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FIGURE 7. Erenna richardi Bedot, 1904. A in A description of two new species of the genus Erenna (Siphonophora: Physonectae: Erennidae), with notes on recently collected specimens of other Erenna species

FIGURE 7. Erenna richardi Bedot, 1904. A. Separate large nematocysts of cnidoband. B. Large nematocysts forming lateral margins of cnidoband. C & D. Smaller nematocysts from central region of cnidoband. See text for details. Scale bars A, B, 100 µm; C, D. 50 µm.

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FIGURE 6. Chnoodes tarsalis Weise, 1904. A–D in Two new species of Chnoodes Chevrolat (Coleoptera: Coccinellidae) from Brazil

FIGURE 6. Chnoodes tarsalis Weise, 1904. A–D. habitus: A. dorsal; B. ventral; C. frontal view; D. lateral. E–G. male genitalia: E. tegmen, ventral; F. tegmen, lateral; G. sipho lateral view. H–I. female genitalia; H. coxites; I. spermatheca.

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FIGURE 6 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 6. Individuals of Euthygomphus yunnanensis in life: a—mature male, Cambodia, Stung Treng Province, Thala Barivat District, 27 vii 2016; b—teneral female, Cambodia, Koh Kong Province, Thma Bang River 5 km SW Thma Bang village, 366 m a.s.l., 15 iv 2010; c—teneral male, Cambodia, Pursat Province, a river at O'Som village, 12°04'40–45'' N 103°12'21–35'', 524–531 m a.s.l., 24 iii 2015. Not to scale.

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FIGURE 7 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 7. Wings of Euthygomphus yunnanensis (a—male, from Cambodia, Mondulkiri Province, 'Loringae' brook downstream of Buu Sraa Waterfalls, b—female from Cambodia, Koh Kong Province, 'Microgomphus River' 6.5 km SW Thma Bang village) and A. (E.) schorri (c—male holotype, from Cambodia, Mondulkiri Province, 3.5–3.8 km ESE of Sen Monorom, 'Culminicola Rivulet; d—paratype female 2 from Cambodia, Mondulkiri Province, downstream of Buu Sraa Waterfalls). Not to scale; c mirrored.

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FIGURE 5 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 5. Details of Cambodian specimens of Euthygomphus yunnanensis: a,e,g,k–m—male from Mondulkiri Province, 'Loringae' brook downstream of Buu Sraa Waterfalls; b–d,f–j,n—females: b–c,f,i–j,n—from the same locality, d,h—from Koh Kong Province, 'Microgomphus River' 6.5 km SW Thma Bang village; a–d—thorax, lateral view; e–f—head, frontal view; g– i—occiput, fromtal view; j—ditto, dorsal view; k—male anal appendages, dorsal view; l—ditto, lateral view; m—ditto, ventral view; n—female vulvar scale, ventral view. Not to scale; b,l mirrored.

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FIGURE 10 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 10. Head and thorax of Euthygomphus schorri; a–d—holotype male; e–i—paratype female 2; j—paratype female 1 from the same locality as the holotype; a,f—head and thorax, lateral view; b,g—synthorax, dorsal view; c,h—head, frontal view; d,i—occiput, frontal view; e,j—occipital horns, posterior view. Not to scale.

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FIGURE 4 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 4. Details of secondary genitalia of males of Euthygomphus yunnanensis from Cambodia: a–c—from Mondulkiri Province, 'Loringae' brook downstream of Buu Sraa Waterfalls; d–f—from Koh Kong Province, 'Microgomphus River' 6.5 km SW Thma Bang village; g–k—from Pursat Province, O'Som village; a—general lateral view; b—hamuli, lateral view; d, anterior hamulus, lateral view; e, posterior hamulus, oblique view; c—penis distal segment (V4), lateral view; f–g—ditto, ventral view; i—ditto, lateral view; j—ditto, dorsolateral view; h—penis (vesica spermalis), lateral view; k—ditto, ventral view. Not to scale.

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FIGURE 12 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 12. Habitats of Euthygomphus schorri: a—the type locality, Cambodia, Mondulkiri Province, 3.5–3.8 km ESE of Sen Monorom, 'Culminicola Rivulet', 689–691 m a.s.l.,; b—the other locality, Cambodia, Mondulkiri Province, downstream of Buu Sraa Waterfalls, 'Loringae brook' just downstream of its own waterfall, 490 m a.s.l., which is also a habitat of Euthygomphus yunnanensis.

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FIGURE 3 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 3. Details of Euthygomphus yunnanensis Zhou & Wu, 1992 as illustrated in literature: a–g—after Zhou & Wu (1992: fig. 19–25), therein as Anisogomphus yunnanensis, h–t—after Asahina (1986: fig. 25–37), therein as Merogomphus parvus (sensu Asahina 1986 nec Krüger, 1899); a–g—the male holotype from China, Yunnan Province, Wandingzhen; h–n—males from Thailand (h–j—from Saraburi Province, Muak Lek); o–t—females from Thailand (p,r—from Mae Hon Son, q,s—from Saraburi Province, Muak Lek) and S. Vietnam, B'Lao; a,h—anal appendages, lateral view; b,i—ditto, dorsal view; c—head, frontal view; j,p,q—ditto, subfrontal view; r–t—occiput enlarged; e—penis (vesica spermalis), lateral view; f—posterior hamulus; g—anterior hamulus; m–n—secondary genitalia, lateral view; l—metafemur; o—end of abdomen, ventral view. Not to scale; a–b and e–g mirrored.

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FIGURE 9 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 9. Euthygomphus schorri, general habitus: a—holotype male; b—paratype female 2 from Cambodia, Mondulkiri Province, downstream of Buu Sraa Waterfalls.

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FIGURE 11 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 11. Abdominal structures of Euthygomphus schorri; a–j—holotype male; k—paratype female 2; a—cerci, lateral view; b—ditto, dorsal view; c—ditto, ventral view; d—right cercus, lateral view; e—secondary genitalia, lateral view; fhamuli, lateral view; g—penis distal part segment (V4), lateral view; h—ditto, dorsal view; i—ditto, oblique view; j—ditto, schematic outline; k—vulvar scale. Not to scale; i mirrored.

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FIGURE 8. A in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 8. A male of Euthygomphus schorri photographed in nature in Cambodia, Mondulkiri Province, 'Loringae brook' downstream of Buu Sraa Waterfalls, 10 vi 2014.

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FIGURE 1 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 1. Diagnostic details of males of some Euthygomphus spp. as illustrated in literature: a–h—anal appendages; i–j,l– m,o—distal part of penis (vesica spermalis); k,n—secondary genitalia; a–b,i–j—E. martini (therein considered as Merogomphus martini): a—after Fraser (1934: fig. 95b); i—after Fraser (1940: plate 3.3); b,j—after Lahiri (1987: fig. 364– 365,314–315); c–d,k—E. parvus (as Merogomphus parvus): c,k—from South Sumatra, Lampoeng residency, Giesting near Talang Padang, Wai Teboe River after Lieftinck (1941: plate 13, fig. 3–5); d—from Malay Peninsula, Central Perak, Sungai Plus, near Lasah after Lieftinck (1964: fig. 46); e–f,l–m,p–q—E. koxingai (as Anisogomphus koxingai): e,l,p—the holotype, after Chao (1954: fig. 293–297); f,m,q—from China, Guandong after Wilson & Xu (2009: fig. 1e–g); g,n—E. fujianensis, the holotype of Anisogomphus fujianensis, after Zhou & Wu (1992: fig. 16–18); h,o,r—A. chaoi, the holotype of Anisogomphus chaoi, after Liu (1991: fig. 3–8). Not to scale; i,b,k mirrored.

