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Subspecies and Distribution. M. e. erminea Linnaeus, 1758 — Finland, Norway, NW Russia, and Sweden. M. e. aestiva Kerr, 1792 — most of mainland N & C Europe to C Asia in Kazakhstan, Kyrgyzstan, and Tajikistan. M. e. alascensis Merriam, 1896 — S Alaska. M. e. anguinae Hall, 1932 — SW Canada (Vancouver I, British Columbia). M. e. arctica Merriam, 1896 — Alaska and NW Canada. M. e. bangsi Hall, 1945 — C Canada and NC USA. M. e. celenda Hall, 1944 — Alaska (Prince of Wales I). M.e. cicognanii Bonaparte, 1838 — SE Canada and NE USA. M. e. fallenda Hall, 1945 — W Canada (British Columbia) and NW USA (N Washington). M. e. ferghanae Thomas, 1895 — Afghanistan, N India, and Pakistan. M.e. gulosa Hall, 1945 — NW USA (E Washington). M.e. haidarum Preble, 1898 — W Canada (Queen Charlotte Is, British Columbia). M.e. hibernica Thomas & Barrett-Hamilton, 1895 — Ireland. M.e. imatis Hall, 1944 — Alaska (Baranof I). M.e. invicta Hall, 1945 — SW Canada (Alberta) and NW USA (Idaho & Montana). M.e. kadiacensis Merriam, 1896 — Alaska (Kodiak I). M.e. kaneti Baird, 1857 — NE China, Russia (E Siberia). M.e. karaginensis Jurgenson, 1936 — NE Russia (Karaginsky I). M.e. lymani Hollister, 1912 — E Russia (Altai Mts, Siberia). M.e. minima Cavazza, 1912 — Switzerland. M.e. mongolica Ognev, 1928 — NW China and Mongolian Altai. M. e. muricus Bangs, 1899 — USA (N California, Colorado, Idaho, New Mexico, Nevada, Oregon, South Dakota, Utah & Wyoming). M. e. nippon Cabrera, 1913 — Japan. M.e. olympica Hall, 1945 — NW USA (Olympic Peninsula, Washington). M.e. polaris Barrett-Hamilton, 1904 — Greenland. M.e.richardsonii Bonaparte, 1838 — N Canada. M.e. ricinae G. S. Miller, 1907 — Scotland (Islay I). M.e.salva Hall, 1944 — SE Alaska (Admiralty I). M.e.seclusa Hall, 1944 — SE Alaska (Suemez I). M.e.sempler Sutton & Hamilton, 1932 — Canada (Franklin & Keewatin Districts). M.e.stabilis Barrett-Hamilton, 1904 — Great Britain. M.e.streatori Merriam, 1896 — W USA (NE California, Oregon & coastal Washington). M.e.teberdina Kornejv, 1941 — Russian Caucasus. M.e. tobolica Ognev, 1923 — W Siberia. Introduced to New Zealand. in Mustelidae

Subspecies and Distribution. M. e. erminea Linnaeus, 1758 — Finland, Norway, NW Russia, and Sweden. M. e. aestiva Kerr, 1792 — most of mainland N & C Europe to C Asia in Kazakhstan, Kyrgyzstan, and Tajikistan. M. e. alascensis Merriam, 1896 — S Alaska. M. e. anguinae Hall, 1932 — SW Canada (Vancouver I, British Columbia). M. e. arctica Merriam, 1896 — Alaska and NW Canada. M. e. bangsi Hall, 1945 — C Canada and NC USA. M. e. celenda Hall, 1944 — Alaska (Prince of Wales I). M.e. cicognanii Bonaparte, 1838 — SE Canada and NE USA. M. e. fallenda Hall, 1945 — W Canada (British Columbia) and NW USA (N Washington). M. e. ferghanae Thomas, 1895 — Afghanistan, N India, and Pakistan. M.e. gulosa Hall, 1945 — NW USA (E Washington). M.e. haidarum Preble, 1898 — W Canada (Queen Charlotte Is, British Columbia). M.e. hibernica Thomas & Barrett-Hamilton, 1895 — Ireland. M.e. imatis Hall, 1944 — Alaska (Baranof I). M.e. invicta Hall, 1945 — SW Canada (Alberta) and NW USA (Idaho & Montana). M.e. kadiacensis Merriam, 1896 — Alaska (Kodiak I). M.e. kaneti Baird, 1857 — NE China, Russia (E Siberia). M.e. karaginensis Jurgenson, 1936 — NE Russia (Karaginsky I). M.e. lymani Hollister, 1912 — E Russia (Altai Mts, Siberia). M.e. minima Cavazza, 1912 — Switzerland. M.e. mongolica Ognev, 1928 — NW China and Mongolian Altai. M. e. muricus Bangs, 1899 — USA (N California, Colorado, Idaho, New Mexico, Nevada, Oregon, South Dakota, Utah & Wyoming). M. e. nippon Cabrera, 1913 — Japan. M.e. olympica Hall, 1945 — NW USA (Olympic Peninsula, Washington). M.e. polaris Barrett-Hamilton, 1904 — Greenland. M.e.richardsonii Bonaparte, 1838 — N Canada. M.e. ricinae G. S. Miller, 1907 — Scotland (Islay I). M.e.salva Hall, 1944 — SE Alaska (Admiralty I). M.e.seclusa Hall, 1944 — SE Alaska (Suemez I). M.e.sempler Sutton & Hamilton, 1932 — Canada (Franklin & Keewatin Districts). M.e.stabilis Barrett-Hamilton, 1904 — Great Britain. M.e.streatori Merriam, 1896 — W USA (NE California, Oregon & coastal Washington). M.e.teberdina Kornejv, 1941 — Russian Caucasus. M.e. tobolica Ognev, 1923 — W Siberia. Introduced to New Zealand.

