North America: Skulls of Tarbosaurus bataar and Tyrannosaurus rex compared
Fig. 6. Left lacrimal of Tarbosaurus bataar ZPAL MgD−I/4in lateral (A, B) and medial (C, D) views.
Fig. 4 in Giant theropod dinosaurs from Asia and North America: Skulls of Tarbosaurus bataar and Tyrannosaurus rex compared
Fig. 4. Left maxilla of Tarbosaurus bataar ZPAL MgD−I/4 in lateral (A, B) and medial (C, D) views.
Fig. 1 in Giant theropod dinosaurs from Asia and North America: Skulls of Tarbosaurus bataar and Tyrannosaurus rex compared
Fig. 1. Skull of Tarbosaurus bataar ZPAL MgD−I/4 in lateral (A) and dorsal (B) views.
Table 1.2 in One Hundred Years of Tyrannosaurus rex: The Skeletons
<p>Table 1.2. Twenty Most <i>Complete</i> <i>Tyrannosaurus</i> Rex Skeletons</p><table><tbody><tr><th><b>Specimen</b></th><th>No. of <b>Bones</b></th><th><b>Percentage</b> of Skeleton</th></tr></tbody><tbody><tr><th>Sue FMNH PR2081</th><td>219</td><td>73%</td></tr><tr><th>Stan BHI 3033</th><td>190</td><td>63%</td></tr><tr><th>MOR 555</th><td>146</td><td>49%</td></tr><tr><th>AMNH 5027</th><td>143</td><td>48%</td></tr><tr><th>Ollie *</th><td>124</td><td>41%</td></tr><tr><th>Scotty RSM 2523.8 *</th><td>120+</td><td>40%+</td></tr><tr><th>Samson (<i>Z-rex</i>)*</th><td>121</td><td>40%</td></tr><tr><th>Pecks rex MOR 980 *</th><td>120</td><td>40%</td></tr><tr><th>Ivan *</th><td>116</td><td>39%</td></tr><tr><th>Wyrex BHI 6230</th><td>114</td><td>38%</td></tr><tr><th>MOR 1125</th><td>111</td><td>37%</td></tr><tr><th>Thomas LACM 7509/10167 *</th><td>110+</td><td>37%+</td></tr><tr><th>Bucky TCM 2001.90.1</th><td>101</td><td>34%</td></tr><tr><th>Black Beauty RTMP.81.6.1</th><td>85</td><td>28%</td></tr><tr><th>SDSM 12047</th><td>82</td><td>27%</td></tr><tr><th>Duffy BHI 4100</th><td>79</td><td>26%</td></tr><tr><th>LACM 23844</th><td>74</td><td>25%</td></tr><tr><th>Tinker, SD, collected by Eatman and Ferrel*</th><td>73</td><td>24%</td></tr><tr><th>UCRC V1 *</th><td>60</td><td>20%</td></tr><tr><th>MOR 009</th><td>58</td><td>19%</td></tr></tbody></table>
Table 1.1 in One Hundred Years of Tyrannosaurus rex: The Skeletons
<p>Table 1.1. Summary of <i>Tyrannousaurus Rex</i> Skeletons <i>by Year</i> of <i>Excavation (</i>as of August 2006)</p><table><thead><tr><th><b>Location</b></th><th>Year Excavated</th><th><b>Specimen</b> No. <b>/</b> Name</th><th><b>Discoverer, Year</b></th><th><b>State/</b> Province</th><th>No. of <b>Elements</b></th><th><b>%</b> of <b>Skeleton</b></th><th>Skull <b>/ Parts</b></th><th>Sex <b>*</b></th><th>Current <b>Location</b></th></tr></thead><tbody><tr><th>1</th><td>1900</td><td>BMNH R7994</td><td>Brown, 1900</td><td>WY</td><td>40</td><td>13</td><td>Y</td><td>-</td><td>Natural History Museum, London, England</td></tr><tr><th>2</th><td>1902-1905</td><td>CM 9380</td><td>Brown, 1902</td><td>MT</td><td>34</td><td>11</td><td>Y</td><td>F</td><td>Carnegie Museum of Natural History, Pittsburgh, PA</td></tr><tr><th>3</th><td>1902</td><td>CM 1400</td><td>Peterson, 1902</td><td>WY</td><td>29</td><td>10</td><td>Y</td><td>?