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Figures 17 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 17 - Line drawings of spermatheca of Guatemalan Geocharidius species. A Geocharidius integripennis (GUATEMALA, Totonicapán, "Totonicapam"), holotype B Geocharidius erwini (GUATEMALA, Quiché, Los Encuentros), paratype C Geocharidius minimus (GUATEMALA, Sacatepéquez, Antigua), paratype D Geocharidius jalapensis (GUATEMALA, Jalapa, Mataquescuintla), paratype E Geocharidius balini (GUATEMALA, Suchitepéquez, Volcano Atitlán), paratype F Geocharidius longinoi (GUATEMALA, El Progreso, Cerro Pinalón), paratype. Scale = 0.05mm.

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zenodo28/100

Figure 22 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figure 22 - Map of southern Mexico, Guatemala and adjacent part of Honduras, showing positions of locality records for the species of Geocharidius: white diamond, Geocharidius andersoni; white circles (black point in a circle shows "terra typica" for the species), Geocharidius zullinii; white squares, Geocharidius vignatagliantii; white triangle, Geocharidius gimlii; black and white triangle, Geocharidius integripennis; green squares, Geocharidius longinoi; green circles, Geocharidius erwini; green flowers, Geocharidius minimus; green diamonds, Geocharidius balini; green triangle, Geocharidius jalapensis; yellow quadrangle, Geocharidius antigua; yellow diamond, Geocharidius lencanus; yellow flower, Geocharidius celaquensis; yellow triangles, Geocharidius disjunctus; yellow circles, Geocharidius comayaguanus. Physiographic features: CH, Chiapas Highlands; GC, Guatemalan Cordillera; HIH, Honduran Interior Highlands; MTZ, Motagua Fault Zone; SC, Sierra de los Cuchumatanes; SMC, Sierra Madre de Chiapas. Elevation scale bar is given in meters.

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Figures 15 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 15 - Schematic line drawings of median lobe and dorsal sclerites of internal sac of two previously described Guatemalan Geocharidius species. A Geocharidius integripennis (GUATEMALA, Totonicapán, "Totonicapam"), holotype B Geocharidius gimlii (GUATEMALA, Huehuetenango, San Juan Ixcoy), holotype, both left lateral aspect.

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Figures 11 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 11 - Line drawings of spermatheca of Mexican Geocharidius species. A Geocharidius zullinii (MEXICO, Chiapas, Reserva Huitepec) B Geocharidius vignatagliantii (MEXICO, Chiapas, Motozintla), paratype C Geocharidius andersoni (MEXICO, Chiapas, Cerro Huitepec), paratype. Legend: c – cornu; n – nodulus; sd – spermathecal duct; sg – spermathecal gland; sp – spermatheca. Scale = 0.05mm.

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Figures 10 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 10 - Line drawings of ring sclerite of Mexican Geocharidius species, male genitalia, dorsal aspect. A Geocharidius zullinii (MEXICO, Chiapas, Reserva Huitepec) B Geocharidius vignatagliantii (MEXICO, Chiapas, Motozintla), paratype C Geocharidius andersoni (MEXICO, Chiapas, Cerro Huitepec), holotype. Legend: hd – handle of ring sclerite. Scale = 0.1mm.

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Figures 21 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 21 - Line drawings of spermatheca of Guatemalan and Honduran Geocharidius species. A Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), paratype B Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), paratype C Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype D Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype E Geocharidius disjunctus (HONDURAS, Yoro, Pico Pijol National Park). Scale = 0.05mm.

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Figures 14 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 14 - Photographs of labels for type specimens of Geocharidius species. A Geocharidius gimlii, holotype B Geocharidius integripennis, lectotype C Geocharidius integripennis, paralectotype.

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Figures 20 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 20 - Line drawings of ring sclerite of Guatemalan and Honduran Geocharidius species, male genitalia, dorsal aspect. A Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), holotype B Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), holotype C Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype D Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype E Geocharidius disjunctus (HONDURAS, Francisco Morazán, La Tigra National Park), holotype. Scale = 0.1mm.

