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Text-fig. 5. Anacardiaceae (a–m), Burseraceae (n–q). Scale bars = 1 cm. a–f: Pentoperculum sp. a–c: USNM PAL 772360. a: Lateral view of endocarp, apex up; three germination valves visible, the central clearly displaying the bipartite nature of the valve, reflected light, palladium coated. b: Apical view displaying six locules, with two preserved germination valves at the lower left, reflected light, palladium coated. c: Basal view of the endocarp, the locules suggested by swellings; note point of attachment, micro-CT scan surface rendering. d–f: Pentoperculum sp. USNM PAL 772359, reflected light, palladium coated. d: Lateral view of a probable 6-loculed endocarp, apex up; a single intact germination valve in the center, displaying the central lineation that divides it in two. e: Apical view; two bi-partite germination valves are visible, indicated by arrows to the middle cleavage line of two of the valves. f: Basal view, the locules suggested by the undulations in the margin. g–i: Indet. Spondioideae. USNM PAL 772358, reflected light, palladium coated. g: Lateral view of multi-locular endocarp, apex up. h: Apical view showing finely punctuate surface and peripheral locule cavities. i: Basal view. j–m: Cf. Pleiogynium USNM PAL 772357. j: Lateral view of the multi-locular endocarp, apex up; note intact germination valve on left and exposed locule facing the viewer, micro-CT scan surface rendering. k: Lateral view, rotated about 30° from (j), showing three exposed locules, reflected light, palladium coated; note bipartite locule lining at center. l: Apical view of the multilocular endocarp; the locule with intact germination valve at the upper right, reflected light, palladium coated. Arrows to each locule. m: Basal view showing central point of attachment and prominent radiating ridges aligned with the locules, micro-CT scan surface rendering. n–q: Canarium, USNM PAL 772361. Scale bar = 1 cm. n: Lateral view of endocarp directly facing one germination valve flanked by two strong ridges; apex up; specimen coated in sodium nitrate and photographed by R. A. Scott. o: Lateral view facing one of the three pronounced ridges, flanked to the left and right by two germination valves; apex up. p: Apical view displaying the three strong ridges, arching over the apex and flanking three deep embayments, covered with germination valves. q: Basal view, the three ridges being less pronounced. o–q: Reflected light, palladium coated. in The Early Middle Eocene Wagon Bed Carpoflora Of Central Wyoming, U.S.A.
Text-fig. 5. Anacardiaceae (a–m), Burseraceae (n–q). Scale bars = 1 cm. a–f: Pentoperculum sp. a–c: USNM PAL 772360. a: Lateral view of endocarp, apex up; three germination valves visible, the central clearly displaying the bipartite nature of the valve, reflected light, palladium coated. b: Apical view displaying six locules, with two preserved germination valves at the lower left, reflected light, palladium coated. c: Basal view of the endocarp, the locules suggested by swellings; note point of attachment, micro-CT scan surface rendering. d–f: Pentoperculum sp. USNM PAL 772359, reflected light, palladium coated. d: Lateral view of a probable 6-loculed endocarp, apex up; a single intact germination valve in the center, displaying the central lineation that divides it in two. e: Apical view; two bi-partite germination valves are visible, indicated by arrows to the middle cleavage line of two of the valves. f: Basal view, the locules suggested by the undulations in the margin. g–i: Indet. Spondioideae. USNM PAL 772358, reflected light, palladium coated. g: Lateral view of multi-locular endocarp, apex up. h: Apical view showing finely punctuate surface and peripheral locule cavities. i: Basal view. j–m: Cf. Pleiogynium USNM PAL 772357. j: Lateral view of the multi-locular endocarp, apex up; note intact germination valve on left and exposed locule facing the viewer, micro-CT scan surface rendering. k: Lateral view, rotated about 30° from (j), showing three exposed locules, reflected light, palladium coated; note bipartite locule lining at center. l: Apical view of the multilocular endocarp; the locule with intact germination valve at the upper right, reflected light, palladium coated. Arrows to each locule. m: Basal view showing central point of attachment and prominent radiating ridges aligned with the locules, micro-CT scan surface rendering. n–q: Canarium, USNM PAL 772361. Scale bar = 1 cm. n: Lateral view of endocarp directly facing one germination valve flanked by two strong ridges; apex up; specimen coated in sodium nitrate and photographed by R. A. Scott. o: Lateral view facing one of the three pronounced ridges, flanked to the left and right by two germination valves; apex up. p: Apical view displaying the three strong ridges, arching over the apex and flanking three deep embayments, covered with germination valves. q: Basal view, the three ridges being less pronounced. o–q: Reflected light, palladium coated.