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FIGURE 2 in Reconsideration of the genera Merogomphus Martin, 1904, and Anisogomphus Selys, 1857, including erection of a new genus, with a new species and discussion of additional specimens from Cambodia

FIGURE 2. Diagnostic details of females of some Euthygomphus spp. as illustrated in literature: a–f—occiput; g–k—vulvar scale; a,g—E. parvus, the female syntype of Leptogomphus parvus after Schmidt (1935: Abb. 69–70) (therein considered as Merogomphus parvus); b—E. parvus from Malay Peninsula, Selangor, upstream Klang Gates after Lieftinck (1964: fig. 47) (as Merogomphus parvus); c,h—E. koxingai from China, Guandong after Wilson & Xu (2009: fig. 1k,m) (as Anisogomphus koxingai); d,i—E. wuzhishanus after Chao (1990: fig.5–9.6.1–2,4), the holotype of Anisogomphus wuzhishanus; e,j—E. chaoi, the allotype of Anisogompus chaoi, after Liu (1991: fig. 9,12); f,k—E. jinggangshanus, the holotype of Anisogomphus jinggangshanus, after Liu (1991: fig. 13–15). Not to scale.

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FIGURE 7 in A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes)

FIGURE 7. Lateral aspect of pectoral girdle (A) and dorsoventral view of pelvic girdle (B) of Styracura schmardae (MZUSP uncat. 508 mm DW). Abbreviations: adf, anterodorsal fenestra; avf, anteroventral fenestra; fct, additional supporting articular facet for propterygium; ip, iliac processes; isp, ischial processes; lpp, lateral prepelvic processes; msc, mesocondyle; mtc, metacondyle; of, obturator foramina; pc, procondyle; pdf, posterodorsal fenestra; pib, puboischiadic bar; ppp, prepelvic process; pvc, pelvic condyles; pvf, posteroventral fenestra; scp, scapular process; spf, scapular process fenestra.

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FIGURE 1 in A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes)

FIGURE 1. Styracura schmardae (ROM 66845, juvenile female, 276 mm DW, Guyana) in dorsal (A) and ventral (B) views. Live color brown or grayish- to olivaceous-brown.

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FIGURE 6 in A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes)

FIGURE 6. Head skeleton of Styracura schmardae (MCZ 37964, 545 mm DW, Cuba), showing small angular elements between hyomandibula and lower jaw (remarkably similar to specimen in Fig. 5); asterisk denotes terminal portion of hyomandibula. Abbreviations: ac, angular cartilage; fpf, frontoparietal fontenelle; hyo, hyomandibula; is, internasal septum; Mc, Meckel's cartilage; na, nasal aperture; pcf, precerebral fontenelle; pq, palatoquadrate; psc, prespiracular cartilage; sp, supraorbital process.

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FIGURE 5 in A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes)

FIGURE 5. Head skeleton of Styracura schmardae (MZUSP uncat. 508 mm DW, Belize). Note small calcified (angular) elements between hyomandibulae and lower jaws, which are slightly variable between both sides; asterisk denotes terminal portion of hyomandibula. Abbreviations: ac, angular cartilage; aoc, antorbital cartilage; fpf, frontoparietal fontenelle; hyo, hyomandibula; Mc, Meckel's cartilage; na, nasal aperture; pcf, precerebral fontenelle; pq, palatoquadrate; prp, preorbital process; psc, prespiracular cartilage.

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FIGURE 8. Phylogenetic inference obtained from a in A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes)

FIGURE 8. Phylogenetic inference obtained from a Maximum Likelihood analysis of the complete protein-coding mitochondrial genome (courtesy of L. Yang and G. Naylor); arrow indicates position of Styracura.

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FIGURE 4 in A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes)

FIGURE 4. Ventral lateral-line canals of Styracura schmardae (MZUSP uncat. 508 mm DW). Abbreviations: HMD, hyomandibular canal (pink); IOC, infraorbital canal (blue); jug, jugular component of hyomandibular canal; NAS, nasal canal (including the prenasal canal anteriorly; yellow); pst, posterior subpleural tubules; spl, subpleural component of hyomandibular canal; SPO, supraorbital canal (green).

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International Brain Laboratory public data

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Last verified 2026-04-29Open record