</td><td>Carnegie Museum of Natural History, Pittsburgh, PA</td></tr><tr><th>4</th><td>1908</td><td>AMNH 5027</td><td>Brown, 1908</td><td>MT</td><td>143</td><td>48</td><td>Y</td><td>M?</td><td>American Museum of Natural History, NY, NY</td></tr><tr><th>5</th><td>1967</td><td>MOR008</td><td>MacMannis, 1967</td><td>MT</td><td>46</td><td>15</td><td>Y</td><td>F?</td><td>Museum of the Rockies, Bozeman, MT</td></tr><tr><th>6</th><td>1967-1969</td><td>LACM 23844</td><td>Garbani, 1966</td><td>MT</td><td>74</td><td>25</td><td>Y</td><td>M</td><td>Natural History Museum of LA County, LA, CA</td></tr><tr><th>7</th><td>1967</td><td><i>LACM</i> 23845</td><td>Garbani. 1969</td><td>MT</td><td>37</td><td>12</td><td>Y</td><td>?</td><td>Natural History Museum of LA County, LA, CA</td></tr><tr><th>8</th><td>1981</td><td>SDSM 12047</td><td>Floden, 1980</td><td>SD</td><td>82</td><td>27</td><td>Y</td><td>M?</td><td>Museum of Geology, South Dakota School of Mines and Technology, Rapid City, SD</td></tr><tr><th>9</th><td>1981</td><td>RTMP 81.12.1</td><td>Sternberg, 1946</td><td>AB</td><td>49</td><td>16</td><td>Y</td><td>M</td><td>Royal Tyrrell Museum of Palaentology, Drumheller, AB</td></tr><tr><th>10</th><td>1981</td><td>RTMP 81.6.1 Black Beauty</td><td>Baker, 1980</td><td>AB</td><td>85</td><td>28</td><td>Y</td><td>F</td><td>Royal Tyrrell Museum of Palaentology, Drumheller, AB</td></tr><tr><th>11</th><td>1981</td><td>MOR 009 Hager rex</td><td>Hager, 1981</td><td>MT</td><td>58</td><td>19</td><td>N</td><td>?</td><td>Museum of the Rockies, Bozeman, MT</td></tr><tr><th>12</th><td>1983, 2003</td><td>NMMNH PR1081</td><td>Staton and LaPoint, 1982</td><td>NM</td><td>10</td><td>3</td><td>Y</td><td>?</td><td>New Mexico Museum of Natural History and Science, Albuquerque, NM</td></tr><tr><th>13</th><td>1990</td><td>MOR 555 Wankel <i>T.</i> rex</td><td>Wankel, 1988</td><td>MT</td><td>146</td><td>49</td><td>Y</td><td>M</td><td>Museum of the Rockies, Bozeman, MT</td></tr><tr><th>14</th><td>1990</td><td>FMNH PR2081 Sue</td><td>Hendrickson, 1990</td><td>SD</td><td>219</td><td>73</td><td>Y</td><td>f</td><td>Field Museum, Chicago, IL</td></tr><tr><th>15</th><td>1992, 1993, 2003</td><td>BHI 3033 Stan</td><td>Sacrison, 1987</td><td>SD</td><td>190</td><td>63</td><td>Y</td><td>M</td><td>Black Hills Institute of Geological Research, Hill City, SD</td></tr><tr><th>16</th><td>1992</td><td>Samson</td><td>Zimmerscheid, 1987</td><td>SD</td><td>121</td><td>40</td><td>Y</td><td>F</td><td>Private, Carnegie Museum of Natural History, Pittsburgh, PA</td></tr><tr><th>17</th><td>1992</td><td>DMNH 2827</td><td>Fickle, 1992</td><td>CO</td><td>10</td><td>3</td><td>N</td><td>7</td><td>Denver Museum of Nature & Science, Denver, CO</td></tr><tr><th>18</th><td>1993</td><td>Bowman</td><td>Pearson, 1992</td><td>ND</td><td>45+</td><td>15+</td><td>N</td><td>7</td><td><i>Pioneer</i> Trails Regional Museum, Bowman, ND</td></tr><tr><th>19</th><td>1993-1996, 2006</td><td>BHI 4100 Duffy</td><td>Sacrison, 1993</td><td>SD</td><td>79</td><td>26%</td><td>Y</td><td>7</td><td>Black Hills Institute of Geological Research, Hill <i>City</i>, SD</td></tr><tr><th>20</th><td>1993</td><td>UWGM 181</td><td>Pallen, 1993</td><td>MT</td><td>20</td><td>7</td><td>Y</td><td>7</td><td>Geological Museum, University of Wisconsin, Madison, Wl</td></tr><tr><th>21</th><td>1994-2001</td><td>RSM 2523.8 Scotty</td><td>Gebhardt, 1991</td><td>SK</td><td>120+</td><td>40+</td><td>Y</td><td>F</td><td><i>T.