Subspecies and Distribution. M. n. nivalis Linnaeus, 1766 — China, North and South Korea, Mongolia, Russia, Taiwan, and Scandinavia. M. n. allegheniensis Rhoads, 1900 — NE USA (Allegheny Mts W to Wisconsin). M. n. boccamela Bechstein, 1800 — Corsica, Italy, Portugal, Sardinia, Sicily, and Spain. M. n. campestris Jackson, 1913 — C Great Plains, USA. M. n. eskimo Stone, 1900 — Alaska and NW Canada (Yukon). M. n. namiyer Kuroda, 1921 — Japan and the Kurile Is. M. n. numidica Pucheran, 1855 — N Africa. M. n. rnixosa Bangs, 1896 — Canada and N Great Plains of USA. M. n. vulgaris Erxleben, 1777 = W & C Europe and most of C Eurasia. Introduced to New Zealand, Malta, Crete, the Azores Is, and apparently also Sao Tome I. in Mustelidae

Subspecies and Distribution. M. n. nivalis Linnaeus, 1766 — China, North and South Korea, Mongolia, Russia, Taiwan, and Scandinavia. M. n. allegheniensis Rhoads, 1900 — NE USA (Allegheny Mts W to Wisconsin). M. n. boccamela Bechstein, 1800 — Corsica, Italy, Portugal, Sardinia, Sicily, and Spain. M. n. campestris Jackson, 1913 — C Great Plains, USA. M. n. eskimo Stone, 1900 — Alaska and NW Canada (Yukon). M. n. namiyer Kuroda, 1921 — Japan and the Kurile Is. M. n. numidica Pucheran, 1855 — N Africa. M. n. rnixosa Bangs, 1896 — Canada and N Great Plains of USA. M. n. vulgaris Erxleben, 1777 = W & C Europe and most of C Eurasia. Introduced to New Zealand, Malta, Crete, the Azores Is, and apparently also Sao Tome I.

Subspecies and Distribution. N. v. vison Schreber, 1776 — E Canada and NE USA (Allegheny Mts). N. v. aestuarina Grinnell, 1916 — SW USA (California & W Nevada). N. v. aniakensis Burns, 1964 — W Alaska. N. v. energumenos Bangs, 1896 — W Canada and NW USA. N. v. evagor Hall, 1932 — SW Canada (Vancouver I). N. v. evergladensis Hamilton, 1948 — SE USA (SW Florida). N. v. ingens Osgood, 1900 — most of Alaska and NW Canada (N Mackenzie & Yukon). N. v. lacustris Preble, 1902 — C Canada (Keewatin Region, Manitoba & Ontario). N. v. letifera Hollister, 1913 — Great Plains of USA. N. v. low Anderson, 1945 — E Canada (Labrador & Quebec). N. v. lutensis Bangs, 1898 — USA (S Carolina to Florida). N. v. melampeplus Elliot, 1903 — Alaska (Kenai Peninsula). N. v. mink Peale & Palisot de Beauvois, 1796 — SE USA. N. v. nesolestes Heller, 1909 — SW Alaska. N. v. vulgivaga Bangs, 1895 — USA (Arkansas & Louisiana). Introduced to Belarus, Belgium, China, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britian, Iceland, Ireland, Italy, Japan (Hokkaido), Latvia, Lithuania, Netherlands, Norway, Poland, Portugal, Russia, Spain, and Sweden. in Mustelidae

Subspecies and Distribution. N. v. vison Schreber, 1776 — E Canada and NE USA (Allegheny Mts). N. v. aestuarina Grinnell, 1916 — SW USA (California & W Nevada). N. v. aniakensis Burns, 1964 — W Alaska. N. v. energumenos Bangs, 1896 — W Canada and NW USA. N. v. evagor Hall, 1932 — SW Canada (Vancouver I). N. v. evergladensis Hamilton, 1948 — SE USA (SW Florida). N. v. ingens Osgood, 1900 — most of Alaska and NW Canada (N Mackenzie & Yukon). N. v. lacustris Preble, 1902 — C Canada (Keewatin Region, Manitoba & Ontario). N. v. letifera Hollister, 1913 — Great Plains of USA. N. v. low Anderson, 1945 — E Canada (Labrador & Quebec). N. v. lutensis Bangs, 1898 — USA (S Carolina to Florida). N. v. melampeplus Elliot, 1903 — Alaska (Kenai Peninsula). N. v. mink Peale & Palisot de Beauvois, 1796 — SE USA. N. v. nesolestes Heller, 1909 — SW Alaska. N. v. vulgivaga Bangs, 1895 — USA (Arkansas & Louisiana). Introduced to Belarus, Belgium, China, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britian, Iceland, Ireland, Italy, Japan (Hokkaido), Latvia, Lithuania, Netherlands, Norway, Poland, Portugal, Russia, Spain, and Sweden.

Subspecies and Distribution. G. g. gulo Linnaeus, 1758 — NE & NW China, Mongolia, Russia, and Scandinavia. G. g. luscus Linnaeus, 1758 — Canada and W USA (Alaska, California, Idaho, Oregon, Montana, Washington & Wyoming). in Mustelidae

Subspecies and Distribution. G. g. gulo Linnaeus, 1758 — NE & NW China, Mongolia, Russia, and Scandinavia. G. g. luscus Linnaeus, 1758 — Canada and W USA (Alaska, California, Idaho, Oregon, Montana, Washington & Wyoming).