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Figures 19 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 19 - Line drawings of aedeagus of Guatemalan and Honduran Geocharidius species. A–D Geocharidius antigua (GUATEMALA, Sacatepéquez, Antigua), holotype: A median lobe with internal sac and dorsal sclerites, right lateral aspect B dorsal sclerite of median lobe, dorsal aspect C left paramere, left lateral aspect D right paramere, right lateral aspect E–G Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), holotype: E median lobe with internal sac and dorsal sclerites, right lateral aspect F left paramere, left lateral aspect G right paramere, right lateral aspect H–J Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype: H median lobe with internal sac and dorsal sclerites, right lateral aspect I left paramere, left lateral aspect J right paramere, right lateral aspect K–N Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype: K median lobe with internal sac and dorsal sclerites, right lateral aspect L variation in a shape of dorsal sclerite of internal sac, right lateral aspect M left paramere, left lateral aspect N right paramere, right lateral aspect O–P Geocharidius comayaguanus (HONDURAS, La Paz, Guajicuiro): variations in a shape of dorsal sclerite of internal sac, right lateral aspect Q–S Geocharidius disjunctus (HONDURAS, Francisco Morazán, La Tigra National Park), holotype: Q median lobe with internal sac and dorsal sclerites, right lateral aspect R left paramere, left lateral aspect S right paramere, right lateral aspect. T Geocharidius disjunctus (HONDURAS, Yoro, Pico Pijol National Park): shape of dorsal sclerite of median lobe, right lateral aspect. Scale = 0.05mm.

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Figures 3 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 3 - SEM illustrations of structural features of body parts of Geocharidius species. A–C prothorax, ventral aspect: A Geocharidius zullinii B Geocharidius erwini C Geocharidius comayaguanus. D–F pterothorax, ventral aspect: D Geocharidius jalapensis E Geocharidius minimus F Geocharidius comayaguanus. G–I abdominal ventrites 3-5: G Geocharidius minimus, male H Geocharidius jalapensis, male I Geocharidius jalapensis female. Legend: asts – abdominal sternal terminal seta; ipa –intercoxal process of abdominal ventrite 2; mscx – mesocoxa; msp – mesosternal process; mtcx – metacoxa; mtp – metasternal process; mtv – metaventrite; pas – prosternal ambulatory seta; pep – proepipleuron; pes – proepisternum; prcx – procoxa; ps – prosternum; psp – prosternal process; v3-v5 – abdominal ventrites 3-5. Scale = 0.05mm.

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Figures 5 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 5 - SEM illustrations of structural features of labial complex of Geocharidius species. A Geocharidius erwini B Geocharidius balini C Geocharidius lencanus D Geocharidius zullinii E Geocharidius longinoi F Geocharidius minimus. Legend: gsc – glossal sclerite; lms – lateral mental seta; lss – lateral submental seta; m – mentum; mss – medial submental seta; pms – paramedial mental seta; prss – primary basal submental seta; sm – submentum. Scale = 0.05mm.

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Figures 23 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 23 - Diagrams illustrating size variation for sympatric pairs of closely and remotely related species of the integripennis group at different geographical localities of Nuclear Central America. Closely related pairs (for shared characters see subchapters on Relationships in the text for corresponding species): A Chiapas Highlands, Mexico B Mataquescuintla, Guatemala C Celaque National Park, Honduras. Remotely related species: D Volcano Agua, Guatemala E Volcano Atitlan, Guatemala F Pico Pijol National Park, Honduras. Legend: black dot – median; box – 25–75% range of values; whiskers – non-outlier range of values; b – coefficient of regression.

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Figures 18 from: Sokolov I, Kavanaugh D (2014) The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61-118. https://doi.org/10.3897/zookeys.443.7880

Figures 18 - Digital images of habitus of Honduran Geocharidius species. A Geocharidius celaquensis (HONDURAS, Lempira, Celaque National Park), paratype B Geocharidius lencanus (HONDURAS, Lempira, Celaque National Park), paratype C Geocharidius comayaguanus (HONDURAS, Comayagua, Comayagua), paratype D Geocharidius disjunctus (HONDURAS, Francisco Morazán, La Tigra National Park), holotype. Scale = 0.5mm.

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Figure 3 from: Liu W, Fabrizi S, Bai M, Yang X, Ahrens D (2014) A taxonomic revision of the Neoserica (sensu lato) pilosula group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 440: 89-113. https://doi.org/10.3897/zookeys.440.8126

Figure 3 - A–D Neoserica lincangensis sp. n. (holotype) E–H Neoserica rangshuiensis sp. n. (holotype). A, E edeagus, left side lateral view C, G aedeagus, right side lateral view B, F parameres, dorsal view D, H habitus. Scale: 0.5 mm, habitus not to scale.

