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Fig. 8 Broeckinella hensoni n in Broeckinella Hensoni N. Sp., A New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran And A Revision Of The Genus Broeckinella Henson, 1948

Fig. 8 Broeckinella hensoni n. sp., upper Maastrichtian Tarbur Formation of the Naghan (a, d–e) and Mandegan (b–c, f) sections, Zagros Zone, SW Iran. a Oblique equatorial section. Note porcelaneous test of Cuvillerinella sp. below. b Detail showing fine subepidermal network (sn, below), followed by zone with radial partitions (rp) only and finally the undivided central chamber part with foramina (f) piercing the septa (s). c Oblique section cutting the initial planispiral part. d–e Oblique equatorial sections showing zones of different internal structure (see Text-Figure 3). f Detail from c showing initial planispirally coiled part with numerous chambers. Note also the fine subepidermal network and main partitions (right side). Thin-sections: 2NG 166 (a), Rt 67-1 (b), RT 72 (c, f), 2NG 118 (d), 2NG 172 (e). Scale bars 1.0 mm for a, c–e, 0.2 mm for b and f.

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Fig. 5 in Maastrichtian Larger Benthic Foraminifera From The Arabian Plate Sensu Lato: New Data From Somalia, Turkey, And Iran

Fig. 5 Cyclopsinella steinmanni (Munier-Chalmas) from the upper Maastrichtian of Somalia (Auradu Formation: a), Iran (Tarbur Formation: b-c, e-f), and C. steinmanni from the upper Santonian of France (d). a from Luger (2018, pl. 4, fig. 10 as Saudia sp.). b-c, e-f Naghan section. Note the aligned pillars (partly fusing laterally) in b and f, and sporadic rudimentary short rafter (r) in e. d from Gendrot (1964, pl. 1, fig. 10). Abbreviations: pi = pillar, r = rafter, s = septum.

Fig. 4 in Broeckinella Hensoni N. Sp., A New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran And A Revision Of The Genus Broeckinella Henson, 1948

Fig. 4 Internal structure of Broeckinella arabica Henson. a Drawing of a slightly oblique equatorial section (slightly modified from Henson, 1947, fig. 13c; see also Cherchi and Schroeder 1978, fig. 1C). 1 = subepidermal layer (network; exoskeleton), 2 = zone with transverse partitions only, 3 = undivided chamber interior (lacking endoskeleton); septa with foramina. b Tangential equatorial section of Broeckinella arabica showing the three zones of Henson (1948). Thin-section Rt 85, Mandegan section. c Tangential equatorial section showing main partitions aligned between successive chambers (detail from Fig. 6d). Thin-section NG 42-1, Naghan section.

Fig. 5 Broeckinella hensoni n in Broeckinella Hensoni N. Sp., A New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran And A Revision Of The Genus Broeckinella Henson, 1948

Fig. 5 Broeckinella hensoni n. sp. (a) and Broeckinella arabica Henson (b–d) from the upper Maastrichtian of the Tarbur Formation, Naghan section. a Foraminiferal wackestone with Broeckinella hensoni n. sp., thin-section 2NG 197. b Foraminiferal wackestone with Broeckinella arabica Henson, 2NG 128-1. c–d Details from b, showing a microspheric specimen. Scale bars in c and d = 1 mm.

Fig. 3 in Broeckinella Hensoni N. Sp., A New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran And A Revision Of The Genus Broeckinella Henson, 1948

Fig. 3 Naghan section, along the road from Naghan to Izeh with the type-level of Broeckinella hensoni n. sp. marked by an asterisk (left side).

Fig. 1 in Broeckinella Hensoni N. Sp., A New Larger Benthic Foraminifera From The Upper Maastrichtian Of Iran And A Revision Of The Genus Broeckinella Henson, 1948

Fig. 1 Peneroplis aragonensis (Peybernès) nov. comb. a Equatorial section. b Broken axial section (both from Peybernès, 1984, pl. 1, fig. 1 and 7, Late Albian of Spanish Pyrenees). c Detail from d showing fine surface ornamentation (striae). d Slightly oblique equatorial section. e Subaxial section showing fine striae of surface ornamentation (c-e from the Late Albian Barcenaciones Formation, Cantabria, N-Spain, leg M. Najarro).