Subspecies and Distribution. M. m. moschata Gray, 1831 — SE China (Guangdong, Guangxi, Guizhou, Yunnan & Hainan I) and N Laos. M. m. ferreogrisea Hilzheimer, 1905 — C China. M. m. millsi Thomas, 1922 — S China (NW Yunnan) through N Myanmar to NE India. M. m. sorella G. M.. Allen, 1929 — E China (Fujian). M. m. subaurantiaca Swinhoe, 1862 — Taiwan. M. m. taxilla Thomas, 1925 — Vietnam. in Mustelidae

Subspecies and Distribution. M. m. moschata Gray, 1831 — SE China (Guangdong, Guangxi, Guizhou, Yunnan & Hainan I) and N Laos. M. m. ferreogrisea Hilzheimer, 1905 — C China. M. m. millsi Thomas, 1922 — S China (NW Yunnan) through N Myanmar to NE India. M. m. sorella G. M.. Allen, 1929 — E China (Fujian). M. m. subaurantiaca Swinhoe, 1862 — Taiwan. M. m. taxilla Thomas, 1925 — Vietnam.

Subspecies and Distribution. M.c.chrysogasterHodgson,1839—alpinezoneat2800-4000mofNIndia(includingSikkim),Nepal,andBhutan. M. c. sifanicus Büchner, 1891 — alpine zone at 3500-4800 m of C & S China (S Ningxia, Qinghai, S Gansu, W Sichuan, SE Xizang & NW Yunnan). in Moschidae

Subspecies and Distribution. M.c.chrysogasterHodgson,1839—alpinezoneat2800-4000mofNIndia(includingSikkim),Nepal,andBhutan. M. c. sifanicus Büchner, 1891 — alpine zone at 3500-4800 m of C & S China (S Ningxia, Qinghai, S Gansu, W Sichuan, SE Xizang & NW Yunnan).

Subspecies and Distribution. E.k.kiangMoorcroft,1841—WChina(SWXinjiang,C&WXizang),NEPakistan(KhunjerabNationalPark),NIndia(Ladakh),andNNepal(Mustang). E.k.holderersMatschie,1911—WCChina(SEXinjiang,Quinghai,Gansu&EXizang). E. k. polyodon Hodgson, 1847 — SW China (S Xizang) and N India (N Sikkim); also possibly in extreme N & NW Bhutan. in Equidae

Subspecies and Distribution. E.k.kiangMoorcroft,1841—WChina(SWXinjiang,C&WXizang),NEPakistan(KhunjerabNationalPark),NIndia(Ladakh),andNNepal(Mustang). E.k.holderersMatschie,1911—WCChina(SEXinjiang,Quinghai,Gansu&EXizang). E. k. polyodon Hodgson, 1847 — SW China (S Xizang) and N India (N Sikkim); also possibly in extreme N & NW Bhutan.

Subspecies and Distribution. C.b.bactrianusLinnaeus,1758—aridlandsanddesertsofC&SAsiaasdomesticatedlivestock. C. b. ferus Przewalski, 1878 — NW China (C & SE Xinjiang, Nei Mongol, NW Qinghai & NW Gansu) and Mongolia. in Camelidae

Subspecies and Distribution. C.b.bactrianusLinnaeus,1758—aridlandsanddesertsofC&SAsiaasdomesticatedlivestock. C. b. ferus Przewalski, 1878 — NW China (C & SE Xinjiang, Nei Mongol, NW Qinghai & NW Gansu) and Mongolia.

Distribution. Fragmented distribution in the Tibetan Plateau in China (Xinjiang, Xizang & Qinghai); one population drifts from S Xinjiang into NW India (Ladakh) and another from Qinghai to Gansu, China. in Bovidae

Distribution. Fragmented distribution in the Tibetan Plateau in China (Xinjiang, Xizang & Qinghai); one population drifts from S Xinjiang into NW India (Ladakh) and another from Qinghai to Gansu, China.

