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Fig. 26 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 26. Frontal view of cephalothorax. A, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; B, Manarma johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; C, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; D, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia.

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Fig. 28 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 28. Chela of Contusarma. A, C. bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, C, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; D, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak. A, B, D, outer view; C, ventro-marginal view.

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Fig. 24. Overall habitus. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 24. Overall habitus. A, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; B, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; C, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; D, Manarma johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; E, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; F, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; G, Bresedium laevimanum, male (18.1 × 16.4 mm) (ZRC 1972.3.7.25), Sarawak, Borneo; H, Bresedium laevimanum, male (21.2 × 18.2 mm) (ZRC 1965.7.29.121), Johor, Malaysia.

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Fig. 21. A–G in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 21. A–G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H–L, Platychirarma buettikoferi, 1 male (10.7 × 8.8 mm) (SMF 25980), West Nimbe Cameroon. A, H, male pleon; B, I, left G1 (ventral view, denuded); C, left G1 (dorso-lateral view) (denuded); D, J, left G1 (dorsal view, denuded); E, K, left distal part of G1 (ventral view, denuded); G, L, left distal part of G1 (dorsal view, denuded); F, left distal part of G1 (dorso-lateral view) (denuded). Scales: A = 5.0 mm; B–D, H = 2.0 mm; E–G, I, J = 1.0 mm; K, L = 0.5 mm.

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Fig. 20. A, D–H in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 20. A, D–H, Cristarma eulimene, male, Tamatave, Madagascar; B, C, C. eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; I, J, C. ortmanni, male, Tuléar, Madagascar; K, C. ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique. A, I, carapace; B, left third maxilliped (denuded); C, anterior thoracic sternites 1–4; D, J, dorso-lateral view showing ridge on chela and dactylus; E, K, male pleon; F, left G1 (ventral view, denuded); G, left G1 (dorsal view, denuded); H, left distal part of G1 (ventral view, denuded). A, D–H, I, J, after Crosnier (1965: figs. 68–71, 73, 85, 107, 108). Scales: B, C = 1.0 mm; K = 5.0 mm.

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Fig. 23. A, E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 23. A, E, Contusarma bocourti, lectotype male (24.0 × 27.2 mm) (MNHN D-10965), Bangkok, Thailand; B, F, C. bocourti, male (25.3 × 23.3 mm) (ZRC 2000.0952), Bangkok, Thailand; C, C. bocourti, female (21.1 × 19.0 mm) (ZRC 2019.1114), Bangkok, Thailand; D, H, C. cheirogonum, neotype male (24.5 × 21.5 mm) (ZRC 1995.225), Bako National Park, Sarawak; G, C. cheirogonum, male (23.0 × 20.7 mm) (ZRC 2000.2018), Pulau Ubin, Singapore. A–D, overall habitus; E–H, anterior half of carapace.

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Fig. 48 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 48. Migmarma lustrum, new species, holotype male (15.7 × 13.9 mm) (ZRC 2010.0021), Irian Jaya, Indonesia. A, thoracic sternites 1–4; B, sternopleonal cavity; C, frontal view of cephalothorax; D, outer view of right chela; E, anterior thoracic sternum and pleon; C, D, left G1; E–G, distal part of left G1.

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Fig. 51 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 51. Migmarma lorentzi, new species, holotype male (15.8 × 14.0 mm) (NMBA 625IIa), Lorentz River, Irian Jaya, Indonesia. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, distal part of left G1 (mesial view); F, left G2. Scales = 0.5 mm.

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Fig. 54 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 54. Colour in life. A, Orisarma intermedium, Pingtung, Taiwan, specimen not collected [photograph: PKL Ng]; B, O. intermedium, female (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan [photograph: PKL Ng]; C, O. intermedium, male (23.2 × 20.0 mm) (ZRC 2019.0540), Hong Kong [photograph: Paul Ng]; D, O. sinense, Shanghai, China, specimen not collected [photograph: Liu Wenliang].