FIGURE 17 in Diptera of the Middle Eocene Kishenehn Formation II
FIGURE 17. Palaeoapsilocephala kishenehnensis Hauser and Greenwalt sp. n., USNM 717195. A, Hind tibia spurs; B, Genitalia. C, Drawing of genitalia. Scale bars equal 0.5 mm (A, B).
FIGURE 9 in Middle Eocene cartilaginous fishes (Vertebrata: Chondrichthyes) of the Dnieper-Donets Basin, northern Ukraine
FIGURE 9. Teeth of Physogaleus secundus (Winkler, 1876) from the middle Eocene deposits of Vyshhorod: A – upper lateral tooth NMNHU-G 391/24/2; B – lower anterolateral tooth NMNHU-G 391/24/1; C – lower lateral tooth NMNHU-G 391/24/5, all in lingual view.
FIGURE 3. Zircon U in Geochronology of the middle Eocene Purple Bench locality (Devil's Graveyard Formation), Trans-Pecos Texas, USA
FIGURE 3. Zircon U-Pb results from tuffaceous sandstones of the Devil's Graveyard Fm. analyzed via LAICP-MS. Results were filtered to exclude single grain ages with>15% discordance between 235U/207Pb age and 238U/206Pb age, or>10% propagated analytical error in 238U/206Pb age. Weighted means were calculated for each sample using Tuff extractor plugin from the ISOPLOT package (Ludwig and Mundil, 2002; Ludwig, 2003). Number of grains used in TuffZirc algorithm displayed as n= # of ages used by TuffZirc / # of ages total for sample. Data are displayed as kernel density estimators (KDEs) with a bandwidth of 1 Myr (Vermeesch, 2018). Individual zircon ages and 2s standard errors are displayed as vertically stacked boxes. Individual ages in red were used in the calculation of the weighted mean. Weighted means and errors are displayed as a grey box with a bar representing the mean.
FIGURE 4 in Geochronology of the middle Eocene Purple Bench locality (Devil's Graveyard Formation), Trans-Pecos Texas, USA
FIGURE 4. Comparison of new and published age estimates for the Devil's Graveyard Fm. and the Purple Bench tuff. Walton (1992) placed the Purple Bench tuff at the C20R-C20N chron transition (43.43 Ma; recalibrated to GST2012; Gradstein et al., 2012), while Prothero (1996) placed the Purple Bench tuff within the C18R chron (41.15- 40.14 Ma; recalibrated to GST2012; Gradstein et al., 2012).
Fig. 3. A. Zygodactylid bird Primozygodactylus major Mayr, 1998 in New specimens of zygodactylid birds from the middle Eocene of Messel, with description of a new species of Primozygodactylus
Fig. 3. A. Zygodactylid bird Primozygodactylus major Mayr, 1998, selected bones of the newly referred specimen SMF−ME 799a (A4) and SMF−ME 799b (A1–A3, A5) from the middle Eocene of Messel in Germany. A1, atlas and axis; A2, third and fourth cervical vertebrae; A3, furcula; A4, distal end of right tarsometatarsus in plantar view; A5, right coracoid in dorsal view in comparison to (B) Zygodactylus luberonensis (SMF Av 519) from the early Oligocene of the Lubéron in France. All specimens coated with ammonium chloride.
FIGURE 11. A-C in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 11. A-C. Ouledia lacuna nov. sp. A.?Anterior tooth KEB 1-180, A1. Occlusal view, A2. Lingual view, A3. Magnificence of crown-root boundary in A2, A4. profile; B.?lateral tooth KEB 1-181, B1. Occlusal view, B2. labial view, C.?Anterior tooth KEB 1-182, C1. Labial view, C2. Basal view; D-H. Pachygymnura attiai nov. gen. D. Anterior tooth KEB 1-183, D1. Lingual view, D2. Occlusal view, D3. Labial view, D4. Basal view; E. Antero-lateral tooth KEB 1-184, E1. Lingual view, E2. Near labial view, F. lateral tooth KEB 1-185, F1. Occlusal view, F2. Profile, F3. Basal view; G. lateral tooth KEB 1-186, G1. Lingual view, G2. Occlusal view; H. Anterior tooth KEB 1-187, H1. Profile, H2. Occlusal view.