Subspecies and Distribution. S. s. scrofa Linnaeus, 1758 — W Europe, from Denmark, Germany, Poland, and Czech Republic to N Italy and N Iberian Peninsula; possibly also Albania. The taxonomic status of animals in Austria, Switzerland, Slovenia, and Slovakia is unclear but presumably these populations are included in scrofa, as are the populations of Sweden, Finland, and the Baltic states. However, restocking of once depleted populations, for example in Italy, has likely involved the introduction and mixing of this subspecies with other subspecies, such as attila. S. s. affinis Gray, 1847 — S India and Sri Lanka. S. s. algirus Loche, 1867 — Tunisia, Algeria, and Morocco, on the coastal side of the mountains or in the low montane areas. S. s. attila Thomas, 1912 — Hungary, Ukraine, C & S Belarus, Romania, Moldova, and S Russia towards the N flank of the Caucasus, but not including the Transcaucasian countries of Georgia, Armenia, and Azerbaijan. The range possibly extends as far S as the Mesopotamian Delta in Iraq, in which case it would likely include W & SW Iran, and possibly E Turkey and Syria, where it borders with lybicus. Such a range could not be easily reconciled with a statement by Groves that "the difference between pigs from N and S of the Caucasus is quite striking; Transcaucasian boars are certainly not attila." This subspecies may also extend into C Asia and include Kazakhstan, Uzbekistan, and Turkmenistan, but no data exist to support this. S. s. baeticus Thomas, 1912 — originally described from Coto Donana, S Spain, and later merged with meridionalis; also S Portugal. Unless evidence is found that these Italian and Iberian populations are the relics of a much larger formerly contiguous range, this subspecies should be kept as distinct. S. s. coreanus Heude, 1897 — Korean Peninsula. S. s. eristatus Wagner, 1839 — Himalayas S to C India and E to Indochina (N of the Kra Isthmus). S. s. davidi Groves, 1981 — the arid zone from E Iran to Gujarat, including Pakistan and NW India, and perhaps N to Tajikistan. S. s. leucomystax Temminck, 1842 — main Is ofJapan (Honshu, Shikoku, Kyushu, Nakadori, Hiburijima, Tojima, Kushima, and other smaller Is). S. s. lybicus Gray, 1868 — Bulgaria, Greece, Turkey, Syria, Jordan, Israel, Palestine, in the past also in Lybia, and Egypt. The former Yugoslavia was included in its range, which would suggest that now Slovenia, Serbia, Croatia, Bosnia and Herzegovina, Montenegro, and Kosovo are within the range of this subspecies, although the exact boundaries are unclear. Pigs from Albania have been assigned to S. s. scrofa. S. s. majori De Beaux & Festa, 1927 — C & S Italian Peninsula. S. s. menidionalis Forsyth Major, 1882 — Corsica and Sardinia, with the proviso that the two populations are very likely to be introduced or feral. S. s. moupinensis Milne-Edwards, 1871 — China, S to Vietnam and W to Sichuan. S. s. nigripes Blanford, 1875 — the flanks of the Tianshan mountains in Kyrgyzstan and NW China (Xinjiang). An animal photographed in NE Iran (Golestan) looked like this subspecies. S. s. nukiuanus Kuroda, 1924 — Iriomote, Ishigaki, Okinawa, Tokunoshima, Amamioshima, and Kakerome Is in the Ryukyu chain in extreme S Japan, though some of these populations have hybridized with introduced domesticates. S. s. sibiricus Staffe, 1922 — Mongolia and Transbaikal (S & E of Lake Baikal). S. s. tawvanus Swinhoe, 1863 — Taiwan. S. s. ussuricus Heude, 1888 — far E Russia and the Manchurian region (China). Korean populations were previously included in this subspecies, but based on new evidence, the Korean taxon seems more similar to moupinensis. S. s. vittatus Boie, 1828 — Malay Peninsula, S of the Isthmus of Kra, the offshore islands of Terutai and Langkawi, Sumatra, Riau Archipelago, Java, Bali, and a range of smaller islands around these, including Babi, Bakong, Batam, Bawean, Bengkalis, Bintan, Bulan, Bunguran, Cuyo, Deli, Durian, Enggano, Galang, Jambongan, Karimon (Riau Is), Kundur, Lagong, Laut, Lingga, Lingung, Mapor, Moro Kecil, North Pagai, Nias, Panaitan, Payong, Penang, Pinie, Rupat, Siantan, Siberut, Simeulue, Singkep, Sugi, Sugi Bawa, Telibon, Tinggi, Tuangku, and the Tambelan Is. This species was originally present from the British Is in the extreme W, through Eurasia from S Scandinavia to S Siberia, extending as far E as Korea and Japan, and SE into some of the Sunda Is and Taiwan. In the S the species ranged along the Nile Valley to Khartoum, and N of the Sahara in Africa, more orless following the continental coasts of S, E, and SE Asia. Within this range it was absent only from extremely dry deserts, e.g. the driest regions of Mongolia and in China W of Sichuan; and alpine zones, such as the high altitudes of Pamir and Tien Shan. In recent centuries, the range of S. scrofa has changed dramatically because of hunting and changes in available habitat. The species disappeared from the British Is in the 17" century, from Denmark in the 19" century, and was greatly reduced in range and numbers in the 20" century from areas as distant as Tunisia, Sudan, Germany, and Russia. Following these severe declines, there were some slight population recoveries in Russia, Italy, Spain, and Germany in the mid-20™ century, and natural and assisted range expansions in Denmark and Sweden. The species has also been inadvertently reintroduced in various locations in the Great Britain via escapees of mixed origin from commercial farming enterprises. Ex-S. scrofa stocks also occur as introduced feral populations in various other parts of the world, including Australia, New Zealand, the eastern Malay Archipelago, and in North, Central, and South America. In all of these areas they are now generally recognized as a major pest. in Suidae