</i> rex Discovery Centre, Eastend, SK</td></tr><tr><th>22</th><td>1994-1995</td><td>BHI 6219007</td><td>Garstka, 1994</td><td>ND</td><td>12</td><td>2</td><td>Y</td><td>7</td><td>Various, incl. Black Hills Inst. Geol. Res., Hill <i>City</i>, SD</td></tr><tr><th>23</th><td>1995</td><td>BHI 6239 Steven</td><td>Sacrison, 1995</td><td>SD</td><td>15</td><td>5</td><td>N</td><td>F?</td><td>Black Hills Inst. Geol. Res., Hill City, SD</td></tr><tr><th>24</th><td>1995</td><td>LDP 977-2 Pete</td><td>Patchus, 1995</td><td>WY</td><td>35+</td><td>12+</td><td>N</td><td>7</td><td>Univ. New Orleans, New Orleans, LA</td></tr><tr><th>25</th><td>1995-1996</td><td>Barnum</td><td>Theisen, 1995</td><td>WY</td><td>47</td><td>16</td><td>N</td><td>7</td><td>Private</td></tr><tr><th>26</th><td>1996-1998</td><td>BHI 4182 Fox</td><td>Fox, 1994</td><td>SD</td><td>29</td><td>10</td><td>Y</td><td>F?</td><td>Black Hills Inst. Geol. Res., Hill <i>City</i>, SD</td></tr><tr><th>27</th><td>1997-2004</td><td>MOR 980 Pecks Rex</td><td>Tremblay, 1997</td><td>MT</td><td>120+</td><td>40+</td><td>Y</td><td>M</td><td>Ft. Peck Paleo. Station, Ft. Peck, MT</td></tr><tr><th>28</th><td>1997</td><td>Tinker</td><td>Eatman, 1997</td><td>SD</td><td>73</td><td>24</td><td>Y</td><td>7</td><td>Private</td></tr><tr><th>29</th><td>1998-1999</td><td>Ollie</td><td>Pfister, 1998</td><td>MT</td><td>124</td><td>41</td><td>Y</td><td>7</td><td>Great Plains Paleo., Madison, Wl</td></tr><tr><th>30</th><td>1998</td><td>Rex B</td><td>Alley, 1998</td><td>SD</td><td>24</td><td>8</td><td>Y</td><td>M</td><td>Black Hills Inst. Geol. Res., Hill City, SD</td></tr><tr><th>31</th><td>1999</td><td>Rex-C</td><td>Alley, 1999</td><td>SD</td><td>18</td><td>6</td><td>Y</td><td>F</td><td>Private, SD</td></tr><tr><th>32</th><td>2000</td><td>BHI 6248 E. D. Cope</td><td>Derlinger, 1999</td><td>SD</td><td>30</td><td>10</td><td>Y</td><td>?</td><td>Black Hills Inst. Geol. <i>Res</i>., Hill City, SD</td></tr><tr><th>33</th><td>2000</td><td>Monty</td><td>Landowner, 1999</td><td>WY</td><td>53</td><td>18</td><td>Y</td><td>?</td><td>Babiarz Inst. Paleon. Studies, Mesa, AZ</td></tr><tr><th>34</th><td>2000-2003</td><td>MOR 1125 B-rex</td><td>Harmon, 2000</td><td>MT</td><td>111</td><td>37</td><td>Y</td><td>F</td><td>Museum of the Rockies, Bozeman, MT</td></tr><tr><th>35</th><td>2000-2001</td><td>MOR 1126 <i>C-rex</i></td><td>Homer, 2000</td><td>MT</td><td>26</td><td>9</td><td>Y</td><td>?</td><td>Museum of the Rockies, Bozeman, <i>MT</i></td></tr><tr><th>36</th><td>2001</td><td>UCRC PV1</td><td>Zerbst, before 1950</td><td>WY</td><td>60+</td><td>20+</td><td>N</td><td>?</td><td>University of Chicago, Chicago, IL</td></tr><tr><th>37</th><td>2001</td><td>UMNH 110000</td><td>Difley and Saharatian, 2001</td><td>UT</td><td>26</td><td>9</td><td>Y</td><td>?