FIGURE 7. A-C in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 7. A-C: Hemipristis curvatus. A. Antero-lateral upper tooth KEB 1-133, A1. Lingual view, A2. Labial view; B. Lateral upper tooth of young specimen KEB 1-134, labial view; C. Anterior lower tooth of young specimen KEB 1-135, labial view; D-E: Moerigaleus sp. D. Lateral upper tooth KEB 1-136, D1. Lingual view, D2. Labial view; E. Lateral upper tooth KEB 1-137, E1. Lingual view, E2. Labial view; F-K: Leptocharias tunisiensis nov. sp. F. (HOLOTYPE) Antero-lateral lower tooth KEB 1-138, F1. Profile, F2. Lingual view, F3. Labial view; G. Antero-lateral upper tooth KEB 1-139, G1. Labial view, G2. Profile, G3. Lingual view; H. Anterro-lateral lower tooth KEB 1-140, labial view; I. Lateral lower tooth KEB 1-141, labial view; J. More lateral tooth KEB 1-142, J1. Labial view, J2. Lingual view, J3 occlusal view; K. Posterior tooth KEB 1-143, K1. Labial view, K2. Lingual view.
FIGURE 5. A-K in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 5. A-K: Sphyrna guinoti nov. sp. A. Anterior upper tooth KEB 1-109, A1. Labial view, A2. Lingual view; B. Lateral upper tooth KEB 1-110, B1. Labial view, B2. Lingual view; C. Anterior upper tooth KEB 1-111, C1. Lingual view, C2. Labial view; D. Lateral lower tooth KEB 1-112, D1. Labial view, D2. Lingual view; E. lateral upper tooth KEB 1-113, E1. Lingual view, E2. Labial view; F. Lateral upper tooth KEB 1-114, F1. Labial view, F2. Lingual view; G. Posterior upper tooth KEB 1-115, lingual view; H. Anterior lower tooth KEB 1-116, H1. Labial view, H2. Lingual view; I. Antero-lateral lower tooth KEB 1-117, I1. Labial view, I2. Lingual view; J. (HOLOTYPE) Antero-lateral lower tooth KEB 1-118, J1. Labial view, J2. Lingual view; K. Lateral lower tooth KEB 1-119, K1. Labial view, K2. Lingual view.
FIGURE 3. A-N in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 3. A-N: Carcharhinus kasserinensis nov. sp. A. Parasymphyseal upper tooth KEB 1-086, A1. labial view, A2. Lingual view; B. HOLOTYPE Anterior upper tooth KEB 1-087, B1. Labial view, B2. Lingual view, B3. Magnificence of mesial cutting edge; C. Antero-lateral upper tooth KEB 1-088, C1. Labial view, C2. Lingual view; D. Antero-lateral upper tooth KEB 1-089, D1. Labial view, D2. Lingual view; E. Antero-lateral upper tooth KEB 1-226, E1 Lingual view, E2. Labial view; F. Antero-lateral upper tooth KEB 1-227; G. Antero-lateral upper tooth KEB 1-090, G1. Labial view, G2. Lingual view; H. Lateral upper tooth KEB 1-091, labial view; I Posterior upper tooth KEB 1-092, lingual view; J. Symphyseal abnormal lower tooth KEB 1-093, J1. Labial view, J2. Lingual view; K. Anterior lower tooth KEB 1-094, labial view; L. Anterior lower tooth KEB 1-095, L1. Labial view, L2. Lingual view; M. Antero-lateral lower tooth KEB 1- 096, M1. Labial view, M2. Lingual view; N. Antero-lateral lower tooth KEB 1-097, N1. Lingual view, N2. Labial view, N3. Profile.
FIGURE 2 in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 2. Locality of KEB-1, Djebel el Kébar, Kasserine region, Tunisia. A, Simplified topographic map of Tunisia locating the Djebel el Kébar in central Tunisia; B, simplified stratigraphical position of fossiliferous level having yielded the fossil-bearing KEB-1 locality; C, photograph showing the typical badlands of variegated clays from the early Tertiary sequence of Djebel el Kébar. See Merzeraud et al. (2016) for precise details about geological settings.