Subspecies and Distribution. S. s. scrofa Linnaeus, 1758 — W Europe, from Denmark, Germany, Poland, and Czech Republic to N Italy and N Iberian Peninsula; possibly also Albania. The taxonomic status of animals in Austria, Switzerland, Slovenia, and Slovakia is unclear but presumably these populations are included in scrofa, as are the populations of Sweden, Finland, and the Baltic states. However, restocking of once depleted populations, for example in Italy, has likely involved the introduction and mixing of this subspecies with other subspecies, such as attila. S. s. affinis Gray, 1847 — S India and Sri Lanka. S. s. algirus Loche, 1867 — Tunisia, Algeria, and Morocco, on the coastal side of the mountains or in the low montane areas. S. s. attila Thomas, 1912 — Hungary, Ukraine, C & S Belarus, Romania, Moldova, and S Russia towards the N flank of the Caucasus, but not including the Transcaucasian countries of Georgia, Armenia, and Azerbaijan. The range possibly extends as far S as the Mesopotamian Delta in Iraq, in which case it would likely include W & SW Iran, and possibly E Turkey and Syria, where it borders with lybicus. Such a range could not be easily reconciled with a statement by Groves that "the difference between pigs from N and S of the Caucasus is quite striking; Transcaucasian boars are certainly not attila." This subspecies may also extend into C Asia and include Kazakhstan, Uzbekistan, and Turkmenistan, but no data exist to support this. S. s. baeticus Thomas, 1912 — originally described from Coto Donana, S Spain, and later merged with meridionalis; also S Portugal. Unless evidence is found that these Italian and Iberian populations are the relics of a much larger formerly contiguous range, this subspecies should be kept as distinct. S. s. coreanus Heude, 1897 — Korean Peninsula. S. s. eristatus Wagner, 1839 — Himalayas S to C India and E to Indochina (N of the Kra Isthmus). S. s. davidi Groves, 1981 — the arid zone from E Iran to Gujarat, including Pakistan and NW India, and perhaps N to Tajikistan. S. s. leucomystax Temminck, 1842 — main Is ofJapan (Honshu, Shikoku, Kyushu, Nakadori, Hiburijima, Tojima, Kushima, and other smaller Is). S. s. lybicus Gray, 1868 — Bulgaria, Greece, Turkey, Syria, Jordan, Israel, Palestine, in the past also in Lybia, and Egypt. The former Yugoslavia was included in its range, which would suggest that now Slovenia, Serbia, Croatia, Bosnia and Herzegovina, Montenegro, and Kosovo are within the range of this subspecies, although the exact boundaries are unclear. Pigs from Albania have been assigned to S. s. scrofa. S. s. majori De Beaux & Festa, 1927 — C & S Italian Peninsula. S. s. menidionalis Forsyth Major, 1882 — Corsica and Sardinia, with the proviso that the two populations are very likely to be introduced or feral. S. s. moupinensis Milne-Edwards, 1871 — China, S to Vietnam and W to Sichuan. S. s. nigripes Blanford, 1875 — the flanks of the Tianshan mountains in Kyrgyzstan and NW China (Xinjiang). An animal photographed in NE Iran (Golestan) looked like this subspecies. S. s. nukiuanus Kuroda, 1924 — Iriomote, Ishigaki, Okinawa, Tokunoshima, Amamioshima, and Kakerome Is in the Ryukyu chain in extreme S Japan, though some of these populations have hybridized with introduced domesticates. S. s. sibiricus Staffe, 1922 — Mongolia and Transbaikal (S & E of Lake Baikal). S. s. tawvanus Swinhoe, 1863 — Taiwan. S. s. ussuricus Heude, 1888 — far E Russia and the Manchurian region (China). Korean populations were previously included in this subspecies, but based on new evidence, the Korean taxon seems more similar to moupinensis. S. s. vittatus Boie, 1828 — Malay Peninsula, S of the Isthmus of Kra, the offshore islands of Terutai and Langkawi, Sumatra, Riau Archipelago, Java, Bali, and a range of smaller islands around these, including Babi, Bakong, Batam, Bawean, Bengkalis, Bintan, Bulan, Bunguran, Cuyo, Deli, Durian, Enggano, Galang, Jambongan, Karimon (Riau Is), Kundur, Lagong, Laut, Lingga, Lingung, Mapor, Moro Kecil, North Pagai, Nias, Panaitan, Payong, Penang, Pinie, Rupat, Siantan, Siberut, Simeulue, Singkep, Sugi, Sugi Bawa, Telibon, Tinggi, Tuangku, and the Tambelan Is. This species was originally present from the British Is in the extreme W, through Eurasia from S Scandinavia to S Siberia, extending as far E as Korea and Japan, and SE into some of the Sunda Is and Taiwan. In the S the species ranged along the Nile Valley to Khartoum, and N of the Sahara in Africa, more orless following the continental coasts of S, E, and SE Asia. Within this range it was absent only from extremely dry deserts, e.g. the driest regions of Mongolia and in China W of Sichuan; and alpine zones, such as the high altitudes of Pamir and Tien Shan. In recent centuries, the range of S. scrofa has changed dramatically because of hunting and changes in available habitat. The species disappeared from the British Is in the 17" century, from Denmark in the 19" century, and was greatly reduced in range and numbers in the 20" century from areas as distant as Tunisia, Sudan, Germany, and Russia. Following these severe declines, there were some slight population recoveries in Russia, Italy, Spain, and Germany in the mid-20™ century, and natural and assisted range expansions in Denmark and Sweden. The species has also been inadvertently reintroduced in various locations in the Great Britain via escapees of mixed origin from commercial farming enterprises. Ex-S. scrofa stocks also occur as introduced feral populations in various other parts of the world, including Australia, New Zealand, the eastern Malay Archipelago, and in North, Central, and South America. In all of these areas they are now generally recognized as a major pest.

Distribution. NE Afghanistan, N Pakistan, NW, N, NE & S India, Nepal, Sri Lanka, N Myanmar, N Vietnam, S, SE & E China, Taiwan, Korean Peninsula, extreme S Russian Far East, and Japan (except Hokkaido); it may occur in Bhutan and Bangladesh. in Miniopteridae

Distribution. NE Afghanistan, N Pakistan, NW, N, NE & S India, Nepal, Sri Lanka, N Myanmar, N Vietnam, S, SE & E China, Taiwan, Korean Peninsula, extreme S Russian Far East, and Japan (except Hokkaido); it may occur in Bhutan and Bangladesh.

