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Figure 46 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figure 46 - Parsimony cladogram of subtribe Homethina with characters indicated; character numbers above cladogram stems, state numbers below stems [cladogram divided into two parts joined at "A"]; geographic abbreviations as in Figs 44–45.
Figures 40-43 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 40-43 - Female gonocoxal ovipositor and female reproductive tract, ventral view for taxa assigned to Odacanthini (see Suppl. material 1): 40, Neoeudalia nigra; 41, Odacantha melanura; 42, Colliuris ludoviciana; 43, Colliuris subdistincta. For abbreviations see Table 1.
Figures 28-33 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 28-33 - 28–32 Male aedeagus with associated abdominal segments VIII and IX, dorsal view, Cyphocoleus spp.: 28, Cyphocoleus heterogenus; 29, Cyphocoleus angustatus; 30, Cyphocoleus mirabilis; 31, Cyphocoleus cordatus; 32, Cyphocoleus subulatus. 33. Abdominal lateroventrites VIII (left) and laterotergites VIII (right), showing membranous division of laterotergites, male Cyphocoleus latipennis. Dorsal gland reservoir is globose without dorsal lobe. For abbreviations see Table 1.
Figures 44-45 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 44-45 - Parsimony cladograms comprising exemplar taxa of Odacanthini used in phylogenetic analysis, rooted at the outgroup Celaenephes linearis (tribe Lebiini): 44, 852-step length strict consensus of 29 equally parsimonious trees of 835-step length (CI = 22, RI = 74) [A = subtribe Actenonycina; H = subtribe Homethina subtrib. n.; P = subtribe Pentagonicina; O = subtribe Odacanthina]; 45, majority-rule consensus of 29 equally parsimonious trees with all resolved nodes present in 100% of the multiple equally parsimonious trees; i.e. the majority-rule consensus is the result of collapsing 8 nodes. Abbreviations for geographical distributions include: Af, Africa; As, Asia; Eu, Europe; NC, New Caledonia; Nea, Nearctic; Neo, Neotropical; NZ, New Zealand; Oz, Australia.
Figures 26-27 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 26-27 - Right maxillary stipes, lacinia, galea, and palp, Cyphocoleus subulatus: 26, dorsal view; 27, ventral view. Single-segmented galea of Cyphocoleus and other Homethina lies laterad the lacinia, with a rabbeted groove on the dorsomedial surface of the galea accommodating the lateral margin of the lacinia, thereby supporting lacinia during alternating, rotary movements of the maxillae. For abbreviations see Table 1.
Figures 34-39 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 34-39 - Female gonocoxal ovipositor and female reproductive tract, ventral view for taxa assigned to Odacanthini (see Suppl. material 1): 34, Actenonyx bembidioides; 35, Homethes gracilis; 36, Stenocheila lacordairei; 37, Pentagonica daimella; 38, Scopodes edwardsii; 39, Eudalia atrata. For abbreviations see Table 1.
Figure 166 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figure 166 - Diversification history of odacanthine subtribes and generic-level taxa within Homethina placed in temporal context defined by major geological events and molecular dating estimates (Ober and Heider 2010). Major events and dating estimates constraining historical relationships of the represented taxa include: 1, 100–105 Ma, estimated age of origin of Odacanthini relative to its proposed adelphotaxon Peleciini (Ober and Maddison 2008, Ober and Heider 2010); 2, 84–65 Ma, estimated divergence times of lineages within Odacanthini, represented in this study by Actenonycina, Pentagonicina, and Odacanthina (Ober and Heider 2010); 3, 84–61Ma, rifting of Zealandia, including New Caledonia and New Zealand, from Australia (McLoughlin 2001, Ladiges and Cantrill 2007); 4, 50–30 Ma, extended rifting and increased isolation of Australia and South America from Antarctica due to opening of the Tasman Sea at 50 Ma (Woodburne and Case 1996, Sanmartín and Ronquist 2004), development of the Australo-Antarctic Gulf at 35–32 Ma (McLoughlin 2001, Lawver and Gahagan 2003), and opening of Drake Passage at 34–30 Ma (Livermore et al. 2005); 5, 46–37 Ma, hypothesized submergence of New Caledonia (Grandcolas et al. 2008); 6, 34–23 Ma, hypothesized Oligocene drowning of New Zealand (Campbell and Hutching 2007); 7, 15–10 Ma, closing of Panamanian Isthmus establishing subaerial contact between South and Central America (Montes et al. 2015). Time of divergence of Cyphocoleus unconstrained due to lack of molecular-dating data. Under vicariance hypothesis (v), the ancestor of Cyphocoleus was emplaced on Zealandia in Late Cretaceous (event 3). Under dispersal scenario (d with dashed line), Cyphocoleus dispersal to New Caledonia is constrained at the lower bound by reemergence of Grande Terre after putative Oligocene submergence (event 5), and at upper bound by amphiantarctic vicariance at Austral-Antarctic Gulf that isolated Antarctica from Australia (event 4).
