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FIGURE 3 in A further new species of Palaemonella Dana, 1852 from the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Palaemonidae)
FIGURE 3. Palaemonella yalla sp. nov., living shrimps in dorsal view; A, holotype, ovigerous female (pocl 3.9 mm, cl 5.2 mm), Thuwal, Saudi Arabia, FLMNH UF 71446; B, non-type, ovigerous female (pocl 3.4 mm, cl 5.6 mm), Masirah Island, Oman, FLMNH UF 71445. Photographs by A. Anker.
FIGURE 15 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 15. Pseudathanas banneri (Kazmi & Kazmi, 2010) comb. nov., male (cl 6.7 mm) from Muscat, Oman (FLMNH UF 68864); a, right cheliped (drawn in situ), lateral; b, same, carpus and chela, mesial; c, left cheliped, distal portion of ischium, merus and carpus (drawn in situ), lateral.
FIGURE 12 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species' habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c].
FIGURE 11 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral
FIGURE 9. Athanopsis saurus Anker, 2011 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 9. Athanopsis saurus Anker, 2011, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [a–e] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably Ochetostoma sp. (Thalassematidae), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author.
FIGURE 8 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 8. Athanopsis brevirostris Banner & Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author.
FIGURE 7 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 7. Athanas parvus De Man, 1910, male (cl 2.9 mm) from Masirah Island, Oman (FLMNH UF 71418); a, shrimp in life, lateral (left side); b, dorsal. Photographs by the author.
FIGURE 13. Athanopsis rubricinctuta Berggren, 1991 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 13. Athanopsis rubricinctuta Berggren, 1991, male (cl 5.4 mm) from Masirah Island, Oman (FLMNH UF 71437) [a, b]; male (cl 5.8 mm) from the same locality (FLMNH UF 71443) [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, shrimp in life (different individual), lateral (left side). Photographs by the author.
FIGURE 6 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 6. Athanas minikoensis Coutière, 1903, ovigerous female (cl 3.2 mm) from Masirah Island, Oman (FLMNH UF 71422); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author.
FIGURE 2 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral.
FIGURE 4 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 4. Athanas cf. dentirostris Anker, Jeng & Chan, 2001, male (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71432); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author.
FIGURE 1 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted).
FIGURE 3 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author.
FIGURE 14 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 14. Athanopsis tarahomii Marin, Sheibani & Sari, 2014, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author.
FIGURE 16 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 16. Pseudathanas banneri (Kazmi & Kazmi, 2010) comb. nov., male (cl 6.7 mm) from Muscat, Oman (FLMNH UF 68864) [a, b]; female (cl 6.45 mm) from the same locality (not deposited, see text) [c–e]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), frontal region, dorsal; d, e, left cheliped, lateral (d) and mesial (e). Photographs by the author.
FIGURE 5 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 5. Athanas iranicus Anker, Naderloo & Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author.
FIGURE 10 in Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)
FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal.
Subspecies and Distribution. V. v. vulpes Linnaeus, 1758 — N Europe (Scandinavia). V. v. abietorum Merriam, 1900 — SW Canada (Alberta & British Columbia). V. v. aegyptiacus Sonnini, 1816 — Egypt, Israel, and Lybia. V. v. alascensis Merriam, 1900 — Alaska and NW Canada (NW Territories & Yukon). V. v. alpheraky: Satunin, 1906 — Kazakhstan. V. v. anatolica Thomas, 1920 — Turkey. V. v. arabica Thomas, 1902 — Arabian peninsula. V. v. atlantica Wagner, 1841 — Algeria (forested Atlas Mts). V. v. bangsi Merriam, 1900 — NE Canada (Labrador). V. v. barbara Shaw, 1800 — NW Africa (Barbary Coast). V. v. beringiana Middendorff, 1875 — NE Siberia (shore of Bering Strait). V. v. cascadensis Merriam, 1900 — NW USA (Cascade Mountains, Oregon & Washington). V. v. caucasica Dinnik, 1914 — SW Russia (Caucasus). V. v. crucigera Bechstein, 1789 — Europe through N & C Russia. V. v. daurica Ognev, 1931 — E Russia (Amur, Siberia & Transbaikalia). V.v. deletrix Bangs, 1898 — NE Canada (Newfoundland). V. v. dolichocrania Ognev, 1926 — SE Siberia (S Ussuri). V. v. flavescens Gray, 1843 — N Iran. V. v. fulva Desmarest, 1820 — E USA. V. v. griffith: Blyth, 1854 — Afghanistan and N Pakistan. V.v. harrimani Merriam, 1900 — Alaska (Kodiak I). V. v. hoole Swinhoe, 1870 — S China (Fujian to Sichuan). V. v. ichnusae G. S. Miller, 1907 — Corsica and Sardinia. V. v. induta G. S. Miller, 1907 — Cyprus. V. v. jakutensis Ognev, 1923 — E Siberia (S of Yakutsk). V. v. japonica Gray, 1868 — Japan. V. v. karagan Erxleben, 1777 — Mongolia, Kazakhstan, and Kirgizstan. V. v. kenaiensis Merriam, 1900 — Alaska (Kenai Peninsula). V. v. kurdistanica Satunin, 1906 — Armenia and NE Turkey. V. v. macroura Baird, 1852 — USA (Mountain States). V. v. montana Pearson, 1836 — Himalayas form China (Yunnan) to C Pakistan. V. v. mecator Merriam, 1900 — SW USA (California & Nevada). V_ v. ochroxantha Ognev, 1926 — E Russian Turkestan, Aksai, Kirgizstan, Semirechie. V. v. palaestina Thomas, 1920 —Jordan and Lebanon. V.v. peculiosa Kishida, 1924 — Korea. V. v. pusilla Blyth, 1854 — NW India to Irak. V.v. regalis Merriam, 1900 — N Great Plains of Canada and USA. V. v. rubricosa Bangs, 1898 — E Canada. V.v. schrencki Kishida, 1924 — N Japan (Hokkaido) and NE Russia (Sakhalin). V. v. silacea G. S. Miller, 1907 — Iberian Peninsula. V.v. splendidissima Kishida, 1924 — E Russia (N & C Kurile Is). V. v. stepensis Brauner, 1914 — steppes of S Russia. V. v. tobolica Ognev, 1926 — Russia (lower basin of Ob River) V. v. tschiliensis Matschie, 1907 — NE China. Foxes of European origin were introduced into E USA and Canada in the 17" century, subsequently mixed with local subspecies. Also introduced to Australia in 1800s, and the Falkland Islands (Malvinas). in Canidae
Subspecies and Distribution. V. v. vulpes Linnaeus, 1758 — N Europe (Scandinavia). V. v. abietorum Merriam, 1900 — SW Canada (Alberta & British Columbia). V. v. aegyptiacus Sonnini, 1816 — Egypt, Israel, and Lybia. V. v. alascensis Merriam, 1900 — Alaska and NW Canada (NW Territories & Yukon). V. v. alpheraky: Satunin, 1906 — Kazakhstan. V. v. anatolica Thomas, 1920 — Turkey. V. v. arabica Thomas, 1902 — Arabian peninsula. V. v. atlantica Wagner, 1841 — Algeria (forested Atlas Mts). V. v. bangsi Merriam, 1900 — NE Canada (Labrador). V. v. barbara Shaw, 1800 — NW Africa (Barbary Coast). V. v. beringiana Middendorff, 1875 — NE Siberia (shore of Bering Strait). V. v. cascadensis Merriam, 1900 — NW USA (Cascade Mountains, Oregon & Washington). V. v. caucasica Dinnik, 1914 — SW Russia (Caucasus). V. v. crucigera Bechstein, 1789 — Europe through N & C Russia. V. v. daurica Ognev, 1931 — E Russia (Amur, Siberia & Transbaikalia). V.v. deletrix Bangs, 1898 — NE Canada (Newfoundland). V. v. dolichocrania Ognev, 1926 — SE Siberia (S Ussuri). V. v. flavescens Gray, 1843 — N Iran. V. v. fulva Desmarest, 1820 — E USA. V. v. griffith: Blyth, 1854 — Afghanistan and N Pakistan. V.v. harrimani Merriam, 1900 — Alaska (Kodiak I). V. v. hoole Swinhoe, 1870 — S China (Fujian to Sichuan). V. v. ichnusae G. S. Miller, 1907 — Corsica and Sardinia. V. v. induta G. S. Miller, 1907 — Cyprus. V. v. jakutensis Ognev, 1923 — E Siberia (S of Yakutsk). V. v. japonica Gray, 1868 — Japan. V. v. karagan Erxleben, 1777 — Mongolia, Kazakhstan, and Kirgizstan. V. v. kenaiensis Merriam, 1900 — Alaska (Kenai Peninsula). V. v. kurdistanica Satunin, 1906 — Armenia and NE Turkey. V. v. macroura Baird, 1852 — USA (Mountain States). V. v. montana Pearson, 1836 — Himalayas form China (Yunnan) to C Pakistan. V. v. mecator Merriam, 1900 — SW USA (California & Nevada). V_ v. ochroxantha Ognev, 1926 — E Russian Turkestan, Aksai, Kirgizstan, Semirechie. V. v. palaestina Thomas, 1920 —Jordan and Lebanon. V.v. peculiosa Kishida, 1924 — Korea. V. v. pusilla Blyth, 1854 — NW India to Irak. V.v. regalis Merriam, 1900 — N Great Plains of Canada and USA. V. v. rubricosa Bangs, 1898 — E Canada. V.v. schrencki Kishida, 1924 — N Japan (Hokkaido) and NE Russia (Sakhalin). V. v. silacea G. S. Miller, 1907 — Iberian Peninsula. V.v. splendidissima Kishida, 1924 — E Russia (N & C Kurile Is). V. v. stepensis Brauner, 1914 — steppes of S Russia. V. v. tobolica Ognev, 1926 — Russia (lower basin of Ob River) V. v. tschiliensis Matschie, 1907 — NE China. Foxes of European origin were introduced into E USA and Canada in the 17" century, subsequently mixed with local subspecies. Also introduced to Australia in 1800s, and the Falkland Islands (Malvinas).
Distribution. Widespread in N and NE Africa, occurring from Senegal on the W coast of Africa to Egypt in the E, in a range that includes Morocco, Algeria, Tunisia, and Libya in the N to Nigeria, Chad, and Tanzania in the S. They have expanded their range from the Arabian Peninsula into Western Europe, to Bulgaria, Austria, and NE Italy and E into Turkey, Syria, Iraq, Iran, Central Asia, the entire Indian subcontinent, then E and S to Sri Lanka, Myanmar, Thailand, and parts of Indochina. in Canidae
Distribution. Widespread in N and NE Africa, occurring from Senegal on the W coast of Africa to Egypt in the E, in a range that includes Morocco, Algeria, Tunisia, and Libya in the N to Nigeria, Chad, and Tanzania in the S. They have expanded their range from the Arabian Peninsula into Western Europe, to Bulgaria, Austria, and NE Italy and E into Turkey, Syria, Iraq, Iran, Central Asia, the entire Indian subcontinent, then E and S to Sri Lanka, Myanmar, Thailand, and parts of Indochina.
Subspecies and Distribution. Vr. rueppellii Schinz, 1825 — Egypt and Sudan (Nubian Desert). V. r. caesia Thomas & Hinton, 1921 — N & W Africa. V. r. cyrenaica Festa, 1921 — SW Egypt, Lybia, extreme NW Sudan. V. r. sabaea Pocock, 1934 — Arabian Peninsula and Middle East. V. r. somaliae Thomas, 1918 — Eritrea, Ethiopia, and Somalia. V. r. zarudny: Birula, 1913 — Baluchistan in Afghanistan, Iran, and Pakistan. in Canidae
Subspecies and Distribution. Vr. rueppellii Schinz, 1825 — Egypt and Sudan (Nubian Desert). V. r. caesia Thomas & Hinton, 1921 — N & W Africa. V. r. cyrenaica Festa, 1921 — SW Egypt, Lybia, extreme NW Sudan. V. r. sabaea Pocock, 1934 — Arabian Peninsula and Middle East. V. r. somaliae Thomas, 1918 — Eritrea, Ethiopia, and Somalia. V. r. zarudny: Birula, 1913 — Baluchistan in Afghanistan, Iran, and Pakistan.
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