Subspecies and Distribution. V. v. vulpes Linnaeus, 1758 — N Europe (Scandinavia). V. v. abietorum Merriam, 1900 — SW Canada (Alberta & British Columbia). V. v. aegyptiacus Sonnini, 1816 — Egypt, Israel, and Lybia. V. v. alascensis Merriam, 1900 — Alaska and NW Canada (NW Territories & Yukon). V. v. alpheraky: Satunin, 1906 — Kazakhstan. V. v. anatolica Thomas, 1920 — Turkey. V. v. arabica Thomas, 1902 — Arabian peninsula. V. v. atlantica Wagner, 1841 — Algeria (forested Atlas Mts). V. v. bangsi Merriam, 1900 — NE Canada (Labrador). V. v. barbara Shaw, 1800 — NW Africa (Barbary Coast). V. v. beringiana Middendorff, 1875 — NE Siberia (shore of Bering Strait). V. v. cascadensis Merriam, 1900 — NW USA (Cascade Mountains, Oregon & Washington). V. v. caucasica Dinnik, 1914 — SW Russia (Caucasus). V. v. crucigera Bechstein, 1789 — Europe through N & C Russia. V. v. daurica Ognev, 1931 — E Russia (Amur, Siberia & Transbaikalia). V.v. deletrix Bangs, 1898 — NE Canada (Newfoundland). V. v. dolichocrania Ognev, 1926 — SE Siberia (S Ussuri). V. v. flavescens Gray, 1843 — N Iran. V. v. fulva Desmarest, 1820 — E USA. V. v. griffith: Blyth, 1854 — Afghanistan and N Pakistan. V.v. harrimani Merriam, 1900 — Alaska (Kodiak I). V. v. hoole Swinhoe, 1870 — S China (Fujian to Sichuan). V. v. ichnusae G. S. Miller, 1907 — Corsica and Sardinia. V. v. induta G. S. Miller, 1907 — Cyprus. V. v. jakutensis Ognev, 1923 — E Siberia (S of Yakutsk). V. v. japonica Gray, 1868 — Japan. V. v. karagan Erxleben, 1777 — Mongolia, Kazakhstan, and Kirgizstan. V. v. kenaiensis Merriam, 1900 — Alaska (Kenai Peninsula). V. v. kurdistanica Satunin, 1906 — Armenia and NE Turkey. V. v. macroura Baird, 1852 — USA (Mountain States). V. v. montana Pearson, 1836 — Himalayas form China (Yunnan) to C Pakistan. V. v. mecator Merriam, 1900 — SW USA (California & Nevada). V. v. ochroxantha Ognev, 1926 — E Russian Turkestan, Aksai, Kirgizstan, Semirechie. V. v. palaestina Thomas, 1920 —Jordan and Lebanon. V.v. peculiosa Kishida, 1924 — Korea. V. v. pusilla Blyth, 1854 — NW India to Irak. V.v. regalis Merriam, 1900 — N Great Plains of Canada and USA. V. v. rubricosa Bangs, 1898 — E Canada. V.v. schrencki Kishida, 1924 — N Japan (Hokkaido) and NE Russia (Sakhalin). V. v. silacea G. S. Miller, 1907 — Iberian Peninsula. V.v. splendidissima Kishida, 1924 — E Russia (N & C Kurile Is). V. v. stepensis Brauner, 1914 — steppes of S Russia. V. v. tobolica Ognev, 1926 — Russia (lower basin of Ob River) V. v. tschiliensis Matschie, 1907 — NE China. Foxes of European origin were introduced into E USA and Canada in the 17" century, subsequently mixed with local subspecies. Also introduced to Australia in 1800s, and the Falkland Islands (Malvinas). in Canidae

Subspecies and Distribution. V. v. vulpes Linnaeus, 1758 — N Europe (Scandinavia). V. v. abietorum Merriam, 1900 — SW Canada (Alberta & British Columbia). V. v. aegyptiacus Sonnini, 1816 — Egypt, Israel, and Lybia. V. v. alascensis Merriam, 1900 — Alaska and NW Canada (NW Territories & Yukon). V. v. alpheraky: Satunin, 1906 — Kazakhstan. V. v. anatolica Thomas, 1920 — Turkey. V. v. arabica Thomas, 1902 — Arabian peninsula. V. v. atlantica Wagner, 1841 — Algeria (forested Atlas Mts). V. v. bangsi Merriam, 1900 — NE Canada (Labrador). V. v. barbara Shaw, 1800 — NW Africa (Barbary Coast). V. v. beringiana Middendorff, 1875 — NE Siberia (shore of Bering Strait). V. v. cascadensis Merriam, 1900 — NW USA (Cascade Mountains, Oregon & Washington). V. v. caucasica Dinnik, 1914 — SW Russia (Caucasus). V. v. crucigera Bechstein, 1789 — Europe through N & C Russia. V. v. daurica Ognev, 1931 — E Russia (Amur, Siberia & Transbaikalia). V.v. deletrix Bangs, 1898 — NE Canada (Newfoundland). V. v. dolichocrania Ognev, 1926 — SE Siberia (S Ussuri). V. v. flavescens Gray, 1843 — N Iran. V. v. fulva Desmarest, 1820 — E USA. V. v. griffith: Blyth, 1854 — Afghanistan and N Pakistan. V.v. harrimani Merriam, 1900 — Alaska (Kodiak I). V. v. hoole Swinhoe, 1870 — S China (Fujian to Sichuan). V. v. ichnusae G. S. Miller, 1907 — Corsica and Sardinia. V. v. induta G. S. Miller, 1907 — Cyprus. V. v. jakutensis Ognev, 1923 — E Siberia (S of Yakutsk). V. v. japonica Gray, 1868 — Japan. V. v. karagan Erxleben, 1777 — Mongolia, Kazakhstan, and Kirgizstan. V. v. kenaiensis Merriam, 1900 — Alaska (Kenai Peninsula). V. v. kurdistanica Satunin, 1906 — Armenia and NE Turkey. V. v. macroura Baird, 1852 — USA (Mountain States). V. v. montana Pearson, 1836 — Himalayas form China (Yunnan) to C Pakistan. V. v. mecator Merriam, 1900 — SW USA (California & Nevada). V. v. ochroxantha Ognev, 1926 — E Russian Turkestan, Aksai, Kirgizstan, Semirechie. V. v. palaestina Thomas, 1920 —Jordan and Lebanon. V.v. peculiosa Kishida, 1924 — Korea. V. v. pusilla Blyth, 1854 — NW India to Irak. V.v. regalis Merriam, 1900 — N Great Plains of Canada and USA. V. v. rubricosa Bangs, 1898 — E Canada. V.v. schrencki Kishida, 1924 — N Japan (Hokkaido) and NE Russia (Sakhalin). V. v. silacea G. S. Miller, 1907 — Iberian Peninsula. V.v. splendidissima Kishida, 1924 — E Russia (N & C Kurile Is). V. v. stepensis Brauner, 1914 — steppes of S Russia. V. v. tobolica Ognev, 1926 — Russia (lower basin of Ob River) V. v. tschiliensis Matschie, 1907 — NE China. Foxes of European origin were introduced into E USA and Canada in the 17" century, subsequently mixed with local subspecies. Also introduced to Australia in 1800s, and the Falkland Islands (Malvinas).