FIGURE 8. A-C in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 8. A-C: Nebrius sp. A. Anterior tooth KEB 1-144, A1. Labial view, A2. Lingual view, A3. profile; B. Antero-lateral tooth KEB 1-145, B1. Labial view, B2. Basal view; C. Lateral tooth KEB 1-146, C1. Labial view, C2. Lingual view; D-G: Stegostoma tethysiensis nov. sp. D. (HOLOTYPE) Anterior tooth KEB 1-147, D1. Labial view, D2. Lingual view, D3. Basal view; E. Antero-lateral tooth KEB 1-148, E1. Labial view, E2. Occlusal viex, E3. profile; F. Lateral tooth KEB 1-149, labial view; G. Posterior tooth KEB 1-150, labial view; H: Hemiscyllium sp. 8. Antero-lateral tooth KEB 1-151, labial view; I-N: Odontorythys pappenheimi I.?Anterior tooth KEB 1-152, I1. Profile, I2. Labial view; J.?Latero-posterior tooth KEB 1-153, lingual view; K.?Latero-posterior tooth KEB 1-154, profile; L.?Latero-posterior tooth KEB 1-155, profile; M.?Latero-posterior tooth KEB 1-156, profile; N.? lateral or posterior tooth KEB 1-157, N1. Profile, N2. Labial view, N3. Lingual view.
FIGURE 13. A-E in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 13. A-E. Coupatezia cristata nov. sp. A. antero-lateral tooth KEB 1-205, HOLOTYPE, A1.occlusal view, A2. Profile, A3. Labial view; B. Anterior tooth KEB 1-206, B1. Lingual view, B2. Basal view, C. lateral tooth KEB 1-207, C1. Occlusal view, C2. Magnificence of C1, C3. Lingual view; D. lateral tooth KEB 1-208, D1. Occlusal view, D2. Magnificence of D1, D3. Lingual view; E. lateral tooth KEB 1-209, E1. Occlusal view, E2., lingual view, E3. Basal view. F-G. Rhinoptera sp. F. medium tooth KEB 1-210, lingual view, G. medium tooth KEB 1-211, labial view; H-J. Myliobatis sp., H. lateral tooth KEB 1-212, lingual view, I. medio-lateral tooth KEB 1-213, lingual view, J lateral tooth KEB 1-214, lingual view; K. Aetobatus sp. KEB 1-215, ligual view, L-M. Garabatis sp., L. KEB 1-216, occlusal view, M. KEB 1-217, L1. occlusal view, L2. Magnificence of central part, L3. Magnificence of marginal part of crown.
FIGURE 6. A-C in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 6. A-C: Sphyrna guinoti nov. sp. A. Anterior upper tooth of Young specimen KEB 1-120, A1. Lingual view, A2. Labial view; B. Lateral lower tooth of Young specimen KEB 1-121, B1. Labial view, B2. Lingual view; C. Lateral upper tooth KEB 1-122, C1. Labial view, C2. Lingual view; D-F:?Sphyrna sp. 4. D.?Antero-lateral upper tooth KEB 1-123, D1. Lingual view, D2. Labial view; E.?Antero-lateral upper tooth KEB 1-124, E1. Labial view, E2. Lingual view; F. Lateral upper tooth KEB 1-125, lingual view.
FIGURE 15 in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 15. Distribution of Elasmobranch taxa within localities of KEB-1, Tunisia (KEB, this work) and El Gedida (EG), Baharia (Egypt, from Strougo et al., 2007 updated) with report of their occurences in the other late Middle – Late Eocene tropical assemblages from Egypt (MI, GE, BQ, QS). Frequencies of taxa are somewhat subjective due to the different sources and/or sampling methods applied. Black-Grey intensity indicating the fossil completeness between KEB/EG and the other localities due to uncertainty/resolution of taxonomic attributions (See systematic palaeontology for detail, black: certain to light: possible). (1) Medium-Large Carcharhinus sp. with serrated teeth, (2) as Misrichtys stromeri, (3) as Moerigaleus vitreodon, (4) as Nebrius blankenhormi, (5) as Nebrius sp., (6) refered to Odontorhytis bahariensis, (7) unnnamed species, (8) possibly as Taeniura sp., (9) two types in Fayum.
FIGURE 4. A-B in Diversity and renewal of tropical elasmobranchs around the Middle Eocene Climatic Optimum (MECO) in North Africa: New data from the lagoonal deposits of Djebel el Kébar, Central Tunisia
FIGURE 4. A-B: Carcharhinus kasserinensis nov. sp., A. Antero-lateral upper tooth of young specimen KEB 1-098, A1. Lingual view, A2. Labial view; B. Antero-lateral upper tooth of young specimen KEB 1-099, labial view. C-G: Carcharhinus frequens. C. Lateral upper tooth KEB 1-100, C1. Labial view, C2. Lingual view; D. Lateral upper tooth KEB 1-101, D1. Labial view, D2. Lingual view; E. Anterior lower tooth KEB 1-102, E1. Labial view, E2. Lingual view; F. Lateral lower tooth KEB 1-103, F1. Labial view, F2. Lingual view; G. Posterior lower tooth KEB 1-104, lingual view; HK: Misrichtys sp. H. Anterior lower tooth KEB 1-105, lingual view; I. Lateral upper tooth KEB 1-106, labial view; J. Posterior lower tooth KEB 1-107, j1. Labial view, J2. Lingual view; K. Lateral upper tooth KEB 1-108, labial view; L-R: Rhizoprionodon sp. L. Anterior upper tooth KEB 1-126, labial view; M. lateral upper tooth KEB 1-127, labial view; N. Lateral upper tooth KEB 1-128, labial view; O. Anterior lower tooth KEB 1-129, labial view; P. Lateral lower tooth KEB 1-130, P1. Labial view, P2. Lingual view; Q. Posterior lower tooth KEB 1-131, labial view; R. Antero-lateral lower tooth KEB 1-132, R1. Labial view, R2. Lingual view.
