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Figs. 1–4. Megasoma sleeperi, third instar. 1 in New Field Record for the Rare Dung Beetle, Phanaeus melampus Harold (Coleoptera: Scarabaeidae, Scarabaeinae)

Figs. 1–4. Megasoma sleeperi, third instar. 1) epipharynx; 2) abdominal spiracle; 3) dorsal rt., lt. mandible; 4) ventral lt., rt. mandible.

opennotspecifiedMar 2006View details →
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Fig. 2 in Developmental Biology and Phenology of a Korean Native Dung Beetle, Copris ochus (Motschulsky) (Coleoptera: Scarabaeidae)

Fig. 2. Scale of tibial wear among field-collected C. ochus. Beetles with little or no tibial wear were taken to indicate recent emergence. (a) Unworn; (b) slightly worn; (c) worn.

opennotspecifiedDec 2004View details →
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Fig. 5 in Leaf-Litter Brood Chambers inDichotomius(Luederwaldtinia)Carbonarius(Mannerheim, 1829) (Coleoptera: Scarabaeidae): A Novel Behavior for Dung Beetles

Fig. 5. Probable sequence of construction of brood chambers: A) Original inclined burrow, B) Distal extreme enlarged, forming a spherical cavity, C) Cavity lined with entire or large fragments of leaves, D) Leaf litter at the distal pole leaving a hemispherical space for the egg, E) Egg chamber closed and more meniscate packets added until cavity completely filled, F) Burrow adjacent to proximal pole blocked with thick, perpendicular litter layers forming cylindrical protuberance and remaining burrow filled with soil.

opennotspecifiedSep 2013View details →
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Fig. 1. A in Leaf-Litter Brood Chambers inDichotomius(Luederwaldtinia)Carbonarius(Mannerheim, 1829) (Coleoptera: Scarabaeidae): A Novel Behavior for Dung Beetles

Fig. 1. A) General aspect of the kennel where brood chambers of Dichotomius carbonarius were collected; the soil is covered by leaf litter and short grasses and the tree trunks are fumo bravo, B) Leaf of fumo bravo on the soil and fragmented in subrectangular pieces, scale bar = 2 cm, C) Brood chamber of D. carbonarius showing contact with soil, scale bar = 2 cm, D) Adult D. carbonarius buried in the soil about 7 cm from the surface, scale bar = 1 cm.

opennotspecifiedSep 2013View details →
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Fig. 3. A in Leaf-Litter Brood Chambers inDichotomius(Luederwaldtinia)Carbonarius(Mannerheim, 1829) (Coleoptera: Scarabaeidae): A Novel Behavior for Dung Beetles

Fig. 3. A) Longitudinal section of a brood chamber of Dichotomius carbonarius, showing the egg chamber and some meniscate packets concavely downwards, scale bar = 1 cm, B) Egg with pharate larva inside and provisions with abundant trichomes (arrows), scale bar = 0.5 cm, C–D) Magnified trichomes, scale bar = 0.25 mm, E) Trichomes on a leaf of fumo bravo, scale bar = 1 mm.

opennotspecifiedSep 2013View details →
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Fig. 1 in Abundance and Seasonal Distribution of Predatory Coprophilous Argentine Rove Beetles (Coleoptera: Staphylinidae), and Their Effects on Dung Breeding Flies

Fig. 1. Predatory Staphylinidae collected in Castelar, Buenos Aires province: comparative numbers of the most common species per sampling 80 kg of cow dung.

opennotspecifiedMar 2003View details →
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Fig. 2 in Abundance and Seasonal Distribution of Predatory Coprophilous Argentine Rove Beetles (Coleoptera: Staphylinidae), and Their Effects on Dung Breeding Flies

Fig. 2. Predatory Staphylinidae collected in El Cadillal, Tucuman province: comparative numbers of the most common species per sampling date in 80 kg of cow dung.

opennotspecifiedMar 2003View details →
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Fig. 4 in The Pupation Chamber of Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae)

Fig. 4. Unambiguous optimization for the character "pupation chamber wall" on an adapted Scarabaeinae phylogenetic tree from Monaghan et al. (2007). Gray circles indicate pupation chamber wall character states: 0 – without visible pellets or pellets without a particular arrangement; 1 – row of pellets arranged in concentric rings; 2 – imbricate pellets grouped in slices oriented longitudinally; 3 – helicoid composed of whorls showing transverse rows of imbricate pellets. Dotted lines represent ambiguous optimization of the character. Node with * indicates when clusters A and B (Scarabaeinae) diverged, and probably when pellets arranged in rows in the construction of pupation chamber wall appeared. Black arrow shows a minimum age of approximately 54 million years for clade B. This is only for illustrative purpose. Branch lengths do not reflect any relative age for the groups. Principal genera within different polyphyletic tribes are: Canthonini 1: Panelus; Canthonini 2 (Ca 2): Temnoplectrom + Lepanus + Cephalodesmius + Saphabius; Canthonini 3 (Ca 3): Circellium; Canthonini 4 (Ca 4): Anachalcos; Neotropical Canthonini: Canthon + Megathoposoma + Malagoniella + Deltochilum; Coprini 1: Copris + Microcopris; Coprini 2 (Co 2): Coptodactyla; Coprini 3 (Co 3): Catharsius + Metacatharsius; Dichotomiini 1: Heliocopris; Dichotomiini 2 (Di 2): Uroxys + Bdelyropsis + Demarziella; Dichotomiini 3 (Di 3): Ateuchus; Dichotomiini 4 (Di 4): Ontherus + Canthidium; Onthophagini 1: Onthophagus + Euonthophagus + Hyalonthophagus + Caccobius + Milichus + Cleptocaccobius; Onthophagini 2: Proagoderus + Onthophagus + Digitonthophagus.

opennotspecifiedSep 2010View details →
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Fig. 2 in The Pupation Chamber of Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae)

Fig. 2. Micromorphology of pupation chambers of Sulcophanaeus imperator (A-C) and Anomiopsoides biloba (D-E). A) Internal (white arrows) and external parts of the wall, the latter represented by a set of small, lenticular, juxtaposed pellets (black arrows) that are composed of amorphous organic matter, dung fiber, and minerals, B) Detail of the internal part of the wall composed of flat, lenticular, juxtaposed pellets (white arrow), smaller than those of the external part of the wall, C) Roof of the pupation chamber showing acicular extensions (white arrow) toward the upper pole of the brood ball, where the egg chamber was originally located; in this specimen, part of the egg chamber wall (black arrow) is preserved, D) External part of the wall formed by poorly defined pellets that are composed of minerals and a scarce amount of dung fibers, which are delimited by amorphous organic matter (black arrows) that also cover mineral grain surfaces, E) Detail of the internal part of the wall that is formed by a smooth, thin layer of amorphous organic matter that covers the internal surface of the pupation chamber.

opennotspecifiedSep 2010View details →
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Fig. 3. Double helicoidal wall design. A in The Pupation Chamber of Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae)

Fig. 3. Double helicoidal wall design. A helicoid develops downward to the inferior pole (A), leaving space between whorls to lodge the whorls of the second helicoid, which develops upwards after changing its orientation in the inferior pole (B). These trajectories are complementary and when they are superposed, they form the helicoidal wall (C).

opennotspecifiedSep 2010View details →
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Fig. 1 in The Pupation Chamber of Dung Beetles (Coleoptera: Scarabaeidae: Scarabaeinae)

Fig. 1. Pupation chambers of scarabaeinae dung beetles. A) Dried brood ball of Ontherus sulcator that shows part of the helicoidal wall, B-D) Pupation chamber of Sulcophanaeus menelas showing (B) the whorls that form the helicoidal wall (white arrows), (C-D) the internal part of its wall (white arrow) and the roof (black arrows), E-F) Brood balls of Sulcophanaeus imperator that show (E) the external surface of the helicoidal wall of the pupation chamber when the wall is removed and (F) packets of pellets (white arrow) deposited by the larva on the layer of uneaten dung (black arrow), G) Brood ball of Malagoniella argentina that preserves an adult, which died during emergence, and shows three whorls (white arrows) of the helicoidal pupation chamber wall, H) Inferior pole of a Malagoniella argentina brood ball that shows the last section of one helicoid of the pupation chamber wall, I) Pupation chamber of Malagoniella argentina that shows the whorls of the helicoidal wall composed of transverse, bilobate rows (black arrows) of imbricate pellets, J-K) Pupation chambers of Malagoniella argentina showing (J) the change of the helicoid orientation in the inferior pole with its trajectory marked by a dotted white line (K), L) Pupation chamber of Anomiopsoides biloba, with helicoidal wall composed of only one whorl of juxtaposed pellets.

opennotspecifiedSep 2010View details →
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Fig. 2 in The Impact of Upgrading Roads on the Conservation of the Threatened Flightless Dung Beetle,Circellum bacchus(F.) (Coleoptera: Scarabaeidae)

Fig. 2. Map of the Addo Elephant National Park study site and the mortality transect routes (darkest lines). The Hapoor Road is tar and the Hapoor – Rooidam Track is gravel. Areas of thicket vegetation are dotted.