Subspecies and Distribution. F. s. silvestris Schreber, 1777 — Europe E to the Carpathian Mts and the River Dnieper N of the Black Sea. F.s. brockmani Pocock, 1944 — Somalia. F. s. cafra Desmarest, 1822 — Zimbabwe, S Mozambique and South Africa. F. s. caucasica Satunin, 1905 — Caucasus Mts and Turkey. F.s. caudata Gray, 1874 — deserts E Caspian Sea to NW China (Xinjiang) and Mongolia. F. s. foxi Pocock, 1944 — Senegal to Lake Chad. F.s. gordoni Harrison, 1968 — Batinah coast of Oman. F.s. grampia G. S. Miller, 1907 — N Scotland. F. s. griselda Thomas, 1926 — Kalahari region to S Angola. F. s. iraki Cheesman, 1920 — Arabian Desert regions. F. s. lybica Forster, 1780 — desert regions of N Africa to Sudan and N Niger. F.s. melland: Schwann, 1904 — SC Africa. F.s. nesterovi Biurla, 1916 — Mesopotamian region to SW Iran. F. s. ocreata Gmelin, 1791 — Ethiopian highlands. F. s. ornata Gray, 1830 — India. Probably W through Iran. F.s. pyrrhus Pocock, 1944 — N Angola and SW Zaire. F. s. sarda Lataste, 1885 — coastal Maghreb region of Morocco and Algeria. F.s. tristrami Pocock, 1944 — Palestine and Red Sea coast of Arabia. F.s. ugandae Schwann, 1904 — E Africa. in Felidae

Subspecies and Distribution. F. s. silvestris Schreber, 1777 — Europe E to the Carpathian Mts and the River Dnieper N of the Black Sea. F.s. brockmani Pocock, 1944 — Somalia. F. s. cafra Desmarest, 1822 — Zimbabwe, S Mozambique and South Africa. F. s. caucasica Satunin, 1905 — Caucasus Mts and Turkey. F.s. caudata Gray, 1874 — deserts E Caspian Sea to NW China (Xinjiang) and Mongolia. F. s. foxi Pocock, 1944 — Senegal to Lake Chad. F.s. gordoni Harrison, 1968 — Batinah coast of Oman. F.s. grampia G. S. Miller, 1907 — N Scotland. F. s. griselda Thomas, 1926 — Kalahari region to S Angola. F. s. iraki Cheesman, 1920 — Arabian Desert regions. F. s. lybica Forster, 1780 — desert regions of N Africa to Sudan and N Niger. F.s. melland: Schwann, 1904 — SC Africa. F.s. nesterovi Biurla, 1916 — Mesopotamian region to SW Iran. F. s. ocreata Gmelin, 1791 — Ethiopian highlands. F. s. ornata Gray, 1830 — India. Probably W through Iran. F.s. pyrrhus Pocock, 1944 — N Angola and SW Zaire. F. s. sarda Lataste, 1885 — coastal Maghreb region of Morocco and Algeria. F.s. tristrami Pocock, 1944 — Palestine and Red Sea coast of Arabia. F.s. ugandae Schwann, 1904 — E Africa.

Subspecies and Distribution. O. m. manul Pallas, 1776 — Lake Baikal region S through Mongolia to N & NW China. O. m. ferrugineus Ognev, 1928 — Kazakhstan S to Iran and Pakistan, including lowlands S of the Caucasus and W of the Caspian Sea to Armenia. O. m. nigripectus Hodgson, 1842 — Kashmir to Nepal, the Tibetan highlands, and E to C & SW China. in Felidae

Subspecies and Distribution. O. m. manul Pallas, 1776 — Lake Baikal region S through Mongolia to N & NW China. O. m. ferrugineus Ognev, 1928 — Kazakhstan S to Iran and Pakistan, including lowlands S of the Caucasus and W of the Caspian Sea to Armenia. O. m. nigripectus Hodgson, 1842 — Kashmir to Nepal, the Tibetan highlands, and E to C & SW China.

Distribution. Verified records based on analyzed specimens are from NE Afghanistan, N Pakistan (Khyber Pakhtunkhwa and Punjab), and NW & N India (Jammu and Kashmir, Punjab, and Sikkim), but distribution likely includes other regions of NW India (Himachal Pradesh, Uttarakhand) and SW China (S Tibet [= Xizang]); it probably also occurs in Nepal, although further sample comparison is needed. in Vespertilionidae

Distribution. Verified records based on analyzed specimens are from NE Afghanistan, N Pakistan (Khyber Pakhtunkhwa and Punjab), and NW & N India (Jammu and Kashmir, Punjab, and Sikkim), but distribution likely includes other regions of NW India (Himachal Pradesh, Uttarakhand) and SW China (S Tibet [= Xizang]); it probably also occurs in Nepal, although further sample comparison is needed.