Figures 14-25 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 14-25 - 14–18 Male aedeagus with associated abdominal segments VIII and IX, dorsal view for taxa assigned to Odacanthini (see Suppl. material 1): 14, Colliuris oglobini; 15, Erectocolliuris fairmairei; 16, Basistichus micans; 17, Archicolliuris papua; 18, Colliuris ludoviciana; 19–24 Male aedeagus with associated abdominal segments VIII and IX, dorsal view, for taxa assigned to Lachnophorini (Erwin and Zamorano 2014; repository indicated in parentheses): 19, Selina westermanni Motschulsky (BMUL); 20, Eucaerus sp. (Trinidad, Simla; CUIC); 21, Anchonoderus subtilis Bates (CUIC); 22, Euphorticus pubescens (Dejean) (CUIC); 23, Lachnophorus leucopterus Chevrolat (CUIC); 24, Calybe argentina Brèthes (CUIC). 25. Male aedeagus with associated abdominal segments VIII and IX, dorsal view, for taxon assigned to Calophaenini (Erwin and Zamorano 2014; repository indicated in parentheses): Calophaena cruciata Bates (CUIC). For abbreviations see Table 1.
Figures 161-164 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 161-164 - Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 161, Cyphocoleus flavipes Yahoué; 162, Cyphocoleus bourailensis, Bourail; 163, Cyphocoleus subulatus, Mt. Koghi; 124, Cyphocoleus iledespinsensis, Île des Pins.
Figures 141-160 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 141-160 - Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141–152, Cyphocoleus globulicollis; 141, Aoupinié; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148–150, Mt. Dzumac Rd; 151–152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoué; 154, Cyphocoleus bourailensis, Bourail; 155–158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159–160, Cyphocoleus iledespinsensis, Île des Pins; 159, dextral view; 160, ventral view.
Figures 137-140 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 137-140 - Distribution maps of Cyphocoleus spp.: 137, Cyphocoleus cychroides, Cyphocoleus cordatus, Cyphocoleus cardiopterus; 138, Cyphocoleus latipennis; 139, Cyphocoleus globulicollis; 140 Cyphocoleus flavipes, Cyphocoleus bourailensis.
Figures 86-98 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 86-98 - Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panié; 88, Cyphocoleus monteithi, Parque Prov. Rivière Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panié; 91–98, Cyphocoleus cychroides; 91, internal sac everted, Mandjélia; 92–93, Mandjélia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation.
Figures 121-125 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 121-125 - Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panié; 123, Cyphocoleus cordatus female, Forêt Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes.
Figures 2-13 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 2-13 - Male aedeagus with associated abdominal segments VIII and IX, dorsal view, for taxa assigned to Odacanthini (see Suppl. material 1): 2, Actenonyx bembidioides; 3, Homethes guttifer; 4, Quammenis spectabilis; 5, Stenocheila lacordairei; 6, Pentagonica daimella; 7, Scopodes edwardsii; 8, Lasiocera orientalis; 9, Eucolliuris fuscipennis; 10, Renneria kamouni; 11, Arame macra; 12, Odacantha melanura; 13, Deipyrodes palustris. For abbreviations see Table 1.
Figure 1 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figure 1 - Cyphocoleus subulatus sp. n. male paratype dorsal view: NEW CALEDONIA, Ciu, near Mt. Canala, 300 m el., 1–3–i–1955, rainforest, E. O. Wilson (MCZ).
Figures 76-81 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 76-81 - Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni.
Figures 82-85 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 82-85 - Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panié; 85, Cyphocoleus monteithi male, Dzumac Rd. jct.
Figures 51-59 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 51-59 - Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoué; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58–59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation.
Figures 66-75 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 66-75 - Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus , Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoué; 69, Cyphocoleus parovicollis, Mandjélia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Forêt Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinié. Scale bar, 0.10 mm.
Figures 60-65 from: Liebherr JK (2016) Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63(2): 211-270. https://doi.org/10.3897/dez.63.10241
Figures 60-65 - Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoué; 63, Cyphocoleus parovicollis, Mandjélia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Forêt Nord. For abbreviations see Table 1.
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