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Figure 2 from: Liu W, Fabrizi S, Bai M, Yang X, Ahrens D (2014) A taxonomic revision of the Neoserica (sensu lato) pilosula group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 440: 89-113. https://doi.org/10.3897/zookeys.440.8126

Figure 2 - A–D Neoserica tianeana sp. n. (holotype) E–H Neoserica shennongjiaensis sp. n. (holotype). A, E aedeagus, left side lateral view C, G aedeagus, right side lateral view B, F parameres, dorsal view D, H habitus. Scale: 0.5 mm, habitus not to scale.

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Figure 4 from: Liu W, Fabrizi S, Bai M, Yang X, Ahrens D (2014) A taxonomic revision of the Neoserica (sensu lato) pilosula group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 440: 89-113. https://doi.org/10.3897/zookeys.440.8126

Figure 4 - A–D Neoserica lushuiana sp. n. (holotype) E–H Neoserica emeishanensis sp. n. (holotype) I–L Neoserica curvipenis sp. n. (holotype). A, E, I aedeagus, left side lateral view C, G, K aedeagus, right side lateral view B, F, J parameres, dorsal view D, H, L habitus. Scale: 0.5 mm, habitus not to scale.

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Figure 1 from: Liu W, Fabrizi S, Bai M, Yang X, Ahrens D (2014) A taxonomic revision of the Neoserica (sensu lato) pilosula group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 440: 89-113. https://doi.org/10.3897/zookeys.440.8126

Figure 1 - A–D Neoserica pilosula Moser, 1915 (Lectotype) E–H Neoserica ludingensis sp. n. (holotype) I–L Neoserica weibaoshanica sp. n. (holotype). A, E, I aedeagus, left side lateral view C, G, K aedeagus, right side lateral view B, F, J parameres, dorsal view D, H, L habitus. Scale: 0.5 mm, habitus not to scale.

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Figure 5 from: Ahrens D, Liu W, Fabrizi S, Bai M, Yang X (2014) A taxonomic review of the Neoserica (sensu lato) abnormis group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 439: 27-82. https://doi.org/10.3897/zookeys.439.8055

Figure 5 - A–D Neoserica putaoana sp. n. (holotype) E–H Neoserica alloputaoana sp. n. (holotype) I–L Neoserica simplicissima sp. n. (holotype) A, E, I Aedeagus, left side lateral view C, G, K Aedeagus, right side lateral view B, F, J parameres, dorsal view D, H, L Habitus. Scale: 1 mm. Habitus not to scale.

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Figure 8 from: Ahrens D, Liu W, Fabrizi S, Bai M, Yang X (2014) A taxonomic review of the Neoserica (sensu lato) abnormis group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 439: 27-82. https://doi.org/10.3897/zookeys.439.8055

Figure 8 - Distribution of the species of the Neoserica (s. l.) abnormis group: Neoserica abnormis, Neoserica abnormoides, Neoserica allolaotica, Neoserica euyunnanica, Neoserica jiangxiensis, Neoserica kanphantensis, Neoserica namthaensis, Neoserica ponderosa, Neoserica putaoana, Neoserica thailandensis, Neoserica tonkinea, Neoserica yaoi.

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Figure 2 from: Ahrens D, Liu W, Fabrizi S, Bai M, Yang X (2014) A taxonomic review of the Neoserica (sensu lato) abnormis group (Coleoptera, Scarabaeidae, Sericini). ZooKeys 439: 27-82. https://doi.org/10.3897/zookeys.439.8055

Figure 2 - A–E Neoserica tonkinea sp. n. (holotype), F–I Neoserica taunggyiana sp. n. (holotype) J–N Neoserica euyunnanica sp. n. (holotype) A, F, J Aedeagus, left side lateral view D, H, M Aedeagus, right side lateral view B, G, K parameres, dorsal view C, L aedeagus, ventral view E, I, N Habitus. Scale: 1 mm. Habitus not to scale.

opencc-by-4.0Sep 2014View details →

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allen-brain-atlas
neuroscienceopenDocumentation, web resources, and API references are available online.
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Last verified 2026-04-30Open record

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dandi-nwb
electrophysiologyopenPublished Dandiset metadata and archive endpoints are available through the production DANDI API.
Last verified 2026-04-30Open record

International Brain Laboratory public data

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behavioral-neuroscienceopenPublic sessions can be searched and loaded from the IBL public data server through ONE.
Last verified 2026-04-29Open record

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openneuro
neuroscienceopenPublished datasets are available on demand over the internet.
Last verified 2026-04-29Open record

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