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Fig. 59 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 59. Phylogenetic relationships among selected taxa of Sesarmidae (91 OTUs), including all representatives of Chiromantes and Pseudosesarma previous to current revision of these genera. In addition, several species of Bresedium and Sesarmops are considered, as the revision also involves some of their former members. Topology and confidence values were determined by means of a phylogenetic reconstruction based on 1,096 basepairs of mtDNA (16S-tRNALeu-ND1), conducted with MrBayes, GTR+I+G as substitution model, and Parasesarma plicatum as outgroup (midpoint rooting). Posterior probabilities were determined after 4 million generations with a burn-in of 25%. All species so far classified within Chiromantes (without obvious anterolateral teeth) are shaded in blue, whereas those species that used to be classified in Pseudosesarma (carrying at least one pair of anterolateral teeth) are shaded in yellow.

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Fig. 52 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 52. Migmarma lorentzi, new species, paratype female (13.4 × 11.8 mm) (NMBA 625IIa), Lorentz River, Irian Jaya, Indonesia. A, overall habitus; B, vulvae.

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Fig. 17 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 17. Chela of species of Platychirarma buettikoferi. A, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia; B–D, male (12.6 × 10.4 mm) (ZRC 2015.0298), Cameroon. A, outer view; B, C, dorso-lateral view; D, ventro-marginal view.

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Fig. 16 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 16. Chela of species of Trapezarma angolense. A, B, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; C–F, male (35.2 × 28.6 mm) (ZRC 2015.0297), Cameroon. A, C, outer view; D, inner view; B, E, F, dorso-lateral view.

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Fig. 18. Male anterior thoracic sternum and pleon. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 18. Male anterior thoracic sternum and pleon. A, Danarma obtusifrons, male (19.7 × 14.7 mm) (ZRC 2002.0220), Oahu, Hawaii; B, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; C, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; D, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; E, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; F, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia.

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Fig. 12. A–E in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 12. A–E, Orisarma intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; F–J, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; K–O, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong. A, F, K, male pleons; B, G, L, left G1 (ventral view, denuded); C, H, M, left G1 (dorsal view, denuded); D, I, N, left distal part of G1 (ventral view, denuded); E, J, O, left distal part of G1 (dorsal view, denuded). Scales: A, F = 5.0 mm; B–E, G–J, K–M = 1.0 mm; N, O = 0.5 mm.

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Fig. 13. Overall habitus. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 13. Overall habitus. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Oahu, Hawaii; B, Danarma silus, holotype male (16.3 × 12.8 mm) (ZRC 2012.0787), Guam; C, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; D, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM), Greta Beach, Christmas Island; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia; I, Platychirarma buettikoferi, lectotype male (14.1 × 11.6 mm) (RMNH D 148), Liberia; J, Platychirarma buettikoferi, male (12.6 × 10.4 mm) (ZRC 2015.0298), Cameroon.

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Fig. 15. Chela. A, B in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 15. Chela. A, B, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028–002), Lanyu Island, Taiwan; C, D, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; E, F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique. A, C, E, outer view; B, inner view; D, F, dorso-lateral view showing pectinated ridge on palm.

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Fig. 14 in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 14. Frontal view of cephalothorax. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Hawaii; B, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM, ex ZRC 2009.0822), Christmas Island; C, D, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia.

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Fig. 6. Chela. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 6. Chela. A, Orisarma intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; B, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; C, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; D, O. intermedium, male (37.5 × 35.8 mm) (ZRC 2014.0265), Kumejima Island, Japan; E, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; F, O. sinense, male (30.7 × 26.7 mm) (ZRC 2010.0421), China; G, H, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China. A–G, outer view; H, inner view.

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Fig. 9. Male anterior thoracic sternum and pleon. A in Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species

Fig. 9. Male anterior thoracic sternum and pleon. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan; C, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; D, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; E, O. intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; F, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; G, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong.

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