Fig. 3 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 3 Dictyoconella complanata Henson from the Maastrichtian of Qatar (a-b) and Iran (c-e). a, isolated specimen, holotype (from Henson, 1948, pl. 6, fig. 2). Section planes 1-----1' and 2-----2' refer to b and d respectively. b, transverse section parallel to the median plane (from Henson, 1948, pl. 6, fig. 3, parataype). c, slightly oblique median section. Note the inclination of the chambers towards the apex. Thin section 2Ng 83. d-e, slightly oblique subaxial sections, perpendicular to the median plane (s = septum, f = foramen). Thin sections 2Ng 81, 2Ng 80-5. deuteroconch, eccentric position with twisted apex. Mi- Larger Benthic Foraminifera associated with dasycladale- crospheric embryo not discernible within a close-coiled an algae (Fig. 2). Gusicella minima occurs more common whorl. Adult chambers arched and rectilinear. Exoskelein the lower part of the Tarbur Fm. associated with Om- ton with several orders of horizontal and vertical partiphalocyclus and Loftusia (Wynd, 1965: Omphalocyclu- tions forming a delicate subepidermal network. The marclus-Loftusia assemblage zone), and other taxa such as ginal zone is separated from the central zone by a mar- Gyroconulina columelliforma Schroeder & Darmoian ginal trough. Central zone with pillars alternating be- (Fig. 2c-d). D. complanata is restricted to the upper part tween subsequent chambers. Foramina multiple in the of the Tarbur Formation in a more inner platform setting central zone and with straight arrangement between (compared to the occurrence of G. minima). Here it is chambers; marginal foramina are present. Wall finely associated with taxa such as Loftusia div. sp., Dicyclina agglutinated, thin, consisting of an epiderm and a delicate schlumbergeri Munier-Chalmas, Tarburina zagrosiana sub-epidermal cellular layer. Schlagintweit & Rashidi and Pseudonummoloculina Comparisons: Morphologically, Dictyoconella is unique kalantarii Schlagintweit & Rashidi (Fig. 2a-b). among all other Orbitolinidae due to its strongly compressed (flattened) test. This makes a detailed comparison SYSTEMATICS to the other Upper Cretaceous genera of the Orbitolinidae superfluous. Douglass (1960, p. 256) stated that "it is Phylum Foraminifera d'Orbigny, 1826 likely" that Dictyoconella "is merely a variant of the ge- Class Globothalamea Pawlowski et al., 2013 nus Iraqia". This assumption must be rejected because of Order Loftusiida Kaminski and Mikhalevich in Kamin- the difference in the central zone: reticulate in Iraqia (e.g. ski, 2004 Moullade, 1965) and pillaroid in Dictyoconella. It is Suborder Loftusiina Kaminski and Mikhalevich in Ka- worth noting at this point that the incorrect view of minski, 2004 Douglass was also approved by BouDagher-Fadel and Superfamily Orbitolinoidea Martin, 1890 Price (2009, p. 7) who included Dictyoconella in a group Family Orbitolinidae Martin, 1890 of "Orbitolinids with radial partitions that became zig- Subfamily Dictyoconinae Moullade, 1965 zagged, thickening and fusing centrally. Last but not Genus Dictyoconella Henson, 1948 emended herein least, neither Dictyononella complanata nor "D." minima Type-species: Dictyoconella complanata Henson, 1948. were treated in the revision of the Orbitolinidae by Cruz- Holotype P.35832 in Henson (1948), in repository at the Abad (2018) although the genus was considered valid by Natural History Museum London. Loeblich and Tappan (1987). Accessible from the origi- Diagnosis: Large-sized, laterally compressed test, flabel- nal samples of Qatar deposited at the Natural History liform. The test displays a distinct asymmetry, with one Museum in London (see pictures at nhm.ac.uk), the specside (with the eccentric embryo) that is wider (with re- imens were labeled Dictyoconella (Dictyoconus) comspect to the central axis) and presents a concave outer planata by Henson. In fact, Dictyconella is best described periphery in equatorial section. Test base distinctly con- as a distinctly compressed (flattened) Dictyoconus vex. Megalospheric embryo simple, biloculine with ellip- Blanckenhorn. Two specimens of Dictyoconella comsoidal protoconch and hemispherical to sickle-shaped planata were interpreted as microspheric specimens of

Fig. 2 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 2 Microfacies with Dictyoconella complanata Henson (a-b) and Gusicella minima (Henson) gen. et comb. nov. (c- d) from the Maastrichtian Tarbur Formation of the Naghan (a-b) and Mandegan sections (c-d), SW Iran (Zagros zone). Abbreviations: D = Dictyoconella, Di = Dicyclina, G = Gyroconulina, Gu = Gusicella, L = Loftusia, O = Omphalocyclus, P = Pseudonummoloculina, Ps = Pseudocymopolia, T = Tarburina. Thin sections: 2Ng 81 (a), 2Ng 81-4 (b), Rt 109 (c), Rt 105 (d).

Fig. 9 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l).

Fig. 6 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 6 Comparison of Gusicella gen. nov., Carinoconus Cherchi & Schroeder, and Dictyoconus Blanckenhorn.

Fig. 1 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 1 Map of the Persian Gulf area showing the location of Dukhan Well in Qatar, the type-locality of Dictyoconella complanata Henson and Gusicella minima (Henson) gen. et comb. nov. (from free maps, https://d-maps.com).

Fig. 5 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 5 Dictyoconus bakhtiari Schlagintweit et al. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section). Scale bar = 0.5 mm.

