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Figs 40–52 in On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm.

Figs 1–26 in On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm.

Figs 53–71 in On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm.

Figs 260–277 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 260–277: Zyras morulus (260–265), Z. variolatus (266–270), Z. montanus (271–275), and Z. preangeranus from Java (276–277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263–264, 266–267, 271–272, 276–277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260–262: 1.0 mm; 263–277: 0.5 mm.

Figs 208–226 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 208–226: Zyras parvicollis (208–211), Z. ambulans (212–215), Z. nitens, holotype (216–221), and Z. gratellus (Malaysia) (222–226): median lobe of aedeagus in lateral and in ventral view (208–213, 216–217, 222–223); paramere (214, 218, 224); male tergite VIII (219, 225); male sternite VIII (220, 226); apical lobe of paramere (215); postero–median portion of male sternite VIII (221). Scale bars: 208–214, 216–220, 222–226: 0.5 mm; 215, 221: 0.1 mm.

Figs 227–244 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm.

Figs 191–207 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm.

Figs 156–173 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 156–173: Zyras longilobatus (156–160), Z. latilobatus (161–165), Z. alternans (166–169), and Z. hirtus (170–173; 170–171: Sri Lanka; 172–173: South India): median lobe of aedeagus in lateral and in ventral view (156–157, 161–162, 166–167, 170–173); paramere (158, 163); male tergite VIII (159, 164, 168); male sternite VIII (160, 165, 169). Scale bars: 0.5 mm.

Figs 174–190 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 174–190: Zyras gardneri (174–177), Z. hirsutiventris (178–181), Z. luteipes (182–186), and Z. russiceps (187–190): median lobe of aedeagus in lateral and in ventral view (174–175, 178–179, 182–183, 187–188); male tergite VIII (176, 180, 185, 189); male sternite VIII (177, 181, 186, 190); paramere (184). Scale bars: 0.5 mm.

Figs 117–133 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm.

Figs 134–155 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 134–155: Zyras perforatus (134–135), Z. pallipes (136–139), Z. nigroaeneus, holotype (140–141), Z. condignus (142–147), Z. morvani, holotype (148–149), and Z. truncatus (150–155): male tergite VIII (134, 138, 140, 146, 148, 152); male sternite VIII (135, 139, 141, 147, 149, 153); median lobe of aedeagus in lateral and in ventral view (136–137, 142–143, 150–151); paramere (144–145); female tergite VIII (154); female sternite VIII (155). Scale bars: 0.5 mm.

FIGURE 6 in Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

FIGURE 6. Variations of tHe spermatHeCa of Zyras (Diaulaconia) artemis sp. n. LoCality of eaCH speCimens: A, Gaupo (Taoyuan Co.); B, Hsilin trail (Hualien Co.); C, DaHasHan (Pingtung Co.); D, E, Dakeng (TaiCHung City); F, Xiaping (Nantou Co.); G, H, LuansHan (Taitung Co.); I, SHeding (Pingtung Co.); J, K, PengHu Islands (PengHu Co.).

FIGURE 5 in Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

FIGURE 5. Zyras (Diaulaconia) artemis sp. n., female. A, eigHt abdominal tergite, dorsal vieW; B 8th abdominal sternite, ventral vieW; C, spermatHeCa; D, tHe pHotograpH of tHe apiCal part and spermatHeCal gland; E, tHe pHotograpH of tHe inner Wall of apiCal 1/3 of tHe basal part; F, tHe base of tHe basal part and tHe spermatHeCal duCt.

FIGURE 3 in Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

FIGURE 3. Zyras (Diaulaconia) artemis sp. n. A, male 4th tergite WitH a pair of projeCtions on posterior margin, large individual, dorsal vieW; B, ditto, projeCtion bilobed apiCally, lateral vieW; C, ditto, small individual, dorsal vieW; D, ditto, projeCtion WitH no bilobed apiCally, lateral vieW; E, male 7th tergite WitH apiCal median tuberCle, dorsal vieW; F, ditto, lateral vieW; G, female 7th tergite WitH no small median tuberCle, dorsal vieW.

FIGURE 2 in Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

FIGURE 2. Zyras (Diaulaconia) artemis sp. n. A, forebody, dorsal vieW; B, rigHt antenna, lateral vieW; C, ditto, dorsal vieW; D, frons of tHe female WitHout granulations; E, frons of tHe male WitH granulations.

FIGURE 4 in Zyras (Diaulaconia) artemis sp. n., a new termitophilous species from Taiwan (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

FIGURE 4. Zyras (Diaulaconia) artemis sp. n., male. A, eigHt abdominal tergite, dorsal vieW; B, 8th abdominal sternite, ventral vieW; C, median lobe of aedeagus, ventral vieW; D, ditto, tHe pHotograpH; E, ditto, lateral vieW; F, ditto, tHe pHotograpH.

Map 8 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Map 8: Revised distributions of Zyras alternans (black circles), Z. luteipes (star), Z. brignolii (white circles), and Z. nitens (white

Map 7 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Map 7: Revised distributions of Zyras gardneri (black circles), Z. hirsutiventris (white diamonds), Z. parageminus (black triangles), Z. hastatus (white triangle), Z. hirtus (white circles), and Z. latilobatus (star) in India and adjacent regions.

Figs 1–41 in On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 1–41: Antennae of Zyras spp.: kraatzi (1), exasperatus (2), pindarae (3–5), perforatus (6), championi (7), pallipes (8), condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27), proximus (28–29), novinversus (30–31), brevilobatus (32), parvicollis (33), ambulans (34), matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm.

Figs 27–39 in On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)

Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm.