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FIGURE 36 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 36. Buthacus arenicola (Simon, 1885), ♂ (AMNH), dorsal (A) and (B) ventral aspects. Scale bar = 1 cm.
FIGURE 38 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 38. Buthacus arenicola (Simon, 1885), ♂ (AMNH), dextral pedipalp chela, dorsal (A) and retrolateral (B) aspects. Scale bar = 2 mm.
FIGURE 40 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 40. Buthacus leptochelys (Ehrenberg, 1829), ♂ (HUJ INVSC 859), dextral pedipalp femur (A, B) and patella (C, E), prolateral (A, C), dorsal (B, D), and retrolateral (E) aspects. Scale bar = 2 mm.
FIGURE 37 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 37. Buthacus arenicola (Simon, 1885), ♂ (AMNH), dextral pedipalp femur (A, B) and patella (C, E), prolateral (A, C), dorsal (B, D), and retrolateral (E) aspects. Scale bar = 2 mm.
FIGURE 50 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 50. Buthacus tadmorensis (Simon, 1892), stat. rev., dextral pedipalp chela, dorsal (A, C) and retrolateral (B, D) aspects. A, B. Lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912. C, D. Paralectotype ♀ (NHMW 2452). Scale bar = 2 mm.
FIGURE 33 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 33. Buthacus arava, sp. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. Paratype ♂ (HUJ INVSC 2559). C, D. Paratype ♀ (HUJ INVSC 2561). Scale bars = 1 cm.
FIGURE 34 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 34. Buthacus arava, sp. nov., ♂♂ paratypes (HUJ INVSC 2559, 3295), dextral pedipalp femur (A, B) and patella (C, E), prolateral (A, C), dorsal (B, D), and retrolateral (E) aspects. Scale bar = 2 mm.
FIGURE 32 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 32. Buthacus amitaii sp. nov., dextral pedipalp chela, dorsal (A, C) and retrolateral (B, D) aspects. A, B. Holotype ♂ (HUJ INVSC 3480). C, D. Paratype ♀ (HUJ INVSC 3205). Scale bar = 2 mm.
FIGURE 35 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 35. Buthacus arava, sp. nov., dextral pedipalp chela, dorsal (A, C) and retrolateral (B, D) aspects. A, B. Paratype ♂ (HUJ INVSC 3295). C, D. Paratype ♀ (HUJ INVSC 2561). Scale bar = 2 mm.
FIGURE 29. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 29. Buthacus Birula, 1908, telson, lateral aspect. A. Buthacus arenicola (Simon, 1885), ♂ (AMNH). B. Buthacus levyi, sp. nov., holotype ♂ (HUJ INVSC 2082). C. Buthacus arava, sp. nov., paratype ♂ (HUJ INVSC 2559). D. Buthacus leptochelys (Ehrenberg, 1829), ♂ (HUJ INVSC 859). E. Buthacus tadmorensis (Simon, 1892), stat. rev., lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912. F. Buthacus nitzani Levy et al., 1973, stat. nov., ♂ (HUJ INVSC 3227). G. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (HUJ INVSC 3302). H. Buthacus amitaii, sp. nov., holotype ♂ (HUJ INVSC 3480). Scale bars = 1 mm.
FIGURE 28. Buthacus Birula, 1908, metasomal segments I–V in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 28. Buthacus Birula, 1908, metasomal segments I–V and telson, ventral aspect. A. Buthacus levyi, sp. nov., paratype ♀ (HUJ INVSC 3274). B. Buthacus leptochelys (Ehrenberg, 1829), ♀ (HUJ INVSC 1807). C. Buthacus nitzani Levy et al., 1973, stat. nov., ♀ (HUJ INVSC 3226). D. Buthacus amitaii, sp. nov., paratype ♀ (HUJ INVSC 3205). Scale bars = 5 mm.
FIGURE 26. Buthacus Birula, 1908, metasomal segments I–V in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 26. Buthacus Birula, 1908, metasomal segments I–V and telson, lateral aspect. A. Buthacus levyi, sp. nov., paratype ♀ (HUJ INVSC 3274). B. Buthacus leptochelys (Ehrenberg, 1829), ♀ (HUJ INVSC 1807). C. Buthacus nitzani Levy et al., 1973, stat. nov., ♀ (HUJ INVSC 3226). D. Buthacus amitaii, sp. nov., paratype ♀ (HUJ INVSC 3205). Scale bars = 5 mm.
FIGURE 25. Buthacus Birula, 1908, metasomal segments I–V in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 25. Buthacus Birula, 1908, metasomal segments I–V and telson, lateral aspect. A. Buthacus arava, sp. nov., paratype ♀ (HUJ INVSC 2561). B. Buthacus tadmorensis (Simon, 1892), stat. rev., paralectotype ♀ (NHMW 2452) of Buthus pietschmanni Penther, 1912. C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♀ (HUJ INVSC 3318). D. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 5 mm.
FIGURE 18. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 18. Buthacus Birula, 1908, carapace, dorsal aspect. A, B. Buthacus levyi, sp. nov., A. Holotype ♂ (HUJ INVSC 2082), B. Paratype ♀ (HUJ INVSC 3274). C. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 2 mm.
FIGURE 24. Buthacus Birula, 1908, metasomal segments I–V in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 24. Buthacus Birula, 1908, metasomal segments I–V and telson, dorsal aspect. A. Buthacus levyi, sp. nov., paratype ♀ (HUJ INVSC 3274). B. Buthacus leptochelys (Ehrenberg, 1829), ♀ (HUJ INVSC 1807). C. Buthacus nitzani Levy et al., 1973, stat. nov., ♀ (HUJ INVSC 3226). D. Buthacus amitaii, sp. nov., paratype ♀ (HUJ INVSC 3205). Scale bars = 5 mm.
FIGURE 17. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 17. Buthacus Birula, 1908, carapace, dorsal aspect. A, B. Buthacus leptochelys (Ehrenberg, 1829), A. ♂ (HUJ INVSC 859), B. ♀ (HUJ INVSC 1807). C, D. Buthacus nitzani Levy et al., 1973, stat. nov., C. ♂ (HUJ INVSC 3227), D. ♀ (HUJ INVSC 3226). E, F. Buthacus amitaii, sp. nov., E. Holotype ♂ (HUJ INVSC 3480), F. Paratype ♀ (HUJ INVSC 3205). Scale bars = 2 mm.
FIGURE 19. Buthacus Birula, 1908 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 19. Buthacus Birula, 1908, sternum, genital opercula and pectines, ventral aspect. A, B. Buthacus arava, sp. nov., A. Paratype ♂ (HUJ INVSC 3287), B. Paratype ♀ (HUJ INVSC 2561). C, D. Buthacus tadmorensis (Simon, 1892), stat. rev., C. Lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912, D. Paralectotype ♀ (NHMW 2452). E, F. Buthacus yotvatensis Levy et al., 1973, stat. rev., E. ♂ (HUJ INVSC 3344), F. ♀ (HUJ INVSC 3318). Scale bars = 2 mm.
FIGURE 14 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 14. Phylogeny of Buthacus Birula, 1908, species, obtained by simultaneous phylogenetic analysis of 22 morphological characters and 2205 bp of DNA sequence from three mitochondrial (12S rDNA, 16S rDNA, and COI) and two nuclear (28S rDNA and ITS2) genes, for 104 samples (Cain et al., in press). Maximum likelihood tree with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles parallel derivations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (appendix 3). Bootstrap support values are indicated at nodes.
FIGURE 30 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 30. Buthacus amitaii, sp. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. Holotype ♂ (HUJ INVSC 3480). C, D. Paratype ♀ (HUJ INVSC 3205). Scale bars = 1 cm.
FIGURE 8 in Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C.L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia
FIGURE 8. Map of the Maghreb, plotting known locality records for Buthacus arenicola (Simon, 1885) in Algeria and Tunisia, based on material examined for the present study and the literature. White symbol denotes locality of lectotype.
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