</td><td>Utah Museum of Natural History, Salt Lake <i>City</i>, UT</td></tr><tr><th>38</th><td>2001-2002</td><td>TCM 2001.90.1 Bucky</td><td>Derflinger, 1998</td><td>SD</td><td>101</td><td>34</td><td>N</td><td>F</td><td>The Children's Museum, Indianapolis, IN</td></tr><tr><th>39</th><td>2001</td><td>MOR 1128 G-rex</td><td>Wilson, 2001</td><td>MT</td><td>23</td><td>7</td><td>N</td><td>F</td><td>Museum of the Rockies, Bozeman, MT</td></tr><tr><th>40</th><td>2001</td><td>MOR 1152 F-rex</td><td>Stewart, 2001</td><td>MT</td><td>25?</td><td>8?</td><td>N</td><td>?</td><td>Museum of the Rockies, Bozeman, MT</td></tr><tr><th>41</th><td>2001-2002</td><td>Otto</td><td>Pfister, 2001</td><td>MT</td><td>32</td><td>11</td><td>N</td><td>?</td><td>Great Plains Paleontology, Madison, Wl</td></tr><tr><th>42</th><td>2002-2003</td><td>MOR/USNM N-rex</td><td>Myrhvold, 2001</td><td>MT</td><td>40</td><td>13</td><td>Y</td><td>?</td><td>National Museum of Natural History, Washington, DC</td></tr><tr><th>43</th><td>2002-2004</td><td>BHI 6230 Wyrex</td><td>Wells and Wyrick, 2002</td><td>MT</td><td>114</td><td>38</td><td>Y</td><td>M</td><td>Black Hills Institute of Geological Research, Hill <i>City</i>, SD</td></tr><tr><th>44</th><td>2003-2005</td><td><i>LACM</i> 7509/10167 Thomas</td><td>Curry, 2003</td><td>MT</td><td>110+</td><td>37+</td><td>Y</td><td>?</td><td>Natural History Museum of Los Angles County, LA, CA</td></tr><tr><th>45</th><td>2004-2005</td><td>Wayne</td><td>Olson, 2004</td><td>ND</td><td>24</td><td>8</td><td>N</td><td>?</td><td>Private, Fargo, ND</td></tr><tr><th></th><td>2005</td><td>Ivan</td><td>Olson, 2005</td><td>SD</td><td>116</td><td>39</td><td>N</td><td>?</td><td>Private, Fargo, ND</td></tr></tbody></table><p>Note—See text and Figure 1.1.</p><p>* <i>Sex</i>: M, gracile morph; F, robust morph. <i>See</i> P. Larson (this volume).</p>
Tyrannosaurus (Daspletosaurus) torosus type NMC 8506 and AMNH 5438 in Predatory Dinosaurs of the World
Tyrannosaurus (Daspletosaurus) torosus type NMC 8506 and AMNH 5438
Figure 1 in Tyrannosaurus was not a fast runner
Figure 1 Explanation of freebody diagram analysis of body segments. Skeletal illustration modified from ref. 6. The initial Tyrannosaurus model (Trex_1) is shown in right lateral view and (x, y)coordinate space, with the origin located at the toe joint. Pelvic pitch, hip, knee, ankle and toe joint angle definitions are shown (see Supplementary Information). One of the angles is redundant (four angles suffice). The ground reaction force (GRF) is vertical (typical for midstance12–14) and acts at a distance R from the toe joints. The body segment weights (Wb, Wt, Ws and Wm for the trunk, thigh, shank and metatarsus) are also shown. A freebody diagram was used to calculate the internal moments about each joint. For example, about the knee joint (see inset), the moment that knee extensor muscles must generate (Mk) is equal to the ankle contact force (Fa) times its respective moment arm, plus the gravitational moment of the shank segment and the ankle moment (Ma).
