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Figure 22 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 22 Detail of the oropharyngeal denticles in Cephaloscyllium sufflans, SAIAB 6242, male, 800 mm TL. cb I–V, ceratobranchials I–V.

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Figure 19 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 19 Detail of the basihyal cartilage. A, Scyliorhinus haeckelii, UERJ 1691, male, 522 mm TL; B, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL. thf, thyroid foramen.

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Figure 21 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 21 Detail of the lateral processi rastriformis (ras) in Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL.

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Figure 17 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 17 Detail of jaws; lighter portion of Meckel's cartilage is less calcified portion. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. lrq, labial ridge of the quadrate process; mck, Meckel's cartilage; opq, orbital process of the palatoquadrate; pq, palatoquadrate.

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Figure 2 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 2 Nasoral region with lifted anterior nasal flap and exposed posterior nasal flap. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Cephaloscyllium isabella, USNM 320594, female, 390 mm TL; Poroderma pantherinum, SAIAB 34577, male, 640 mm TL. anf, anterior nasal flap; ful, flap on the upper lip margin; llf, lower labial furrow; mrd, mesonarial crest; nb, nasal barbel; pnf, posterior nasal flap; pog, postoral groove; ulf, upper labial furrow. Scale bar: 20 mm.

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Figure 16 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 16 Neurocranium; lateral view. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Figaro boardmani, CSIRO H989-5, female, 465 mm TL; C, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. enc, external nasal cartilage; hf, hyomandibular facet; lrc, lateral rostral cartilage; mrc, medial rostral cartilage; nc, nasal capsule; pog, postorbital groove.

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Figure 18 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 18 Detail of the articular region of the quadratomandibular joint of Meckel's cartilage. A, Cephaloscyllium sufflans, SAIAB 6242, male, 800 mm TL; Galeus antillensis, UF 77853, female, 370 mm TL; C, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. lac, labial condyle; lic, lingual condyle; mfa, mandibular facet.

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Figure 15 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 15 Neurocranium; ventral view. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Cephaloscyllium variegatum, AMS I.43762-001, female, 670 mm TL; C, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL; D, Atelomycterus fasciatus, CSIRO H1298-7, male, 370 mm TL; E, Asymbolus rubiginosus, AMS I.30393-004, male, 527 mm TL; F, Figaro boardmani, CSIRO H989-5, female, 465 mm TL; G, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. bp, basal plate; enc, external nasal cartilage; exc, excurrent aperture; hf, hyomandibular facet; icf, internal carotid foramen; inc, incurrent aperture; lrc, lateral rostral cartilage; mrc, medial rostral cartilage; nf, nasal fontanelle; sbp, subnasal plate; sf, stapedial foramen; ss, suborbital shelf.

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Figure 24 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 24 Coracoid bar; ventral view. A, Scyliorhinus haeckelii, UERJ 1691, male, 522 mm TL; B, Halaelurus natalensis, SAIAB 26951, male, 400 mm TL; C, Holohalaelurus regani, SAIAB 25717, male, 610 mm TL; D, Cephalurus cephalus, USNM 221527, female, 285 mm TL. mpc, medial projection of the coracoid bar; lpc, lateral processes on pectoral girdle.

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Figure 8 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 8 Detail of the caudal crest in Galeus antillensis, UF 77853, female, 370 mm TL. A, general view, B, closed view. Scale bar:10 mm (A); 200 µm (B).

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Figure 14 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 14 Detail of rostral region and anterior fontanelle of Holohalaelurus regani, SAIAB 25717, male, 610 mm TL. af, anterior fontanelle; eph, epiphyseal notch; lrc, lateral rostral cartilage; mrc, medial rostral cartilage; nc, nasal capsule; pep, preorbital process.

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Figure 26 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 26 Detail of the rhipidion. A, Scyliorhinus boa, USNM 221563, 348 mm TL; B, Scyliorhinus canicula, BMNH 1983.8.3.1–4, 585.7 mm TL. Modified from Soares & de Carvalho (2019). rd, dorsal marginal cartilage; rh, rhipidion; rv, ventral marginal cartilage; t3, accessory terminal cartilage; td, dorsal terminal cartilage; td2, dorsal terminal 2 cartilage; tv, ventral terminal cartilage; tv2, ventral terminal 2 cartilage.