Text-fig. 1. Geological map of the site Kučlín, Trupelník Hill. 1 – dumpsite body, exploitation faces, 2 – landslide body, 3 – occurrence of the basalt in the landslide body, 4 – tuff, tuffite with gneiss particles, 5 – 11 – Late Eocene rocks. 5 – basalts, 6 – tuff, tufite, 7 – diatomaceous cherts layer enclosing lenses of diatomaceous earth, 8 – diatomitic breccia, 9 – diatomaceous shale, 10 – diatomaceous clays to diatomites, 11 – calcareous diatomaceous clays to diatomaceous limestones and re-deposited Cretaceous material, 12 – Late Turonian marlstone to limestone, 13 – Middle Turonian marlstone, 14 – Middle Turonian organo-detritic limestone to conglomerate, 15 – Late Proterozoic orthogneiss, 16 – Syčivka creek, 17 – road, field drives, 18 – Kučlín village. in Geology Of The Site Kučlín, Trupelník Hill Near Bílina In North Bohemia
Text-fig. 1. Geological map of the site Kučlín, Trupelník Hill. 1 – dumpsite body, exploitation faces, 2 – landslide body, 3 – occurrence of the basalt in the landslide body, 4 – tuff, tuffite with gneiss particles, 5 – 11 – Late Eocene rocks. 5 – basalts, 6 – tuff, tufite, 7 – diatomaceous cherts layer enclosing lenses of diatomaceous earth, 8 – diatomitic breccia, 9 – diatomaceous shale, 10 – diatomaceous clays to diatomites, 11 – calcareous diatomaceous clays to diatomaceous limestones and re-deposited Cretaceous material, 12 – Late Turonian marlstone to limestone, 13 – Middle Turonian marlstone, 14 – Middle Turonian organo-detritic limestone to conglomerate, 15 – Late Proterozoic orthogneiss, 16 – Syčivka creek, 17 – road, field drives, 18 – Kučlín village.
Fig. 2 in Tarsal morphology of the pleuraspidotheriid mammal Hilalia from the middle Eocene of Turkey
Fig. 2. Hilalia saribeya Maas, Thewissen, Sen, Kazanci, and Kappelman, 2001 from the middle Eocene Uzunçarşıdere Formation, Central Turkey; right astragalus (EOU-UCF-1) in proximal (A), plantar (B), dorsal (C), medial (D), and lateral (E) views. Each homologous articular facet is marked by dashed line.
FIGURE 23. Claiborne Fagaceae Fruit Type 5 in Fruits, seeds and flowers from the Puryear clay pit (middle Eocene Cockfield Formation), western Tennessee, USA
FIGURE 23. Claiborne Fagaceae Fruit Type 5: 1) UF15820-004015, fruit showing branched spines (indicated by arrows); 2) UF15820-004025, fruit showing two superimposed valves with clay filled in between them (indicated by lower left arrows) and branching and intertwining spines (indicated by top arrow); 3) UF15820- 032853, fruit showing valves with distal portions of spines missing and only proximal web-like valve preserved (indicated by arrow). All scale bars equal 3 mm.
Text-fig. 9. Bivariate graph (length/width) of lower teeth of Masillamys mattaueri (HARTENBERGER) from Mas de Gimel (Hérault, France; MP 10, late early Eocene). in A Reevaluation Of The Taxonomic Status Of The Rodent Masillamys Tobien, 1954 From Messel (Germany, Late Early To Early Middle Eocene, 48-47 M.Y.)
Text-fig. 9. Bivariate graph (length/width) of lower teeth of Masillamys mattaueri (HARTENBERGER) from Mas de Gimel (Hérault, France; MP 10, late early Eocene).
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