FIGURE 4. A in A digitally-rendered endocast for Tyrannosaurus rex
FIGURE 4. A, line interpretation of endocast of FMNH PR2081 (Tyrannosaurus rex) in dorsal view, showing full estimated extent of olfactory bulbs; B, transverse CT section (2 mm thick) through skull immediately dorsal to orbits, showing expanded olfactory bulbs anterior to endocranial cavity; C, diagram of skull showing location of section in D. Abbreviations: fm, foramen magnum; I, olfactory tract; olb, olfactory bulb; orb, orbit; se, sphenethmoid.
FIGURE 3 in A digitally-rendered endocast for Tyrannosaurus rex
FIGURE 3. Horizontal slices through the endocranial cavity of FMNH PR2081, Tyrannosaurus rex. Each slice is 0.5 mm thick, synthetically generated from 2 mm thick coronal slices by VoxBlast, ver. 2.2; spacing between slices is 5 mm, with the dorsalmost slice at A and ventralmost at F. Broad separation of the ophthalmic (V1) and maxillarymandibular (V2,3) branches of the trigeminal nerve can be seen. Abbreviations: asc, anterior semicircular canal; ep, epipterygoid;?fov, fenestra ovalis; pf, pituitary fossa; ps, pneumatic sinus; psc, posterior semicircular canal; res, external otic recess; tg, trigeminal ganglion; us, utricular sinus.
FIGURE 1 in A digitally-rendered endocast for Tyrannosaurus rex
FIGURE 1. Digital endocast for FMNH PR2081, Tyrannosaurus rex. A, stereopair of endocast in ventrolateral view; B, stereopair of endocast in dorsolateral view; C, D, line interpretations of endocast in ventrolateral (C) and dorsolateral (D) view. Cranial nerves indicated with Roman numerals; note separation between ophthalmic (V1) and maxillomandibular (V2,3) branches of the trigeminal nerve. The left facial (VII) and vestibular statoacoustic (VIII.v) are shown. Abbreviations: cblm, cerebellar region (cerebellum itself not visible); cbm, cerebrum; f, forebrain; fm, foramen magnum; fpr, cast of perilymphatic foramen; fr, mold of floccular recess; h, hindbrain; m, midbrain; mf, cast of metotic fissure; olb, olfactory bulb; orb, right orbit; pf, pituitary fossa; ssc, semicircular canals; us, utricular sinus; vcd, dorsal cerebral vein; vcm, middle cerebral vein; vs, vascular structure;? unknown structure ventral to right trigeminal ganglion.
FIGURE 3 in Maximum Bite Force and Prey Size of Tyrannosaurus rex and Their Relationships to the Inference of Feeding Behavior
FIGURE 3 Cranial morphology and jaw adductor musculature. Differences in the posterior width of the skull, relative to the tooth rows, may be constrained, together with tooth shape, by biting strategies such as crushing or penetration. The rather broadly-set tooth rows of Allosaurus fragilis (illustration modified after Madsen, 1976) appear to provide most theropods with a sufficiently concentrated bite force to facilitate lethal, penetrating bites. The expanded posterior part of the skull of Tyrannosaurus rex allowed for jaw adductor muscles with more cross-sectional area (illustration modified after Osborn, 1912). This suggests an atypically high bite force in this taxon, compared to most other theropods. The combination of a high bite force with comparatively broad, conical teeth implies adaptation for crushing the relatively greater mass of flesh and/or bones of large prey. This suggests that T. rex may have fed in significantly different ways from the majority of large theropods.