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Figure 25 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 25 Clasper; external morphology. A, Scyliorhinus boa, USNM 221563, 348 mm TL; B, Scyliorhinus canicula, BMNH 1983.8.3.1–4, 585.7 mm TL; C, Scyliorhinus duhamelii, MCZ S-63, 338.7 mm TL; D, Scyliorhinus retifer, UF 36359, 372 mm TL; E, Scyliorhinus stellaris, BMNH 1976.7.30.10, 476 mm TL; F, Scyliorhinus torazame, NSMT 50632, 427.9 mm TL. ch, clasper hooks; crh, cover rhipidion; dd, dermal denticles; en, envelope; erh, exorhipidion; hp, hypopyle; rh, rhipidion; tdc, terminal dermal cover. Modified from Soares and de Carvalho (2019).

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Figure 7 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 7 Dermal denticles above the origin of the first dorsal fin. A, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL; B, Asymbolus rubiginosus, AMS I.30393-004, male, 527 mm TL; C, Atelomycterus fasciatus, CSIRO H1298-7, male, 370 mm TL; D, Cephaloscyllium sufflans, SAIAB 6242, male, 800 mm TL; E, Cephalurus cephalus, USNM 221527, female, 285 mm TL; F, Galeus antillensis, UF 77853, female, 370 mm TL; G, Halaelurus natalensis, SAIAB 26951, male, 400 mm TL; H, Haploblepharus edwardsii, AMNH 40988, male, 480 mm TL; I, Holohalaelurus regani, SAIAB 25717, male, 610 mm TL; J, Parmaturus xaniurus, CAS 232152, female, 450 mm TL; K, Poroderma africanum, SAIAB 25343, male, 920 mm TL; L, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL. Scale bar: 250 µm.

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Figure 20 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 20 Detail of the hyomandibular cartilage; internal surface. A, Scyliorhinus haeckelii, UERJ 1691, male, 522 mm TL; B, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. 2358.

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Figure 12 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 12 Detail of muscle coracobranchialis. A, Cephaloscyllium umbratile, USP uncatalogued, male, 409 mm TL; B, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL. bbq, basibranchial; cb I–V, ceratobranchials I–V; cbb, basibranchial copula; cor, coracoid bar; hb II–III, hypobranchials II–III; mcb II–IV, muscles coracobranchialis I–IV; mpc, pericardial membrane.

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Figure 13 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 13 Neurocranium; dorsal view. A, Scyliorhinus ugoi, USNM 221611, male, 432 mm TL; B, Cephaloscyllium variegatum, AMS I.43762-001, female, 670 mm TL; C, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL; D, Atelomycterus fasciatus, CSIRO H1298-7, male, 370 mm TL; E, Asymbolus rubiginosus, AMS I.30393-004, male, 527 mm TL; F, Figaro boardmani, CSIRO H989-5, female, 465 mm TL; G, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL. af, anterior fontanelle; asc, anterior semicircular canal; eph, epiphyseal notch; foe, external foramen of preorbital canal; ins, internasal septum; iof, infraorbital canal of the lateral line; lrc, lateral rostral cartilage; mrc, medial rostral cartilage; nc, nasal capsule; pep, preorbital process; prf, parietal fossa; psc, posterior semicircular canal; pt, pterotic process; ptp, postorbital process; sc, supraorbital crest.

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Figure 10 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 10 Detail of the insertion region of the muscle coracomandibularis. A, Cephaloscyllium umbratile, USP uncatalogued, male, 409 mm TL; B, Haploblepharus edwardsii, AMNH 40988, male, 480 mm TL. cbm, basimandibular cartilage; cor, coracoid bar; mca, muscle coracoarcualis; mch, m. coracohyoideus; mck, Meckel's cartilage; mcm, m. coracomandibularis.

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Figure 11 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 11 Hypobranchial musculature. A, Cephaloscyllium umbratile, uncatalogued, male, 409 mm TL; B, Galeus antillensis, UF 77853, female, 370 mm TL; C, Holohalaelurus regani, SAIAB 25717, male, 610 mm TL; D, Apristurus longicephalus, HUMZ 170382, male, 475 mm TL; bh, basihyal; bhf, adjacent flap of the basihyal; ch, ceratohyal; cor, coracoid bar; mca, muscle coracoarcualis; mch, m. coracohyoideus; mcm, m. coracomandibularis; ocm, origin of the m. coracomandibularis.

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Figure 1 from: Soares KDA, de Carvalho MR (2020) Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution 96(2): 345-395. https://doi.org/10.3897/zse.96.52420

Figure 1 Ventral view of the head. A, Scyliorhinus canicula, MNHN 1999–1732, female, 418.5 mm TL; B, Schroederichthys saurisqualus, UERJ uncatalogued, female, 564 mm TL; C, Holohalaelurus regani, SAIAB 25717, male, 610 mm TL. anf, anterior nasal flap; llf, lower labial furrow; mrd, mesonarial crest; ulf, upper labial furrow. Scale bar: 20 mm.

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