opennotspecifiedMar 2010View details →
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Fig. 2 in Arboreal Foraging Height in a Common Neotropical Dung Beetle, Canthon subhyalinus Harold (coleoptera: Scarabaeidae)

Fig. 2. Mean temperature at the ground and 41 m on tower over nine collecting days. Error bars are ±1 SD.

opennotspecifiedMar 2007View details →
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Fig. 1 in Arboreal Foraging Height in a Common Neotropical Dung Beetle, Canthon subhyalinus Harold (coleoptera: Scarabaeidae)

Fig. 1. Mean height versus abundance of beetles (numbers per trap) from ground level to top of the tower (columns, error bars 5 ±1 SD) and vertical temperature gradient at each trap level of the tower at 1100 hr, 10 December (line). Arrow denotes the average height of primates foraging in the area.

opennotspecifiedMar 2007View details →
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FIGURE 3 in A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa

FIGURE 3. Map of South Africa showing the type locality of Hathor spinosa Deschodt, new genus and species (red dot west of Pretoria).

opennotspecifiedNov 2023View details →
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FIGURE 1 in A new genus and species in the diverse dung beetle tribe Onthophagini Streubel, 1846 (Scarabaeidae: Scarabaeinae) from South Africa

FIGURE 1. Hathor spinosa Deschodt, new genus and species. (A) habitus in dorsal view showing the reduced fourth denticle of the protibiae; (B) habitus in ventral view; (C) habitus in lateral view showing the spine on the pronotum and; (D) frontal view showing the characteristic horns on the vertex and the triangular spine-like process situated anteromedially on the prothorax.

opennotspecifiedNov 2023View details →
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Figs. 41– 46. Reserva Ducke dung beetles. 41 in The Dung- and Carrion-Feeding Scarabs (Coleoptera: Scarabaeoidea) of an Amazonian Blackwater Rainforest: Results of a Continuous, 56-Week, Baited-PitfallTrap Study

Figs. 41– 46. Reserva Ducke dung beetles. 41) Deltochilum carinatum. Photograph by Jim McClarin; 42) Deltochilum septemstriatum. Photograph by Denis Faure; 43) Deltochilum pseudoicarus. Photograph by Trond Larsen; 44) Ateuchus sp. Photograph by Jim McClarin; 45) Canthidium bicolor with phoretic mite. Photograph by Trond Larsen; 46) Canthidium gerstaeckeri. Photograph by Trond Larsen. All images used by permission.

opennotspecifiedDec 2013View details →
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Figs. 35–40. Reserva Ducke dung beetles. 35 in The Dung- and Carrion-Feeding Scarabs (Coleoptera: Scarabaeoidea) of an Amazonian Blackwater Rainforest: Results of a Continuous, 56-Week, Baited-PitfallTrap Study

Figs. 35–40. Reserva Ducke dung beetles. 35) Onthophagus bidentatus. Photograph by Denis Faure; 36) Onthophagus rubrescens. Photograph by Trond Larsen; 37) Eurysternus caribaeus. Photograph by Trond Larsen; 38) Canthon quadriguttatus rolling a small piece of howler dung spattered on a leaf about 1 m above the ground. Photograph by Trond Larsen; 39) Canthon triangularis. Photograph by Trond Larsen; 40) Cryptocanthon peckorum. Photograph by Trond Larsen. All images used by permission.

opennotspecifiedDec 2013View details →
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Figs. 47–52. Reserva Ducke dung beetles. 47 in The Dung- and Carrion-Feeding Scarabs (Coleoptera: Scarabaeoidea) of an Amazonian Blackwater Rainforest: Results of a Continuous, 56-Week, Baited-PitfallTrap Study

Figs. 47–52. Reserva Ducke dung beetles. 47) Dichotomius boreus. Photograph by Trond Larsen; 48) Dichotomius mamillatus. Photograph by Trond Larsen; 49) Uroxys pygmaeus. Photograph by Trond Larsen; 50) Coprophanaeus lancifer. Photograph by Trond Larsen; 51) Oxysternon conspicillatum. Photograph by Doug Emlen and Mark Rowland; 52) Phanaeus chalcomelas. Photograph by Trond Larsen. All images used by permission.

opennotspecifiedDec 2013View details →
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FIGURE 3 in Onthophagus acernorus (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini) a new dung beetle species from Jalisco, Mexico

FIGURE 3. Onthophagus acernorus holotype male: A, aedeagus; B, medial endophallite and additional medial endophallite. Onthophagus chevrolati: C, aedeagus; D, medial endophallite and additional medial endophallite.

opennotspecifiedNov 2021View details →

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International Brain Laboratory public data

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Last verified 2026-04-29Open record

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Last verified 2026-04-29Open record