FIGURE 1 in Maximum Bite Force and Prey Size of Tyrannosaurus rex and Their Relationships to the Inference of Feeding Behavior
FIGURE 1 Effect of prey size on penetrating bites. Non-vital tissues of prey animals, such as integument and muscle, serve as barriers to predators seeking to inflict lethal damage on their prey. Assuming some minimum depth of penetration is required for a lethal bite, the depth of tissue through which a predator's teeth must penetrate increases linearly with increasing prey size. However, frictional resistance between flesh and teeth increases more rapidly than this. For a simple cone, frictional resistance increases with the square of the linear increase in depth of penetration. This results, for example, in four times the amount of frictional resistance to penetration that is twice as deep. A subconical tooth of Tyrannosaurus rex was modeled by computer and its engaged surface area was calculated for bites to 2 and 4 cm depth. In the latter case, the surface area of the tooth engaged is approximately three times that for a 2cmbite, due to the moderate lateral compression of T. rex teeth.
FIGURE 2 in Maximum Bite Force and Prey Size of Tyrannosaurus rex and Their Relationships to the Inference of Feeding Behavior
FIGURE 2 Skull morphology and crushing bites. Typical theropods have dentary teeth that are aligned just medial to the maxillary teeth, (a) This results in forces that are almost directly opposed when the teeth, exemplified here by those of Allosaurus fragilis, engage a food item, (b) In contrast, the dentary and maxillary teeth of Tyrannosaurus rex do not generate directly opposed forces in biting. Rather, the medially aligned dentaries tend to concentrate force near midline. In bilateral biting, the resulting stress on the prey is well-suited to crushing either bone or soft tissues.
FIGURE 99 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 99. FMNH PR2081, Tyrannosaurus rex. Right second metatarsal (MTII) in medial (A), lateral (B), dorsal (C), and ventral (D) view. Scale =10 cm. Photographs by J. Weinstein.
FIGURE 85 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 85. FMNH PR2081, Tyrannosaurus rex. Right humerus in dorsal (A), ventral (B), proximal (C), and distal (D) views. Closeups of the anterior (E) and posterior (F) surfaces of the proximal end are included. Scale = 5 cm. Abbreviations in Appendix 1. Photographs by J. Weinstein.
FIGURE 101 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 101. FMNH PR2081, Tyrannosaurus rex. Right fourth metatarsal (MTIV) in medial (A), lateral (B), dorsal (C), and ventral (D) view. Scale =10 cm. Photographs by J. Weinstein.
FIGURE 72 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 72. Putative proatlas arch or atlantal rib in tyrannosaurid theropods. A-D, FMNH PR2081, Tyrannosaurus rex. Dorsal (left) and ventral (right) views of putative right (A, C) and left (B, D) elements; scale = 5 cm. Specimen orientation would be appropriate for either the proatlas or rib interpretation. E, FMNH PR308 (either Albertosaurus or Daspletosaurus), left dorsolateral view of craniovertebral joint showing putative proatlas elements in articulation. Photographs by J. Weinstein.
FIGURE 91 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 91. FMNH PR2081, Tyrannosaurus rex. Left pelvic girdle articulated. Scale = 30 cm. Photograph by J. Weinstein.
FIGURE 23 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 23. FMNH PR2081, Tyrannosaurus rex. Isolated left postorbital in medial (A) and lateral (B) view. Scale = 5 cm.
FIGURE 76 in Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull
FIGURE 76. FMNH PR2081, Tyrannosaurus rex. Anterior view of head of left rib from p17. Scale = 5 cm. See Appendix 1 for abbreviations.
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