Fig. 4 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 4 Dictyoconella complanata Henson from the Maastrichtian of the Tarbur Formation of Iran (Naghan section, a-i, k) and the Simsima Formation of Qatar (j). a, d, oblique sections perpendicular to plane of compression. Note the inclination of the apical-juvenile part expressed by the parallel-sectioning of septum (s) and foramina (f) in d. b, oblique section passing through the initial part of a microspheric specimen. c, f, i-k, oblique sections in the plane of compression (j, paratype from Henson, 1948, pl. 10, fig. 14). Detail from i shows the fine subepidermal cellular network, marginal foramina and undivided marginal trough. e, g-h, oblique sections passing through the initial part of megalospheric specimens. Thin sections: 2Ng 81- 2 (a, d), Ng 76 (b), 2Ng 81-3 (c), Ng 85 (e), 2Ng 87-2 (f, i), 2Ng 80-1 (g), 2Ng 85-4 (h), 2Ng 81-1 (k). Abbreviations: b = beam, deu = deuteroconch, f = foramen m.f. = marginal foramen, m.t. = marginal trough, pi = pillar, pr = proloculus, T = Tarburina zagrosiana Schlagintweit et al. (in k).

Fig. 8 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p).

Fig. 7 in Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)

Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l).

Fig. 3 in Agglutinated Conical Foraminifera (Orbitolinidae, Coskinolinidae) From The Upper Cretaceous (Campanian) Of Greece, With Description Of Paracoskinolina Klokovaensis N. Sp.

Fig. 3 Upper Cretaceous (middle Campanian) microfacies of sample GKL 39. Pa = Paracoskinolina klokovaensis n. sp., Pal = Paleodictyoconus senonicus Moullade & Viallard, Di = Dicyclina schlumbergeri Munier-Chalmas.

Fig. 6 in Agglutinated Conical Foraminifera (Orbitolinidae, Coskinolinidae) From The Upper Cretaceous (Campanian) Of Greece, With Description Of Paracoskinolina Klokovaensis N. Sp.

Fig. 6 Comparison of Paracoskinolina species with one horizontal partition in the marginal zone: a P. klokovaensis sp. nov., b P. reicheli Guillaume, 1956. c P. arcuata (Arnaud-Vanneau 1976). D = test diameter, H = test height, n = number of chambers per last 0.5 mm axial length, h = chamber height (all data except D/H in mm). * measured from original illustrations. Stratigraphy of the two Lower Cretaceous species according to Clavel et al. (2014).

Fig. 2 in Agglutinated Conical Foraminifera (Orbitolinidae, Coskinolinidae) From The Upper Cretaceous (Campanian) Of Greece, With Description Of Paracoskinolina Klokovaensis N. Sp.

Fig. 2 Upper Cretaceous successions of Klokova Mountain, SW Greece (see Fleury, 1970, fig. 1, for details) with the distribution of some benthic foraminifers (A-B: Senalveolina aubouini Fleury, sample GKL 43; C-D Accordiella conica Farinacci, sample GKL 43x, without scale). Note: the lower/middle Campanian boundary is approximately placed at the LAD of K. tergestina (Frijia et al., 2015, fig. 15). For microfacies of sample GKL 39 see Figure 3.

Fig. 1 in Agglutinated Conical Foraminifera (Orbitolinidae, Coskinolinidae) From The Upper Cretaceous (Campanian) Of Greece, With Description Of Paracoskinolina Klokovaensis N. Sp.

Fig. 1 Geographic sketch map showing the location of Klokova mountains on the opposite side of the Gulf of Corinth facing Patra. This is the type-locality of Paracoskinolina klokovaensis n. sp.

Fig. 4 in Agglutinated Conical Foraminifera (Orbitolinidae, Coskinolinidae) From The Upper Cretaceous (Campanian) Of Greece, With Description Of Paracoskinolina Klokovaensis N. Sp.

Fig. 4 Coskinolinidae (a, d, g, j) and Orbitolinidae (b-c, e-f, h-i, k-l) from the Campanian of Klokova Mountain SW Greece (a-f, h-l) and Brač Island, Croatia (g). Lepinoconus chiocchinii Cruz-Abad et al. a Oblique section. d Broken transverse section. g, j Subaxial sections. Paleodictyoconus senonicus Moullade & Viallard b, e Oblique transverse sections. c, f Subaxial sections. Calvezicionus lecalvezae Caus & Cornella. h, k Oblique sections. i Oblique section passing the biloculine, slightly inclined megalospheric embryo. Dictyoconella complanata Henson. l Oblique section in the plane of compression. Abbreviations: b = beam, de = deuteroconch, f = foramen, p = protoconch, pi = pillar, s = septum. Thin sections: GKL 43 (a, j, i, l), GKL 43x (d, k), GKL 39 (b-c, e, h), GKL 39x (f).