Distribution. NC, C & SW China (Tibet [= Xizang], Qinghai, Gansu, Ningxia, and Sichuan), Bangladesh (Sylhet Division), NE India (Meghalaya and Mizoram), and NW Thailand; possibly N Myanmar (although this record has been mapped dubiously). in Vespertilionidae

Distribution. NC, C & SW China (Tibet [= Xizang], Qinghai, Gansu, Ningxia, and Sichuan), Bangladesh (Sylhet Division), NE India (Meghalaya and Mizoram), and NW Thailand; possibly N Myanmar (although this record has been mapped dubiously).

Subspecies and Distribution. E.s.serotinusSchreber,1774—muchofEuropeexceptforSIberianPeninsulaandmostofScandinavia,beingfoundinSBritainandfromN&CIberianPeninsulaEtoextremeSScandinavia,SWEuropeanRussia,Caucasus,andN&ETurkeyalongwithnumerousMediterraneanIs(BalearicIs,Corsica,Sardinia,andSicily),includingIonian(Corfu)andAegeanIs(Samothrace,Lesbos,Skyros,Euboea,Samos,Crete,andRhodes). E.s.mirzadeFilippi,1865—STurkey,WSyria,Lebanon,Israel,andNW,C&SWIran. E. s. turcomanus Eversmann, 1840 — Central Asia in SC Russia, Kazakhstan, Uzbekistan, Turkmenistan, N Iran, Kyrgyzstan, Tajikistan, N Afghanistan, and NW China (Xinjiang). in Vespertilionidae

Subspecies and Distribution. E.s.serotinusSchreber,1774—muchofEuropeexceptforSIberianPeninsulaandmostofScandinavia,beingfoundinSBritainandfromN&CIberianPeninsulaEtoextremeSScandinavia,SWEuropeanRussia,Caucasus,andN&ETurkeyalongwithnumerousMediterraneanIs(BalearicIs,Corsica,Sardinia,andSicily),includingIonian(Corfu)andAegeanIs(Samothrace,Lesbos,Skyros,Euboea,Samos,Crete,andRhodes). E.s.mirzadeFilippi,1865—STurkey,WSyria,Lebanon,Israel,andNW,C&SWIran. E. s. turcomanus Eversmann, 1840 — Central Asia in SC Russia, Kazakhstan, Uzbekistan, Turkmenistan, N Iran, Kyrgyzstan, Tajikistan, N Afghanistan, and NW China (Xinjiang).

Distribution. NW Thailand (Chiang Mai Province), N Laos, N & C Vietnam, and S China (Guizhou); it may occur in Cambodia. in Vespertilionidae

Distribution. NW Thailand (Chiang Mai Province), N Laos, N & C Vietnam, and S China (Guizhou); it may occur in Cambodia.

Subspecies and Distribution. N. n. noctula Schreber, 1774 — throughout Europe from Great Britain, France, and Spain E to W Russia, W Kazakhstan, and SW Turkmenistan, including S Scandinavia, Gotland and Oland Is, and Cyprus (Cyprus records somewhat tentatively regarded as this subspecies). Absent throughout much of Iberia and is locally extinct in Portugal. N. n. lebanoticus D. L.. Harrison, 1962 — WC & SW Syria, Lebanon, and NE Israel. N. n. mecklenburzevi Kuzyakin, 1934 — SC & E Kazakhstan, SC Russia, W Uzbekistan, Tajikistan, Kyrgyzstan, and NW China (Xinjiang). The species may be present in N Africa, with two records claimed from Algeria in 1858, but these may represent N. lasiopterus; further sampling is needed. in Vespertilionidae

Subspecies and Distribution. N. n. noctula Schreber, 1774 — throughout Europe from Great Britain, France, and Spain E to W Russia, W Kazakhstan, and SW Turkmenistan, including S Scandinavia, Gotland and Oland Is, and Cyprus (Cyprus records somewhat tentatively regarded as this subspecies). Absent throughout much of Iberia and is locally extinct in Portugal. N. n. lebanoticus D. L.. Harrison, 1962 — WC & SW Syria, Lebanon, and NE Israel. N. n. mecklenburzevi Kuzyakin, 1934 — SC & E Kazakhstan, SC Russia, W Uzbekistan, Tajikistan, Kyrgyzstan, and NW China (Xinjiang). The species may be present in N Africa, with two records claimed from Algeria in 1858, but these may represent N. lasiopterus; further sampling is needed.

Distribution. NE Uzbekistan, N Kyrgyzstan, SE Kazakhstan, and NW China. Occurs in the W of Tianshan Mt System (except W Karatau), just E of the cities ofJambul and Shimkent in Kazakhstan and the city of Tashkent in Uzbekistan, and extends E through Tianshan Mts (excluding inner Tianshan) surrounding Issyk-Kul Lake and Jungarian Alatau, and in China at the edge of the distribution of the Gobi Argali (O. darwini) on the Chinese-Mongolian border. in Bovidae

Distribution. NE Uzbekistan, N Kyrgyzstan, SE Kazakhstan, and NW China. Occurs in the W of Tianshan Mt System (except W Karatau), just E of the cities ofJambul and Shimkent in Kazakhstan and the city of Tashkent in Uzbekistan, and extends E through Tianshan Mts (excluding inner Tianshan) surrounding Issyk-Kul Lake and Jungarian Alatau, and in China at the edge of the distribution of the Gobi Argali (O. darwini) on the Chinese-Mongolian border.